Dolichopus, Latreille, 1796

Zhang, Lili & Yang, Ding, 2008, New species of Dolichopus Latreille, 1796 from China (Diptera: Dolichopodidae), Journal of Natural History 42 (39 - 40), pp. 2515-2535 : 2516-2521

Key to species of Dolichopus from China (male)

1. Clypeus reaching or beyond lower margin of eyes................... 2

- Clypeus distinctly not reaching lower margin of eyes........................................................ 3 (subgenus Dolichopus )

2. Scape and pedicel distinctly elongated, cylindrical...................................................... 69 (subgenus Hygroceleuthus )

- Scape and pedicel short, conical.......... 72 (subgenus Macrodolichopus )

3. Femora mostly or wholly black................................ 4

- Femora mostly or wholly yellow.............................. 19

4. Postocular bristles wholly black................................ 5

- Middle and lower postocular bristles yellow...................... 11

5. Hind femur without long ventral bristles.......................... 6

- Hind femur with ventral bristles................................ 7

6. Apical 2/5 of fore femur and apical 1/3–1/4 of mid and hind femora yellow; hind tarsomere 1 with two d (Qinghai)................................................. Dolichopus (Dolichopus) turanicus Stackelberg, 1933

- Femora black; hind tarsomere 1 with three d (Xinjiang)................................... Dolichopus (Dolichopus) xinjianganus Yang, 1998

7. Mid and hind femora with two preacpical bristles; tibiae wholly black (Xinjiang).............. Dolichopus (Dolichopus) altayensis Yang, 1998

- Mid and hind femora with one preapical bristle; fore and mid tibiae yellow, at most with dark tip......................................... 8

8. Squama with yellow hairs (Heilongjiang, Xinjiang).................................. Dolichopus (Dolichopus) Mannerheimi Zetterstedt, 1838

- Squama with black hairs..................................... 9

9. Fore tarsomeres 1–2 with strong lateral bristles nearly at apical 1/3, tarsomere 3 with lateral bristles rather weak, tarsomeres 4–5 with strong lateral bristles (Heilongjiang)........... Dolichopus (Dolichopus) bonsdorfii Frey, 1915

- Fore tarsus without bristles as above (Shaanxi, Beijing).............. 10

10. Costal callus distinct....... Dolichopus (Dolichopus) lepidus Staeger, 1842

- Costal callus indistinct or stigma-like............................................... Dolichopus (Dolichopus) geniculatus Stannius, 1831

11. Fore tarsomere 5 flattened................................... 12

- Fore tarsomere 5 normal.................................... 13

12. Fore tarsomere 5 deeply bilobated ( China).............................................. Dolichopus (Dolichopus) sagittarius Loew, 1848

- Fore tarsomere 5 weakly incised apically (Qinghai).......................................... Dolichopus (Dolichopus) meyeri Yang, 1998

13. Hind femur with long ventral bristles........................... 14

- Hind femur without ventral bristles............................ 15

14. Hind femur with yellow ventral bristles; hind tibia black with basal 1/3 yellow (Xinjiang)............. Dolichopus (Dolichopus) clavipes Haliday, 1832

- Hind femur with black ventral bristles; hind tibia yellow with apical 1/3 black (Xinjiang).............. Dolichopus (Dolichopus) xinyuanus Yang, 1998

15. Tibiae wholly black........................................ 16

- At least fore and mid tibiae yellow, hind tibia not wholly black........ 17

16. Hind tarsomere 1 with one d (Qinghai)............................................ Dolichopus (Dolichopus) roborovskii Stackelberg, 1933

- Hind tarsomere 1 with three d (Qinghai, Xizang, Sichuan)........................................ Dolichopus (Dolichopus) shii Yang, 1996

17. Hind tibia and tarsomere 1 (except tip of the latter) yellow (Yunnan).......................... Dolichopus (Dolichopus) zhongdianus Yang, 1998

- Hind tibia with at least apex and entire hind tarsus black............ 18

18. First flagellomere distinctly obtuse (Heilongjiang)............................................... Dolichopus (Dolichopus) yangi sp. nov.

- First flagellomere more or less acute (Xinjiang)............................................ Dolichopus (Dolichopus) basisetus Yang, 1998

19. Postocular bristles wholly black............................... 20

- Middle and lower postocular bristles yellow...................... 26

20. Hind femur with ventral bristles............................... 21

- Hind femur without ventral bristles............................ 23

21. Mid and hind femora with one preapical bristle; hind femur with yellow ventral bristles................................................. 22

- Mid femur with three preapical bristles; hind femur with two preapical bristles; hind femur with black long hair-like ventral bristles (Xinjiang).............................. Dolichopus (Dolichopus) ungulatus Linnaeus, 1758

22. Hind femur with ventral hairs slightly shorter than thickness of femur; fore tibia with one apv 1/3 as long as fore tarsomere 1 (Shandong)..................................... Dolichopus (Dolichopus) robustus Stackelberg

- Hind femur with ventral hairs distinctly longer than thickness of femur; fore tibia without distinct apv (Xinjiang)................................................. Dolichopus (Dolichopus) brevipennis Meigen, 1824

23. Fore tarsomeres 4–5 flattened and tarsomere 5 deeply bilobated (Xinjiang)....................... Dolichopus (Dolichopus) grunini Smirnov, 1948

- All tarsi simple........................................... 24

24. Arista nearly apical, swollen apically (Shandong).............................. Dolichopus (Dolichopus) romanovi Smirnov and Negrobov, 1973

- Arista dorsal, not swollen apically............................. 25

25. Squama yellow with yellow hairs (Yunnan).............................................. Dolichopus (Dolichopus) yunnanus Parent, 1930

- Squama yellow with black hairs (Xinjiang)............................................. Dolichopus (Dolichopus) rupestris Haliday, 1833

26. Fore or mid tarsus broadened, plumose, silvered or white............ 27

- All tarsi simple........................................... 35

27. Fore tarsus with segment(s) modified........................... 29

- Mid tarsus with segment(s) modified........................... 28

28. Mid tarsomere 1 flattened with thin lateral bristles; mid tibia with brown dorsal stripe (Heilongjiang, Hebei, Henan, Shanxi, Inner Mongolia, Xinjiang, Qinghai, Xizang)........ Dolichopus (Dolichopus) plumipes Scopoli, 1763

- Mid tarsus with apex of tarsomere 3 and tarsomeres 4–5 flattened (Jilin)............................. Dolichopus (Dolichopus) jilinensis sp. nov.

29. Arista swollen apically (Henan) Dolichopus (Dolichopus) henanus Yang, 1999

- Arista simple............................................ 30

30. Fore tarsomeres 4 and (or) 5 flattened.......................... 31

- Fore tarsomere 5 bilobated.................................. 34

31. Hind femur with short hair group; squama pale-haired (Heilongjiang)............................ Dolichopus (Dolichopus) galeatus Loew, 1871

- Hind femur without ventral hair group; squama black-haired.......... 32

32. Costal callus long and thick ( China)................................................... Dolichopus (Dolichopus) calceatus Parent, 1927

- Costal callus indistinct..................................... 33

33. Fore tarsomere 4 thick, 1.5–2.0 times as long as wide (Heilongjiang, Hebei, Beijing, Inner Mongolia, Xinjiang)................................................... Dolichopus (Dolichopus) plumitarsis Fallén, 1823

- Fore tarsomere 4 slender, 2.5 times as long as wide (Heilongjiang, Inner Mongolia)............... Dolichopus (Dolichopus) simius Parent, 1927

34. Costal callus long and thick (Xinjiang)................................................. Dolichopus (Dolichopus) hejingensis Yang, 1998

- Costal callus indistinct (Xinjiang)................................................. Dolichopus (Dolichopus) reichardti Stackelberg, 1933

35. Hind femur with ventral hairs or bristles........................ 36

- Hind femur without ventral hairs or bristles, at most with sort hairs..... 46

36. Hind femur with black ventral hairs (bristles)..................... 37

- Hind femur with yellow or brown ventral hairs (bristles)............. 38

37. Mid and hind femora with two preapical bristles; fore tibia without distinct apv ( China)............ Dolichopus (Dolichopus) angustipennis Kertész, 1901

- Mid and hind femora with one preapical bristle; fore tibia with one brown thin apv (Yunnan)............ Dolichopus (Dolichopus) ancistrus Yang, 1996

38. Fore tibia with distinct apv.................................. 39

- Fore tibia without apv..................................... 40

39. Hind femur with yellow ventral hairs much longer than thickness of femur. Fore tibia with pale and slender apv about half as long as fore tarsomere 1; hind tibia with brown long lateral hairs, without distinct dorsal bristles (Xinjiang).................. Dolichopus (Dolichopus) jaxarticus Stackelberg, 1927

- Hind femur with yellow ventral hairs as long as thickness of femur. Fore tibia with apv dark brown and short (1/3 as long as fore tarsomere 1); hind tibia with dorsal bristles, without long lateral hairs (Xinjiang)........................................ Dolichopus (Dolichopus) clavipes Haliday, 1832

40. M with rudimentary M 2 .................................... 41

- M without rudimentary M 2 .................................. 43

41. Hind femur with short ventral hairs; costal callus moderately long (Henan)............ Dolichopus (Dolichopus) luoshanensis Yang and Saigusa, 1999

- Hind femur with long ventral hairs; costal callus punctiform.......... 42

42. Hind tibia wholly yellow (Zhejiang, Guizhou)..................................... Dolichopus (Dolichopus) zhejiangensis Yang and Li, 1998

- Apical half of hind tibia black (Heilongjiang, Jilin, Hebei, Inner Mongolia, Ningxia, Shaanxi, Xinjiang)..................................................... Dolichopus (Dolichopus) martynovi Stackelberg, 1930

43. Hind tarsomere 1 with three d................................ 44

- Hind tarsomere 1 with one d................................. 45

44. Costal callus distinct and thick (Heilongjiang, Xinjiang)........................................ Dolichopus (Dolichopus) hilaris Loew, 1862

- Costal callus punctiform (Heilongjiang, Inner Mongolia).................................... Dolichopus (Dolichopus) orientalis Parent, 1927

45. Hind femur with ventral hairs shorter than half thickness of femur; hind tarsomere 1 yellow; male genitalia yellow (Heilongjiang)............................... Dolichopus (Dolichopus) xanthopyga Stackelberg, 1930

- Hind femur with ventral hairs much longer than thickness of femur; hind tarsus black; male genitalia black ( Mongolia)................................................... Dolichopus (Dolichopus) longipilosus sp. nov.

46. Fore tibia with distinct apv.................................. 47

- Fore tibia with apv very short or indistinct....................... 65

47. M with rudimentary M 2 .................................... 48

- M without rudimentary M 2 .................................. 56

48. Squama with pale hairs..................................... 49

- Squama with black hairs.................................... 50

49. Wing distinctly brown on anterior part (Hebei, Beijing, Tianjin)............................... Dolichopus (Dolichopus) aubertini Parent, 1936

- Wing hyaline............ Dolichopus (Dolichopus) erroneus Parent, 1926

50. Cercus with marginal bristles very long (longer than cercus) (Henan, Shanghai)........................ Dolichopus (Dolichopus) nitidus Fallén, 1823

- Cercus with marginal bristles not longer than cercus................ 51

51. Hind tibia with extreme base (basal 1/8) yellow (Yunnan)................................... Dolichopus (Dolichopus) legendrei Parent, 1930

- Hind tibia at least with basal 1/3 yellow......................... 52

52. Hypandrium curved apically; inner epandrial lobe club-like and densely pubescent near apex (Beijing, Henan, Shandong, Shaanxi, Sichuan, Anhui, Jiangsu, Zhejiang)...... Dolichopus (Dolichopus) bigeniculatus Parent, 1926

- Hypandrium not curved apically; inner epandrial lobe not as stated above 53

53. Basal half of hind tarsomere 1 yellow (Henan, Shaanxi, Sichuan, Hubei, Yunnan, Guizhou, Fujian, Guangxi, Shanghai, Zhejiang).................................... Dolichopus (Dolichopus) simulator Parent, 1926

- Hind tarsomere 1 wholly black............................... 54

54. M 1 with one short appendent vein M 2 .......................... 55

- M 1 without appendent vein.................................. 56

55. Fore tibia with one distinct apv.................................................... Dolichopus (Dolichopus) cuneipennis Parent, 1926

- Fore tibia without distinct apv.................................................... Dolichopus (Dolichopus) howjingleei Olejníček, 2002

56. Inner epandrial lobe rounded apically (Henan, Yunnan, Guizhou, Guangxi)..................... Dolichopus (Dolichopus) meridionalis Yang, 1996

- Inner epandrial lobe acute apically (Guizhou, Taiwan) ........................................ Dolichopus (Dolichopus) exsul Aldrich, 1922

57. All coxae yellow (Heilongjiang, Jilin, Beijing, Inner Mongolia, Gansu, Xinjiang, Qinghai)............... Dolichopus (Dolichopus) linearis Meigen, 1824

- Usually only fore coxa yellow, not all coxae yellow................. 58

58. Arista near apical. All coxae black; fore femur black except apex (Yunnan)....................... Dolichopus (Dolichopus) subapicalis Yang, 1998

- Arista dorsal. Mid and hind coxae black; fore femur usually yellow..... 59

59. Antenna with first flagellomere elongated, two times longer than wide, acute apically................................................ 60

- Antenna with first flagellomere at most 1.5 times longer than wide...... 63

60. Costal callus indistinct (Beijing)......................................... Dolichopus (Dolichopus) zhoui Zhang, Yang and Grootaert, 2004

- Costal callus long and thick.................................. 61

61. Arista arising near base of first flagellomere; hind tarsomere 1 with 2 d (Beijing, Gansu, Shaanxi).......... Dolichopus (Dolichopus) tewoensis Yang, 1998

- Arista arising at middle of first flagellomere...................... 62

62. Hind coxa black; hind tarsomere 1 with one d (Xinjiang).................... Dolichopus (Dolichopus) qinghensis Zhang, Yang and Grootaert, 2004

- Hind coxa mostly yellow with brown spot, hind tarsomere 1 with 2 d ( China)................... Dolichopus (Dolichopus) longicornis Stannius, 1831

63. Wing with apical half distinctly brownish; costal callus long and distinct; squama with pale hairs (Xinjiang)...................................................... Dolichopus (Dolichopus) zernyi Parent, 1927

- Wing hyaline; squama with black hairs.......................... 64

64. Hind tibia yellow with black apex (Heilongjiang, Hebei, Beijing, Shaanxi)...................... Dolichopus (Dolichopus) uniseta Stackelberg, 1929

- Hind tibia black with extreme base yellow (Hunan)......................................... Dolichopus (Dolichopus) alticola Parent, 1930

65. Squama with pale hairs..................................... 66

- Squama with black hairs.................................... 67

66. Antenna wholly black. Hypandrium bifurcate; cercus nearly triangular (Xinjiang)............... Dolichopus (Dolichopus) nubilis Meigen, 1824

- Antenna with first flagellomere black. Hypandrium simple; cercus roundish ( China)................. Dolichopus (Dolichopus) divisus Becker, 1917

67. First flagellomere as long as wide (Jilin)............................................. Dolichopus (Dolichopus) ringdahli Stackelberg, 1929

- First flagellomere distinctly elongated........................... 69

68. First flagellomere elongated, two times longer than wide, acute apically; fore and hind coxae yellow, mid coxa black (North China)....................................... Dolichopus (Dolichopus) vaillanti Parent, 1927

- First flagellomere 1.7 times longer than wide, obtuse apically; fore coxa yellow, mid and hind coxae black (Hebei, Inner Mongolia, Ningxia, Gansu).............................. Dolichopus (Dolichopus) agilis Meigen, 1824

69. Fore and hind femora mostly black (Xizang, Qinghai).............................. Dolichopus (Hygroceleuthus) brevifacies Stackelberg, 1925

- All femora yellow......................................... 70

70. First flagellomere triangular in lateral view (North China)............................... Dolichopus (Hygroceleuthus) tenuicornis Parent, 1927

- First flagellomere not triangular............................... 71

71. Wing with an incision on basal half (Qinghai)....................................... Dolichopus (Hygroceleuthus) rotundipennis Loew, 1848

- Wing without incision ( China).................................................... Dolichopus (Hygroceleuthus) latipennis Fallén, 1823

72. First flagellomere elongated, nearly two times longer than wide. Fore and mid femora with black apex (North China)........................................ Dolichopus (Macrodolichopus) obscuripes Stackelberg, 1925

- First flagellomere nearly as long as wide. All femora yellow ( China)....................... Dolichopus (Macrodolichopus) diadema Haliday, 1832