Ptomaphaminus hanskii Schilthuizen & Perreau

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108: 88-89

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Ptomaphaminus hanskii Schilthuizen & Perreau

sp. n.

Ptomaphaminus hanskii Schilthuizen & Perreau  sp. n. Figure 11a, 12k, l, 13i, j


Holotype: Malaysia, Sarawak, 4th Division, Gunung Mulu National Park, Slope, 4.5.78, 9.1, 9.3, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, in NHMUK, B.M.1978-49), 1 male. Paratypes: Sarawak. Gunung Mulu National Park, Slope, 4.5.78, 9.1, 9.3, v–viii.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M.1978-49), 5 males, 8 females.


Length: 1.6-2.1 mm. General colour brown; the tarsi and two first antennomeres yellowish. Winged. Eyes well developed. Pronotum 1.65 times as wide as long. Elytra 1.30 times as long as wide. Sutural angle of female elytra angular. Apex of the spiculum gastrale of the male genital segment dilated into a diamond-like form (Figure 13i). Aedeagus elongated, approximately 3.5 times shorter than the body length. Median lobe regularly narrowed from base to the apex in dorsal view (Figure 12l), thick and strongly arcuate, with a long ventrally deflexed but not retroverted apical hook in lateral view (Figure 12k). Six lateroventral preapical setae and one lateroapical seta on each side. Internal stylus of the endophallus moderately sinuate. Female genital segment with reduced gonocoxites. Spermiduct membranous, vaguely helical at least near the base of the spermatheca. Apex of the spermatheca rounded, without apical sclerotised plate (Figure 13j).

Differential diagnosis.

Species with normally developed eyes and a long aedeagus. Very similar to P. sarawacensis  , from which it cannot be confidently distinguished in the female sex. Males of P. hanskii  have a thicker median lobe of the aedeagus than P. sarawacensis  .

Habitat and distribution.

Known from lowland forests of Gunung Mulu, Sarawak, Malaysia.


The external morphology and the female genital morphology are extremely similar to P. sarawacensis  , so that females are nearly impossible to distinguish. However, the male aedeagi of these species are very different.


Named in honour of Ilkka Hanski, the Finnish ecologist who played an important role in the Royal Geographical Society expedition to Mulu of 1978, and who passed away in 2016.