Erymanthus Spinola, 1841

Gerstmeier, Roland & Salvamoser, Max, 2014, Revision of the checkered beetle genus Erymanthus Spinola, 1841 (Coleoptera, Cleridae, Clerinae), Zootaxa 3755 (6), pp. 501-548 : 503

Erymanthus Spinola, 1841

Type species: Erymanthus gemmatus Klug, 1842

Spinola 1841: 75; Klug 1842: 327; Lacordaire 1857: 457; Desmarest in Chenu 1860: 269; Schenkling 1903: 28, 64; Corporaal 1949: 355; Opitz 2010: 66.

Head: Head including eyes wider than base of pronotum, eyes coarsely facetted, reniform, protruding laterally, deeply emarginated at antennal insertion, separated by one to two eye widths, ocular notch distinct; frons with two grooves or two gibbosities; labrum bilobed, terminal segment of labial palpomeres securiform; terminal maxillary palpomeres spatulate, widened distally; gular sutures apically converging, gular process broad with two papillae ( Fig. 1 ); antennae comprised of 11 antennomeres; antennomere 1 with one long hair; antennomeres 2–8 filiform, 9– 10 triangular, 9–11 forming a distinct club with dense light pubescence; male specimens with a small pit-like sensilla on antennomere 11 ( Fig. 2 ).

Thorax: Procoxal cavities closed behind; pro-intercoxal process narrow, broadly dilated posteriorly ( Fig. 3 ); pronotum longer than broad; pronotum subparallel, widest in middle, basal collar conspicuously narrower, pronotal arch slightly narrower, sometimes with strong lateral gibossities; pronotal disc with more or less conspicuously gibbosities and always with a longitudinal elongate-ovate groove centrally; mesocoxal cavities open laterally; anterior mesoventral process present, forming a small convexity; intercoxal mesoventral process more or less wide, protruding laterally; mesepisternum with fine punctation anteriorly, surface smooth posteriorly; scutellum oblongovate; hypomeron tapering apically towards intercoxal process, forming a typical “edge” towards base; metendosternite with conspicuous furcal arms, laminae broadly triangular ( Fig. 4 ).

Elytra: Elongate, almost two times longer than broad, base not margined, usually with strong humeri; apex strongly dilated, declivous; elytral punctation in basal part arranged into ten striae, punctation behind humeral part mostly irregular; punctation crateriform, circular to oblong-ovate, sometimes rasp-like; in the posterior part many gibbosities without regular arrangement; posterior half usually with five large gibbosities with dense setae and more or less regular distribution ( Fig. 6 ); hind wing with AA3+4 and CuA2 absent (hence wedge cell absent), MP4 absent or with vetige evident at junction of MP3 and MP3+4.

Legs: Legs stout, femora conspicuously swollen, especially profemora; pro- and mesotibiae strongly bent, metatibiae only slightly bent, almost straight; carinae of pro- and mesotibiae over entire length, carinae of metatibiae extending from two-thirds to full length; inner edge of pro- and mesofemora and -tibiae with a row of short, thick bristles; claws simple; tarsal pulvillar formula 3–3–3; pulvilli deeply bilobed; tibial spur formula 1–1– 1 or 1–2–1 ( E. pustulatus only).

Abdomen: Six visible abdominal ventrites; abdominal metaintercoxal process narrow; pygidium broadest at the beginning of the processes, pygidial index (PL:PW) between 1:0.8 and 1:1; tegminal struts conspicuously long and variable in shape and width.

Figures

FIGURES 1 – 7. Morphological characters of the genus Erymanthus. (1) Head, lower surface (scale bar 1 mm), (2) Antennae, left female, right male with pit-like sensilla (scale bar 1 mm), (3) Pro- and Mesothorax, lower surface (scale bar 1 mm), (4) Metendosternite (scale bar 0.5 mm), (5) Protarsus (scale bar 1 mm), (6) Gibbosity diagram, (7) Wing venation (scale bar 2 mm).