Merodon serrulatus (Wiedemann in Meigen, 1822)

Vujic, Ante, Likov, Laura, Radenkovic, Snezana, Tubic, Natasa Kocis, Djan, Mihajla, Sebic, Anja, Perez-Banon, Celeste, Barkalov, Anatolij, Hayat, Ruestem, Rojo, Santos, Andric, Andrijana & Stahls, Gunilla, 2020, Revision of the Merodon serrulatus group (Diptera, Syrphidae), ZooKeys 909, pp. 79-158 : 79

publication ID	https://dx.doi.org/10.3897/zookeys.909.46838
publication LSID	lsid:zoobank.org:pub:22B7FF16-D0A2-40F9-B20E-F7C6E0AF1842
persistent identifier	https://treatment.plazi.org/id/13F007B4-C081-58EC-892E-77228DCD6BAF
treatment provided by	ZooKeys by Pensoft
scientific name	Merodon serrulatus (Wiedemann in Meigen, 1822)
status

Treatment

Merodon serrulatus (Wiedemann in Meigen, 1822) Figs 1A-G View Figure 1 , 2A-D View Figure 2 , 3A-J View Figure 3 , 4A-F View Figure 4 , 5A-D View Figure 5 , 6A-G View Figure 6 , 14C View Figure 14

Merodon alexeji Paramonov, 1925: 155 - syn. published in Vujić et al. 2011: 84.

Merodon lusitanicus Hurkmans, 1993: 181 - syn. published in Marcos-García et al. 2007: 566.

Merodon tener Sack, 1913: 443 syn. nov.

Diagnosis.

Medium sized (7.1-10.9 mm), short pilose dark species with olive-brown reflection; antennae dark brown; legs mostly black; body pile predominantly pale yellow, except black pile on vertex and scutum, terga 2-4 in some specimens and apical one third of femora in some specimens and populations; basoflagellomere elongated (1.7-2.2 times as long as wide) obviously concave dorsally, arista short (Fig. 3 View Figure 3 ); tergum 2 usually with small pale orange-yellow lateral maculae (Fig. 2 View Figure 2 ); metafemur incrassate, ca. three times longer than wide, with short pilosity, except few long pile on postero- and anteroventral surface of metafemur (Fig. 4 View Figure 4 ); male genitalia: apical part of anterior surstyle lobe triangular (Fig. 1A View Figure 1 : al, C, D); posterior surstyle lobe with lateral hump (Fig. 1A, B View Figure 1 : lp); lingula large (Fig. 1E View Figure 1 : l).

Redescription

(based on the types and specimens from the type area of nominal taxon, Iberian Peninsula; variability includes populations from all of the range). Male. Head. Antennae black to dark brown; basoflagellomere (Fig. 3A-C, G-J View Figure 3 ) elongated, 1.7-2.2 times as long as wide, and 2.5-3 times as long as pedicel, concave dorsally, tapering to the apex; dorsolateral and dorsomedial (if present) fossette large with variable shape (see variability) (as on Fig. 3 View Figure 3 ); arista dark and thickened at basal one third, covered with dense microtrichia; arista short, 1.2-1.5 times as long as basoflagellomere (Fig. 3 View Figure 3 ); face and frons black with gray microtrichia, face covered with dense whitish, and frons with yellowish gray pile; oral margin shiny, with small lateral microtrichose area; lunule shiny black, bare; eye contiguity 8-10 facets long; vertex isosceles, shiny black, except in front of anterior ocellus, covered with microtrichia; vertex with long, pale whitish yellow pile, in some cases mixed with few black pile on the ocellar triangle; ocellar triangle from equilateral to isosceles (see variability); occiput with gray-yellow pile, ventrally covered with a dense, gray microtrichia; eyes covered with dense whitish pile (Fig. 5A, B View Figure 5 ); ratio of length of vertical triangle: eye contiguity: frons = 3: 1: 3.

Thorax. Scutum and scutellum black with bronze luster, covered with dense, erect, usually yellow pile; scutum at wing basis in some specimens and populations with patch of black pile, or with fascia of black pile between wing basis; scutum usually with two or four microtrichose vittae (see variability), anteriorly connected and posteriorly reaching the scutellum (Fig. 2A View Figure 2 ); anterior half of scutum from dull until shiny black (see variability); posterodorsal part of anterior anepisternum, posterior anepisternum (except anteroventral angle), anterior anepimeron, dorsomedial anepimeron, and posterodorsal and anteroventral parts of katepisternum with long, dense pale yellow pile and grayish microtrichia; wings entirely covered with microtrichia; wing veins brown; calypteres pale yellow; halteres yellow, in some cases with dark capitulum; legs (Fig. 4 View Figure 4 ) without spinae or other protuberances; legs mostly black, except brown tarsi ventrally in some specimens; pile on legs pale yellow, except black pile at apical one third of metafemur in some populations (see variability); metafemur moderately incrassate, ca. three times longer than wide; long pile on postero- and anteroventral surface sparse, and ca. one third to one fourth (see variability) of width of metafemur, approximately the same length as pile on dorsal surface (Fig. 4 View Figure 4 ).

Abdomen. Tapering posteriorly, ca. 1.2 times longer than mesonotum; terga dark brown to black, except for a pair of pale yellow-orange, triangular, lateral maculae on tergum 2 (in some specimens less visible: see variability); terga 3 and 4 each with a pairs of white microtrichose, oblique fasciae (on tergum 2 triangular); color of pile on terga variable, from all yellow to specimens with many black pile on terga 2-4 (see variability) (Fig. 2C, D View Figure 2 ); sterna dark brown, covered with long whitish yellow pile.

Male genitalia. Apical part of anterior surstyle lobe triangular shape, 1.1-1.4 times longer than wide, covered with dense, short pile (Fig. 1A View Figure 1 : al, C, D); posterior surstyle lobe oval with basolateral protrusion (lateral hump) (Fig. 1A, B View Figure 1 : bp, 14C: bp); cercus rectangular (Fig. 1A View Figure 1 : c); hypandrium sickle-shaped, without lateral projections; lingula large (Fig. 1E View Figure 1 : l).

Female. Similar to the male except for normal sexual dimorphism and for the following characteristics: antennae with rounded tip, basoflagellomere ca. two times longer than wide, fossette dorsal (Fig. 3D-F View Figure 3 ); frons with microtrichose vittae along eye margins variable in shape and size (see variability); frons covered with variable color of pilosity, from mostly gray-yellow until predominately black (see variability) (Fig. 5C, D View Figure 5 ); ocellar triangle covered with black pile; lateral side of terga, anterior two third of tergum 2 and all tergum 5 with yellow pile; central part of terga 2-4 with short adpressed black pile; microtrichose fasciae on terga 3 and 4 conspicuous (Fig. 2D View Figure 2 ).

Variability.

There is some intra- and interpopulation variability in the morphological characters of Merodon serrulatus , which are summarized in Table 1 View Table 1 .

Distribution.

As shown in Fig. 7 View Figure 7 , this Merodon taxon is characterized by the greatest range, extending from Iberian Peninsula in the south-west, through Greece and eastern Turkey to the south, and eastward to Siberia and Mongolia (Doczkal pers. comm.).

Ecology.

Preferred environment: forest/open ground; thermophilous Quercus forest; Castanea forest ( Ståhls et al. 2009), evergreen oak forest ( Quercus ilex L. and Q. suber L.), dry Pinus forest; lentisc scrub; dry, well-vegetated, calcareous and non-calcareous unimproved grassland and tracksides; hedgehog heath ( Speight 2018); Pinus , Picea , and Larix forests (Siberia) (Fig. 35B View Figure 35 ). Flowers visited: Umbelliferae ; Cirsium spp., Helianthemum spp., Potentilla spp., Rosa spp., Thapsia spp., and Thymus spp. ( Speight 2018). Flight period: April-August.

Type material.

Holotype of Merodon serrulatus [original designation in Meigen (1822: 360)]: Wiedemann in Meigen (1822) as Lampetia serrulata : "Portugal / Hoffmannsegg S." 1 ♀, (ZHMB) (studied).

Merodon alexeji : Described by Paramonov (1925) based on two specimens (male and female). Lectotype [designated by Marcos-García et al. (2007)]: male, " Merodon / alexeji n. sp. / Typus / Paramonov d.", "Kohanovka / Baltsk. u. / Odes. g. [in Cyrillic] 1.VI.24. Ucraina" (PC) (SIZK) (studied).

Merodon lusitanicus : Holotype [original designation by Hurkmans (1993: 181)]: female, "Portugal, Algarve, Quarteira 27.iv.1985, J.A.W. Lucas" (NBCN) (studied). Paratypes. Portugal • 1 ♀; Algarve, Quarteira; 37°03'29"N, 8°04'47"W; 27 Apr. 1985; NBCN • 1 ♀; Algarve, Vilamoura; 37°04'35"N, 8°07'46"W; 27 Apr. 1985; NBCN.

Merodon tener : Described by Sack (1913: 443) based on three male and three female syntypes. Lectotype [designated by Hurkmans (1993)]: female "Sarepta [= Krasnoarmeysk near Volgograd, after Peck 1988] / M. tener Sack det. Sack / coll. Lichtwardt / coll. D. E. I. Eberswalde" (ZHMB) (studied). Original label: "LECTOTYPE of / M. tener Sack / des. 1988 Hurkmans" [red label handwritten], “Sarepta” [yellow label handwritten], "M. tener Sack / ♀ det. Sack" [label partly handwritten], "Coll. DEI / Eberswalde", "Coll. Lichtwardt". (See Supplementary file 6: Figure 6A View Figure 6 ). Lectotype is conspecific with type of M. serrulatus, sharing the same morphological characters.

Other material.

Croatia • 1 ♀; Velebit, Brušane; 44°29'55"N, 15°16'43"E; 600 m a.s.l.; 13 Jun. 1969; NBCN 02489.

France • 1 ♀; Languedoc-Roussillon, Corbieres, Carcassonne; 43°13'00"N, 2°21'00"E; 18 Jun. 1974; NBCN • 1 ♂; Provence Alpes Cote d’Azur, Saint-Maximin-la-Sainte-Baume; 43°25'23"N, 5°50'11"E; 17-20 Jun. 1951; M. Bequaert leg.; NBCN 02491 • 1 ♀; same data as for preceding; NBCN 02492 • 1 ♀; Source du Lez, Saint Clement; 43°43'05"N, 3°50'39"E; 24 May 1989; Maldes leg.; MNHN 22629 • 1 ♂; Provence Alpes Cote d’Azur, Montagne du Luberon, W from Bonnieux; 43°48'00"N, 5°22'00"E; 3 Jun. 1993; NBCN • 1 ♀; Feuilla, Route de Treilles, en face du village Panais; 43°53'53"N, 2°00'16"E; 7 Jun. 1988; J. Hamon leg.; MNHN 17973 • 3 ♂♂; Departement du Gard, Mas Mejean; 44°05'24"N, 3°35'26"E; 29 May 1952; NBCN • 2 ♂♂; P. N. Mercantour, Le Bor, on, Umgebung, mesophiles pot. Argent; 44°06'47"N, 7°16'42"E; 1380 m a.s.l.; 21 Jun. 2011; A. Ssymank leg.; A. S. coll. G1057 • 1 ♂; Causse de Sauveterre; 44°22'03"N, 3°13'49"E; 20 Jul. 1971; MNHN 22628 • 1 ♀; Larche (Basses Alpes); 44°26'59"N, 6°50'60"E; 22 Jul. 1925; R. Benoist leg.; MNHN PM0383 • 1 ♂; same data as for preceding; MNHN PM0429 • 1 ♂; same data as for preceding; 3 Jul. 1925; MNHN 22630 • 1 ♂; Provence Alpes Cote d’Azur, Larche; 44°26'59"N, 6°50'60"E; 22 Jul. 1923; NBCN • 1 ♂; Drome, La Chapelle en Vercors; 44°58'12"N, 5°23'39"E; 28 Jun. 1970; Roman Emile leg.; MNHN PM0377 • 2 ♂♂; Isere, Villars de Lans Pic st Michel hill top; 45°05'24"N, 5°37'12"E; 1970 m a.s.l.; 20 Jul. 2010; J. van Steenis leg.; J. v. S. coll. • 1 ♀; L’Arselle; 45°23'10"N, 7°04'19"E; 14 Jun. 1909; MNHN PM0357.

Greece • 2 ♀♀; Mountain Taygetos; 22 km SW Sparta; 36°58'60"N, 22°24'09"E; 6 May 1990; NBCN • 1 ♂; Mountain Taygetos; 37°05'20"N, 22°18'55"E; 950-1800 m a.s.l.; 15-19 May 1990; ZMUC 00513256 • 7 ♂♂; Laconia, Karyes, 25 km N from Sparta; 37°18'15"N, 22°25'16"E; 930 m a.s.l.; 23 May 2014; A. Vujić, J. Ačanski leg.; FSUNS 06535, 06542, 06547, 06549, 06556, 06563, 06560 • 6 ♀♀; Laconia, Karyes, 25 km N from Sparta; 37°18'15"N, 22°25'16"E; 930 m a.s.l.; 23 May 2014; A. Vujić, J. Ačanski leg.; FSUNS 06543, 06544, 06546, 06553, 06555, 06565 • 3 ♂♂; Chelmos, Kalavryta ski center; 38°00'25"N, 22°11'40"E; 6 Jun. 2017; A. Vujić, Z. Nedeljković, L. Likov, M. Miličić, T. Tot, leg.; FSUNS 15980 to 15982 • 2 ♀♀; Chelmos, Kalavryta ski center; 38°00'25"N, 22°11'40"E; 6 Jun. 2017; A. Vujić, Z. Nedeljković, L. Likov, M. Miličić, T. Tot leg.; FSUNS 15983, 15984 • 1 ♂; Achaia, Mountain Chelmos above Kalavryta; 38°00'31"N, 22°07'08"E; 1700 m a.s.l.; 17-19 Jun. 1982; B. Skule, S. Langemark leg.; ZMUC 00513264 • 3 ♀♀; Achaia, Mountain Chelmos above Kalavryta; 38°00'31"N, 22°07'08"E; 1700 m a.s.l.; 17-19 Jun. 1982; B. Skule, S. Langemark leg.; ZMUC 00513265, 00513273, 00513301 • 2 ♂♂; Corfu; 39°40'00"N, 19°45'00"E; NHMW 02485, 02486 • 2 ♂♂; Corfu; 39°40'00"N, 19°45'00"E; 1400 m a.s.l.; NHMW • 1 ♀; Peristeri mountain; 39°40'36"N, 21°07'06"E; 2030 m a.s.l.; 24-28 May 1994; V. Michelsen leg.; ZMUC 00513259; • 1 ♂; 15 km NO Metsovo; 39°47'19"N, 21°11'58"E; 4 Jun. 1994; M. Ohl leg.; ZHMB • 4 ♂♂; Mountain Pindos, Katara Pass; 39°47'46"N, 21°13'44"E; 20 May 1997; S. Radenković leg.; FSUNS • 3 ♂♂; same data as for preceding; S. Šimić leg.; FSUNS • 4 ♀♀; same data as for preceding; A. Vujić leg.; FSUNS • 6 ♂♂; Mountain Pindos, Katara Pass; 39°47'48"N, 21°13'45"E; 1700 m a.s.l.; 20 May 1997; FSUNS 01779, 01781 to 01785 • 3 ♀♀; same data as for preceding; FSUNS 01786, 01787, 01780 • 1 ♀; Mountain Pindos, Katara Pass; 39°54'00"N, 21°11'00"E; 13 Jul. 1979; M. C. D Day, G. R. Else, D. Morgan leg.; NHMUK • 1 ♂; Mountain Pindos, "Iznad Panagije" [Panagia]; 39°48'25"N, 21°19'44"E; 850 m a.s.l.; 15 May 2011; A. Vujić leg.; FSUNS H38 • 15 ♂♂; Mountain Olympos, Litochoras-Prionia 3, "proplanak pored puta"; 40°04'36"N, 22°00'46"E; 17 May 2012; A. Vujić leg.; FSUNS H82, H83, H86 to H90, H92, H93, H95 to H97, H99, I6, I7 • 1 ♂; Mountain Olympos, Litochoras-Prionia 3, "proplanak pored puta"; 40°04'36"N, 22°00'46"E; 17 May 2012; FSUNS H94 • 1 ♂; Mountain Olympos, Near Litochoro; 40°06'30"N, 22°28'41"E; 21 May 2014; A. Vujić, J. Ačanski leg.; FSUNS 06499 • 1 ♀; Mt Olympos, Litochoro; 40°06'41"N, 22°28'37"E; 650 m a.s.l.; 17 May 2016; A. Vujić, J. Ačanski, M. Miličić, Z. Nedeljković leg.; FSUNS 11679 • 8 ♂♂; Mountain Olympos, Litochoras-Prionia 4; 40°06'43"N, 22°28'08"E; 18 May 2011; A. Vujić leg.; FSUNS I11 to I14, I17, I20 to I22 • 2 ♀♀; same data as for preceding; FSUNS I15, I16.

Italy • 1 ♀; Sicily, Etna, Rifugio Filiciusa; 37°43'14"N, 15°02'51"E; 1400-1500 m a.s.l.; 22-28 Jul. 1961; V. S. van der Goot leg.; NBCN • 1 ♂; Toscana, Florence, Careggi; 43°48'45"N, 11°15'07"E; 19 May 1986; NBCN • 2 ♂♂; Piedmont, Colle di Sestrieres; 44°57'00"N, 6°52'60"E; 1800/2100 m a.s.l.; 23-31 Jul. 1837; Zerny leg.; NHMW.

MONTENEGRO • 1 ♂; Lovćen, Lovćen 1; 42°22'59"N, 18°53'54"E; 17 May 2018; A. Vujić, A. Šebić, M. Ranković leg.; FSUNS 19017 • 1 ♂; Boka Kotorska, Morinj; 42°29'25"N, 18°38'56"E; 16-18 May 1998; FSUNS 03600 • 1 ♂; same data as for preceding; 18-19 May 1998; FSUNS 03601 • 2 ♀♀; same data as for preceding; FSUNS 03602, 03603.

Kazakhstan • 1 ♀; East Kazakhstan, Markakol’ District, 20 km N settlement Alekseevka, Souther slop of Matobaj Mountain range; 48°42'22"N, 85°57'00"E; 2318 m a.s.l.; 6 Jul. 1996; V. Zinchenko leg.; SZMN • 1 ♀; Kazakhstan, 9 km S settlement Karaoj, Kyzyl-Tass Mountain; 29 Jun. 1996; V. Zinchenko leg.; SZMN.

NORTH MACEDONIA • 1 ♂; Kožuf, Golema poljana; 41°10'54"N, 22°12'05"E; 15 Jun. 1955; FSUNS 00165 • 1 ♂; same data as for preceding; 18 Jun. 1956; FSUNS 00166 • 5 ♂♂; Kožuf, Golema poljana; 41°10'54"N, 22°12'05"E; 14 Jun. 1975; FSUNS 00154, 00156 to 00158, 00162 • 1 ♀; Kožuf, Golema poljana; 41°10'54"N, 22°12'05"E; 15 Jun. 1968; FSUNS 00167 • 2 ♀♀; same data as for preceding; 17 Jun. 1956; FSUNS 00168, 00169 • 6 ♀♀; Kožuf, Golema poljana; 41°10'54"N, 22°12'05"E; 14 Jun. 1975; FSUNS 00155, 00159 to 00161, 00163, 00164 • 2 ♀♀; Kožuf; 41°25'38"N, 21°30'45"E; 14 Jun. 1975; FSUNS 00152, 00153.

Russia • 1 ♂; Sarepta, "Russia, RUS, Sarepta, now a suburb of Volgograd city (Christoph)"; 48°31'40"N, 44°29'01"E; ZHMB 02501 • 4 ♀♀; same data as for preceding; ZHMB 02500, 02502, 02508, 02509 • 32 ♂♂; Altai, 10 km S-W of Katanda; 50°06'51"N, 86°07'21"E; 6 Jul. 1983; A. Barkalov leg.; SZMN • 9 ♀♀; same data as for preceding; SZMN • 1 ♂; SW Altai, Katun valley 10 km W Katanda; 50°09'46"N, 86°06'50"E; 7 Jul. 1983; H. Hippa leg.; MZH • 1 ♂; same data as for preceding; 22-27 Jun. 1983; MZH; "exp. Mikkola, Hippa et Jalava" • 1 ♂; Altai, Kurayskaya Step’; 50°12'00"N, 87°47'60"E; 1662 m a.s.l.; 9 Jul. 2006; A. Barkalov leg.; SZMN • 17 specimens; Altai, Terekta; 50°16'12"N, 85°58'12"E; 1098 m a.s.l.; Jun. 1973; SZMN • 1 ♂; Tuva, Erzin river; 50°19'12"N, 95°30'00"E; 1288 m a.s.l.; 27 Jun.-1. Jul. 1989; D. Logunov leg.; SZMN • 16 ♀♀; same data as for preceding; SZMN • 2 ♂♂; Tuva, Tere Khol’ Lake; 50°42'11"N, 97°20'2"E; 27 Jun.-1 Jul. 1989; D. Logunov leg.; SZMN • 1 specimen; Altai, Tuyekta; 50°51'00"N, 85°49'48"E; 944 m a.s.l.; Jun. 1979; SZMN • 2 specimens; Tuva, Chagytay; 50°58'30"N, 94°38'47"E; 1963; SZMN • 1 specimen; Altai, Baragash; 51°16'48"N, 85°12'36"E; 927 m a.s.l.; Jun. 1973; SZMN • 2 ♀♀; Altaj, Erlagol; 51°22'21"N, 86°05'22"E; 27 Jun. 1995; FSUNS 00171, 00173 • 3 ♂♂; Altaj, Erlagol; 51°22'23"N, 86°05'29"E; 27 Jun. 1995; A. Tepavčević leg.; FSUNS 00170, 00172, 02476 • 1 ♀; same data as for preceding; FSUNS 02477 • 1 ♂; Gornyi Altai, Turochaksky r-n kordon Obogo; 51°35'47"N, 87°05'45"E; 950 m a.s.l.; 15 Jun. 2003; D. Kropačeva leg.; SZMN 22631 • 18 ♂♂; Altai mountains, Teletskoye Lake; 51°41'20"N, 87°33'43"E; 23-25 Jun. 2013; A. Vujić, S. Radenković leg.; FSUNS NJ56, NJ57, NJ59 to NJ62, NJ64 to NJ71, NJ73 to NJ75, NJ77 • 5 ♀♀; same data as for preceding; FSUNS NJ63, NJ72, NJ76, NJ78, NJ79 • 2 ♂♂; Altai, Turochaksky r-n, Teletskoe Lake, 14 km S of Iogach; 51°42'0"N, 87°17'60"E; 598 m a.s.l.; 27 Jun. 2006; V. Zinchenko leg.; SZMN • 2 ♀♀; same data as for preceding; SZMN • 1 ♀; Mountain Ural, Orenburg; 51°46'12"N, 54°59'53"E; ZHMB 02519 • 15 ♂♂; Siberia, Altaya, Teletskoe Lake; 51°46'60"N, 87°18'00"E; 24-19 Jun. 2006; A. Barkalov, V. Zinchenko leg.; SZMN • 6 ♀♀; same data as for preceding; SZMN • 2 ♂♂; same data as for preceding; 27 Jun. 2006; J. T. Smit leg.; FSUNS 03972, 03973 • 1 ♀; same data as for preceding; 24 Jun. 2006; FSUNS 03974 • 1 ♂; Siberia, Republic Alatai, Teletskoe lake, Artybash.; 51°47'57"N, 87°14'58"E; 25 Jun. 1990; G. Ståhls leg.; MZH • 8 ♂♂; Altai, Teletskoe Lake, Artibash; 51°47'57"N, 87°14'58"E; 12 Jun. 1990; A. Barkalov, Čekanov leg.; SZMN • 1 ♀; same data as for preceding; SZMN • 17 ♂♂; same data as for preceding; 11-25 Jun. 1990; SZMN • 8 ♀♀; same data as for preceding; SZMN • 13 ♂♂; same data as for preceding; 18-20 Jun. 1990; SZMN • 10 ♀♀; same data as for preceding; SZMN • 1 ♀; same data as for preceding; 23 Jul. 1979; SZMN • 82 specimens; same data as for preceding; Jun. 1979; SZMN • 1 ♂; Altaj, Gorno-Altaysk; 51°57'08"N, 85°57'19"E; 21 Jun. 1983; MZH; "exp. Mikkola, Hippa et Jalava" • 152 specimens; Altai, Gorno-Altaysk; 51°57'08"N, 85°57'19"E; Jun.-Jul. 1979; A. Barkalov leg.; SZMN • 2 ♂♂; same data as for preceding; 22 Jun. 1983; SZMN • 3 ♀♀; same data as for preceding; SZMN • 1 ♂; Altai Republic; 52°30'00"N, 83°00'00"E; 25 Jun. 1979; NBCN • 1 ♀; same data as for preceding; NBCN • 2 ♂♂; Sayan Mountains, Abaza; 52°41'17"N, 90°05'27"E; 30 May-11 Jun. 1981; A. Barkalov, T. Varlamova leg.; SZMN • 7 specimens; same data as for preceding; Jun. 1969; SZMN • 2 specimens; Novosibirsk; 55°06'56"N, 82°51'33"E; 1972-1974; SZMN • 1 specimen; Tuva, Sosnovka; 56°18'18"N, 51°14'44"E; 1949; SZMN.

Spain • 1 ♂; Sierra Nevada, second valley; 37°06'10"N, 3°27'19"W; 1430 m a.s.l.; 17 Jun. 2014; A. Vujić, S. Radenković, C. Pérez-Bañón leg.; FSUNS 07410 • 3 ♂♂; Sierra Nevada, Ski Centar Sierra Nevada; 37°06'45"N, 3°25'10"W; 2190 m a.s.l.; 16 Jun. 2014; A. Vujić, S. Radenković, C. Pérez-Bañón leg.; FSUNS 07275, 07287, 07302 • 1 ♂; prov. Granada Sierra Nevada ri. Valetta; 37°06'55"N, 3°29'32"W; 1 Jun. 1982; NBCN • 11 ♂♂; Sierra Nevada, First valley; 37°07'40"N, 3°26'44"W; 1630 m a.s.l.; 17 Jun. 2014; A. Vujić, S. Radenković, C. Pérez-Bañón leg.; FSUNS 07325, 07326, 07328, 07338, 07342, 07344, 07358, 07368, 07371, 07373, 07384 • 10 ♀♀; same data as for preceding; FSUNS 07324, 07340, 07341, 07343, 07355, 07375, 07381, 07385, 07392, 07401 • 4 ♂♂; Sierra Nevada, road to hotel Duque; 37°08'17"N 3°25'46"W; 16 Jun. 2014; A. Vujić, S. Radenković, C. Pérez-Bañón leg.; FSUNS 07248, 07251, 07255, 07256 • 3 ♀♀; same data as for preceding; FSUNS 07262, 07263, 07265 • 1 ♂; Sierra Nevada Lugros, Horcajo del Camarate; 37°11'50"N, 3°15'13"W; 1370 m a.s.l.; 18 Jun. 2014; A. Vujić, S. Radenković, C. Pérez-Bañón leg.; FSUNS 07428 • 1 ♂; Andalusia, Sierra de Baza, Prados del Roy; 37°22'33"N, 2°51'06"W; 2000/2100 m a.s.l.; 9 Jun. 2003; D. Doczkal leg.; D. D. coll. 04805 • 1 ♀; same data as for preceding; D. D. coll. 04808 • 2 ♂♂; Andalusia, Sierra de Baza, Santa Barbara; 37°23'16"N, 2°50'43"W; 1890 m a.s.l.; 9 Jun. 2003; D. Doczkal leg.; D. D. coll. 04806, 04807 • 1 ♀; La Corte; 37°57'41"N, 6°49'09"W; 28 Apr. 2015; A. Vujić, D. Obreht leg.; FSUNS 09333 • 1 ♂; Alicante, Alcoy-Font Roja; 38°42'00"N, 00°28'00"W; 31 May 1994; P. M. Isidro leg.; FSUNS 02494 • 1 ♀; Valensija, Utiel; 39°34'13"N, 1°11'15"W; 9 May 1994; C. Pérez-Bañón leg.; FSUNS 02495 • 1 ♂; Val de Cabras; 40°09'23"N, 2°01'48"W; 10 Jun. 1980; H. G. M. Tenuissen leg.; NBCN • 1 ♂; between Leon and Oviedo, Puerto de Pajares; 43°00'00"N, 5°46'00"W; 12 Jul. 1972; NBCN.

Turkey • 5 ♂♂; "Kop Dağı geçidi” [Kop mountain pass], Bayburt; 40°15'00"N, 40°15'00"E; 16 Jul. 1992; NBCN.