OXYPODINI

Leschen, Richard A. B. & Newton, Alfred F., 2015, Checklist and type designations of New Zealand Aleocharinae (Coleoptera: Staphylinidae), Zootaxa 4028 (3), pp. 301-353: 337-344

publication ID

http://dx.doi.org/10.11646/zootaxa.4028.3.1

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scientific name

OXYPODINI
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OXYPODINI  subtribe incertae sedis

Anocalea Fenyes 1921 

Anocalea Fenyes 1921: 27  . Type species: Anocalea thaxteri Fenyes, 1921  , by original designation. Notes: The genus Anocalea  includes 13 valid species from Chile, and was hitherto considered endemic to Chile, but one unidentified species has been reported from New Zealand.

Anocalea  undet. sp. (Klimaszewski et al. 1996). Distribution: New Zealand.

Austrocalea Cameron 1948 

Austrocalea Cameron 1948: 728  . Type species: Austrocalea lewisi Cameron, 1948  , by monotypy. Note: as synonym of Makara, Klimaszewski et al. (1996: 149)  .

Notes: The genus Austrocalea  includes only two species, both endemic to New Zealand.

Austrocalea brookesi Cameron 1950  Austrocalea brookesi Cameron 1950: 25  . Type locality: Auckland: Mt. Roskill. Type depository: NZACAbout NZAC LT, BMNH 1 PLT (designated below). Note: New Zealand, Kuschel (1990) (as "introduced", in error), Sivasubramaniam et al. (1997); as “ Austrocalea brookesi ( Cameron)  " in Klimaszewski et al. (1996: 149), implying not true Austrocalea  . Distribution: New Zealand (AK, MC).

Remarks. The number of specimens examined for naming this species was not indicated by Cameron. In order to

stabilize this name a lectotype and one paralectotype are here designated for two syntypes of Austrocalea 

brookesi  .

Type material examined. Lectotype ( NZACAbout NZAC): card-mounted (10 handwritten at base of card), “Coll. A. E.

Brookes. March. 1948. [in Brookes’ hand] // Mt. Roskill. Auckland. [in Brookes’ hand] // under garden rubbish [in

Brookes’ hand] // Austrocalea  brooksei Cam. TYPE [in Cameron’s hand, “ type ” in red ink] // A. E. Brookes

Collection // NZACAbout NZAC specimen database record 1007112 // NZ Arthropod Collection Private Bag 92170 Auckland

New Zealand NZACAbout NZAC 04065714 [barcode label]”. Paralectotype ( BMNH): card-mounted, “Para-type [round label

with yellow border] // N. Zealand Auckland M t. Roskill [in Cameron’s hand] // Austrocalea  brooksei Cam.

COTYPE [in Cameron’s hand, “ type ” written in red ink] // M. Cameron. Bequest. B.M. 1955 - 147.”.

Austrocalea lewisi Cameron 1948 

Austrocalea lewisi Cameron 1948: 729  . Type locality: Westland: Otira Gorge. Type depository: NZACAbout NZAC LT, 1 PLT, BMNH 1 PLT (designated below). Note: in genus Makara  in NZACAbout NZAC (nov.?) and implied in Klimaszewski et al. (1996: 149). Distribution: New Zealand (WD).

Remarks. Cameron did not indicate the number of specimens he based this species, but three were probably the

original number because on a card accommodating three specimens labeled as type in the NZACAbout NZAC, the right

specimen was removed and re-mounted separately, relabeled and and retained from the series by Cameron. In order

to stabilize this name a lectotype and 2 paralectotypes are here designated for these specimens of Austrocalea 

lewisi  .

Type material examined. Lectotype ( NZACAbout NZAC): card-mounted (left specimen, pink dot below it), “Coll. J. H. Lewis

1907. [in Brookes’ hand] // Otira Gorge. Westland [in Brookes’ hand] // Austrocalea lewisi Cam TYPE [in

Cameron’s hand, “ type ” in red ink] // Kept 1 [in Cameron’s hand] // A. E. Brookes Collection // NZACAbout NZAC specimen

database record 1007114 // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZACAbout NZAC 04065715

[barcode label]”. Paralectotypes: 1 ( NZACAbout NZAC), card-mounted (right specimen), same data as lectotype; 1 ( BMNH),

card-mounted, “Para-type [round label with yellow border] // N. Zealand Westland Otira Gorge [in Cameron’s

hand] // Austrocalea lewisi Cam.  COTYPE [in Cameron’s hand, “ type ” written in red ink] // M. Cameron. Bequest.

B.M. 1955 - 147.”.

Brouniana Blackwelder 1952 

Brouniana Blackwelder 1952: 86  new name for Brounia Blackwelder  (attributed to Cameron). Type species: Brounia vulcanica Cameron, 1945  , by autotypy.

= Brounia  Cameron 1945: 176 (nomen nudum, no type species designated; preoccupied, not Sharp 1878, Raffray 1898). Type species: none designated.

=Botromana Cameron 1948: 731 (new name for Brounia Cameron  ; nomen nudum, no type species designated). Type species: none designated.

= Brounia Blackwelder 1952: 85  (validation of Brounia Cameron  by designation of type species; preoccupied, not Sharp 1878, Raffray 1898). Type species: Brounia vulcanica Cameron, 1945  , by original designation.

Notes: Brouniana  includes only the following two species, endemic to New Zealand.

Brouniana lucida ( Cameron 1945) 

Brounia lucida Cameron 1945: 178  . Type locality: New Zealand: Rangitota [sic, Rangitoto]. Type depository: BMNH HT. Distribution: New Zealand (AK).

Remarks. Cameron based his description on a “unique” specimen that we confirmed as the holotype.

Type material examined. Holotype ( BMNH): card-mounted, “near- 169. [in Broun’s hand] // New Zealand. [red

underline] Broun Coll. Brit. Mus. 1922 - 482. // Rangitota. 28.10. 1912. [in Broun’s hand] // Brounia lucida Cam. 

TYPE [in Cameron’s hand, “ type ” written in red ink] // Holo-type [round label with red border].”.

Brouniana vulcanica ( Cameron 1945) 

Brounia vulcanica Cameron 1945: 177  (attributed to Broun in litt. (in Calodera  )). Type locality: New Zealand: Makara  ( LT). Type depository: BMNH LT, 4 PLT (designated below). Distribution: New Zealand (AK,WO?).

Remarks. Cameron did not explicitly state the numbers of specimens he based this species. In order to stabilize

this name a lectotype (in the Main Collection) and 4 paralectotypes (in the Broun Collection) are here designated

for specimens of Brouniana vulcanica  .

Type material examined. Lectotype ( BMNH): card-mounted (♂ printed at base of card), “Holo-type [round label

with red border] // Makara N. Zealand Hudson 1210  [in Cameron’s hand] // Brounia vulcanica Cam.  TYPE [in

Cameron’s hand, “ type ” written in red ink].” Paralectotypes ( BMNH): 1, card-mounted missing opart of the right

antenna), “Epsom. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Para-type

[round label with yellow border]”; 1, card-mounted missing opart of the right antenna), “n. sp. antenna [in Broun’s

hand] // Howick. Dec r 1885. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. //

Para-type [round label with yellow border]”; 1, card-mounted missing opart of the right antenna), “Hunua Maketu.

[in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Calodera vulcanica  [in

Broun’s hand] // Para-type [round label with yellow border].”; 1, card-mounted missing opart of the right antenna),

“Ngatira Rotorua. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Calodera 

vulcanica  [in Broun’s hand] // Para-type [round label with yellow border]”.

Colle Steel 1964 

Colle Steel 1964: 373  . Type species: Colle campbellensis Steel, 1964  , by original designation. Notes: Colle  includes a single species endemic to Campbell Island.

Colle campbellensis Steel 1964 

Colle campbellensis Steel 1964: 373  . Type locality: Campbell I.: Mt. Honey, 550 m, near summit. Type depository: NZACAbout NZAC HT, 10 PT; BPBMAbout BPBM, BMNH 7 PT. Distribution: Campbell I. (CA).

Gastrolamprusa Pace 1982 

Gastrolamprusa Pace 1982: 174  . Type species: Leptusa helmsi Bernhauer, 1941  , by original designation. Notes: Gastrolamprusa  includes a single species and is endemic to New Zealand.

Gastrolamprusa helmsi (Bernhauer 1941) 

Leptusa (Pasilia) Helmsi Bernhauer 1941: 38  . Type locality: "Neu-Seeland". Type depository: FMNHAbout FMNH HT ♀. Note: to Gastrolamprusa, Pace (1982)  . Distribution: New Zealand.

Remarks. Bernhauer based his description on a single specimen that we confirmed as the holotype.

Type material examined. Holotype female ( FMNHAbout FMNH): card-mounted, with mouthparts and spermatheca mounted in

balsam slide below, “Now. Zeeland Helms Reitter [bordered white label] // Helmsi  Brnh. Typus un. Leptusa  [in

Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // Gastrolamprusa  GEN. NOV. MIHI

helmsi (Bernhauer)  [in Pace's hand] det. R. Pace 1980 // HOLOTYPE teste D. J. Clarke 2014 GDI Imaging Project

[violet label] // Photographed Kelsey Keaton 2014 Emu Catalog [blue label] // FMNHINS 2819195 FIELD

MUSEUM [barcode label]”.

Makara Cameron 1948 

Makara Cameron 1948: 731  (new name for Maoria Cameron  ). Type species: Maoria hudsoni Cameron, 1945  , by autotypy.

= Maoria  Cameron 1945: 171 (preoccupied, not Laporte de Castelnau 1868, Pillsbry 1892, Warren 1912; also Franz 1975). Type species: Maoria hudsoni Cameron, 1945  , by monotypy.

=Malcama Blackwelder 1952: 230 (new name for Maoria Cameron  ). Type species: Maoria hudsoni Cameron, 1945  , by autotypy.

Notes: Makara  includes a single species and is endemic to New Zealand.

Makara hudsoni ( Cameron 1945) 

Maoria hudsoni Cameron 1945: 172  . Type locality: New Zealand: Mt. Aurum, Lake Makatipu [sic, =Wakatipu]. Type depository: BMNH LT, 3 PLT (designated below). Distribution: New Zealand (OL?).

Remarks. Cameron did not explicitly state the number of specimens on which he based this species. In order to

stabilize this name a lectotype and three paralectotypes are here designated for syntypes of Brounia vulcanica  .

Type material examined. Lectotype ( BMNH): card-mounted, “M t. Aurum N. Zealand (Hudson 1149) [in

Cameron’s hand] // Maoria hudsoni Brounia Cam  TYPE [in Cameron’s hand, “ type ” written in red ink] // M.

Cameron. Bequest. B.M. 1955 - 147.”. Paralectotypes ( BMNH): 1, card-mounted, “ NEWAbout NEW ZEALAND. Mt. Aurum.

Lake Wakatipu 300 ft. G. V. Hudson 1149. [in Hudson’s hand, partly printed] // M. Cameron. Bequest. B.M. 1955 -

147.”; 1, card-mounted, “M t. Aurum N. Zealand (Hudson 1149) [in Cameron’s hand] // Not in Broun coll. [in

Cameron’s hand] // M. Cameron. Bequest. B.M. 1955 - 147.”; 1, card-mounted (partly dissected), “M t. Aurum Lake

Wakatipu N. Zealand [in Cameron’s hand] // Maoria hudsoni Brounia Cam  [in Cameron’s hand] // M. Cameron.

Bequest. B.M. 1955 - 147.”.

Marecon Blackwelder 1952  STATUS NOV.

Marecon Blackwelder 1952: 230  (new name for Eurynotus Cameron  ). Type species: Gyrophaena rufipenne Broun, 1880  , by autotypy.

= Eurynotus  Cameron 1945: 170 (preoccupied, not Kirby 1817, Agassiz 1835, Scott 1898). Type species: Gyrophaena rufipenne Broun, 1880  , by original designation.

= Gyronotus  Cameron 1948: 731 (new name for Eurynotus Cameron  ; preoccupied, not Lansberge 1874). Type species: Gyrophaena rufipenne Broun, 1880  , by autotypy. SYN. NOV.

Notes: Marecon  is the only available name for this genus and must replace the preoccupied name Gyronotus  . Marecon  includes a single species and is endemic to New Zealand.

Marecon rufipennis (Broun 1880)  COMB. RESURR. (ex Gyronotus  )

Gyrophaena rufipenne Broun 1880: 92  . Type locality: Parua. Type depository: BMNH LT (designated below). Note: ex Calodera, Cameron 1945  ; as Gyronotus, Kuschel (1990: 46)  , Klimaszewski et al. (1996: 149), Frank & Ahn (2011: 28). Distribution: New Zealand (AK,ND).

Remarks. Broun based his description on two specimens, and one in BMNH was identified as a syntype. In order

to stabilize this name a lectotype is here designated for this specimen of Gyrophaena rufipenne  .

Type material examined. Lectotype ( BMNH): card-mounted (specimen with original card, recarded), “ Type

[round label with red border] // 170. [green label] // Parua. [in Broun’s hand] // New Zealand. [red underline] Broun

Coll. Brit. Mus. 1922 - 482. // Calodera  . rufipennis  [in Broun’s hand] // Eurynotus Cam.  [in Cameron’s hand]”.

Mecrona Blackwelder 1952 

Mecrona Blackwelder 1952: 232  (new name for Calonotus Cameron  ). Type species: Calodera algophila Broun, 1886  , by autotypy.

= Calonotus  Cameron 1945: 171 (preoccupied, not Agassiz 1846, Jan 1863). Type species: Calodera algophila Broun, 1886  , by original designation.

= Dasydera  Cameron 1948: 731 (new name for Calonotus Cameron  ; preoccupied, not LeConte 1861). Type species: Calodera algophila Broun, 1886  , by autotypy.

Notes: Mecrona  includes a single species and is endemic to New Zealand.

Mecrona algophila (Broun 1886) 

Calodera algophila Broun 1886: 941  . Type locality: Mokohinau Island. Type depository: BMNH LT, 1 PLT

(designated below). Note: as Dasydera, Frank & Ahn (2011: 28)  . Distribution: New Zealand (ND). Remarks. Broun did not mention the number of specimens he based this species on, and we located two syntypes in the BMNH. In order to stabilize this name a lectotype and one paralectotype are here designated for these specimens of Gyrophaena rufipenne  .

Type material examined. Lectotype ( BMNH): card-mounted, “ Type [round label with red border] // 1693. [in Broun’s hand] // Mokohinau // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Calodera algophila  [in Broun’s hand]”. Paralectotype ( BMNH): card-mounted, “ 1693. [in Broun’s hand] // Mokohinau // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // algophila  [in unknown hand]”.

Neodoxa Klimaszewski, Marris & Thorpe 2003 

Neodoxa Klimaszewski, Marris & Thorpe 2003: 119  (new name for Heterodoxa Cameron 1950  ). Type species: Heterodoxa secreta Cameron, 1950  , by autotypy.

= Heterodoxa  Cameron 1950: 25 (preoccupied, not Malloch 1932). Type species: Heterodoxa secreta Cameron, 1950  , by original designation.

Notes: Neodoxa  includes only two species and is endemic to New Zealand (including Three Kings Islands).

Neodoxa giachinoi Pace 2008 

Neodoxa giachinoi Pace 2008 b: 86  . Type locality: New Zealand, N. Island, 670m, Urewera N.P., Onepoto Cave (vaglio). Type depository: NZACAbout NZAC HT ♂, 1 PT ♀. Distribution: New Zealand (GB,TK).

Neodoxa secreta ( Cameron 1950) 

Heterodoxa secreta Cameron 1950: 25  . Type locality: Great King Island. Type depository: AMNZAbout AMNZ LT, 2 PLT (designated by Klimaszewski et al. 2003: 122). Distribution: New Zealand (TH).

Remarks. Cameron stated that the types are in the AMNZAbout AMNZ and co-types are in his collection. We located an

additional five syntypes in the BMNH and six syntypes in the NZACAbout NZAC that were not labelled as paralectotypes by

Klimaszewski et al. (2003) who fixed the name by lectotypy and labelled the specimen in the AMNZAbout AMNZ as a

lectotype. We affixed the following label to each of the BMNH and NZACAbout NZAC specimens: “ PARALECTOTYPE

Heterodoxa secreta Cameron 1950  , teste Leschen 2014 ”.

Neozelandusa Pace 2008

Neozelandusa Pace 2008 b: 88. Type species: Neozelandusa giachinoi  Pace, 2008, by original designation. Notes: Neozelandusa includes a single species and is endemic to New Zealand.

Neozelandusa giachinoi  Pace 2008

Neozelandusa giachinoi  Pace 2008 b: 89. Type locality: New Zealand, N. isl., 850m, Tongariro N.P., Erua for[est]. Type depository: NZACAbout NZAC HT ♂. Distribution: New Zealand (TO).

Paraphytopus Cameron 1948 

Paraphytopus Cameron 1948: 730  . Type species: Paraphytopus brookesi Cameron, 1948  , by monotypy. Notes: Paraphytopus  includes only two species, both endemic to New Zealand.

Paraphytopus brookesi Cameron 1948 

Paraphytopus brookesi Cameron 1948: 730  . Type locality: Canterbury: Bealy Flat. Type depository: NZACAbout NZAC LT, 2 PLT; BMNH 1 PLT (designated below). Distribution: New Zealand (MC).

Remarks. Cameron did not explicitly state the number of specimens on which he based this species. In order to

stabilize this name a lectotype and 3 paralectotypes are here designated for syntypes of Paraphytopus brookesi  .

Type material examined. Lectotype ( NZACAbout NZAC): card-mounted (far left specimen with pink dot underneath; “ 11 ”

handwritten in pencil at base of card), “Coll. E. S. Brookes 21 - 5-1945. [in Brookes’ hand] // Leaf-mould [in

Brookes’ hand] // Bealy Flat. Canterbury [in Brookes’ hand] // Paraphytopus brookesi Cam  TYPE [in Cameron’s

hand, “ type ” in red ink] // A. E. Brookes Collection // NZACAbout NZAC specimen database record 1007117 // NZ Arthropod

Collection Private Bag 92170 Auckland New Zealand NZACAbout NZAC 04065736 [barcode label]”. Paralectotypes: 2

( NZACAbout NZAC), card-mounted (middle specimen mounted dorsally and right specimen together with lectotype), same data

as lectotype; 1 ( BMNH), card-mounted (dorsally), “SYN-TYPE [round label with powder blue border] // Bealy

Flat, Canterbury [in Brookes’ hand] // Leaf-mould. [in Brookes’ hand] // Coll. E.S. Brookes. 21.5-1945. [in Brookes’ hand] // Paraphytophus brookesi Cam  COTYPE [in Cameron’s hand, “cotype” in red ink] // M. Cameron. Bequest. B.M. 1955 - 147.”.

Paraphytopus minutus Cameron 1948 

Paraphytopus minutus Cameron 1948: 731  . Type locality: Canterbury: Te Moaka [sic, =Te Moana], Geraldine.

Type depository: BMNH NZACAbout NZAC LT, BMNH 1 PLT (designated below). Distribution: New Zealand (SC). Remarks. Cameron did not explicitly state the number of specimens on which he based this species. In order to stabilize this name a lectotype and a paralectotype are here designated for syntypes of Paraphytopus minutus  . Type material examined. Lectotype ( NZACAbout NZAC): card-mounted (“ 23 ” handwritten in pencil at base of card), “Coll. E. S. Brookes 4 - 1-1946. [in Brookes’ hand] // Leaf-mould [in Brookes’ hand] // Te Moana, Geraldine. Canterbury [in Brookes’ hand] // Paraphytopus minutus Cam  TYPE [in Cameron’s hand, “ type ” in red ink] // A. E. Brookes Collection // NZACAbout NZAC specimen database record 1007116 // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZACAbout NZAC 04065735 [barcode label]”. Paralectotype ( BMNH): card-mounted (right antenna missing), “SYN-TYPE [round label with powder blue border] // Te Moana, Geraldine, Canterbury. [in Brookes’ hand] // Leaf-mould. [in Brookes’ hand] // Coll. E.S. Brookes. 7 - 1-1946. [in Brookes’ hand] // Paraphytophus minutus Cam  COTYPE [in Cameron’s hand, “cotype” in red ink] // M. Cameron. Bequest. B.M. 1955 - 147.”.

Polylobus Solier 1849 

Polylobus Solier 1849: 354  . Type species: Polylobus maculipennis Solier, 1849  , designated by Fenyes (1918: 24). Notes: Polylobus  contains 75 valid species from the Neotropical region, Australia and New Zealand (one endemic species).

Polylobus sternalis (Broun 1880) 

Gyrophaena sternale Broun 1880: 88  . Type locality: near Whangarei Harbour. Type depository: BMNH LT, 1 PLT (designated below). Distribution: New Zealand (ND).

Remarks. Broun indicated that two specimens were the basis for this species, and these were located in the

BMNH. In order to stabilize this name a lectotype and one paralectotype are here designated for these specimen of

Gyrophaena sternale  .

Type material examined. Lectotype ( BMNH): card-mounted (right specimen, left antenna missing), “ Type [round

label with red border] // 160. [green label] // Parua –Whangarei [in Broun’s hand] // New Zealand. [red underline]

Broun Coll. Brit. Mus. 1922 - 482. // Polylobus sternalis  [in Broun’s hand] // Gen. dub. [in Cameron’s hand]”.

Paralectotype ( BMNH): card-mounted (left specimen mounted dorsally with head missing), shared labels with

lectotype.

Sytus Blackwelder 1952 

Sytus Blackwelder 1952: 368  (new name for Dasynotus Broun  ). Type species: Dasynotus fulgens Broun, 1880  , by autotypy.

= Dasynotus  Broun 1880: 93 (preoccupied, not Wagler 1830). Type species: Dasynotus fulgens Broun, 1880  , designated by Fenyes (1918: 22).

subgenus Bunandrogaster Pace 1982: 176  . Type species: Leptusa curiosa Bernhauer, 1941  , by original designation.

Notes: The genus Sytus  is endemic to New Zealand, with seven species.

Sytus aerarius (Broun 1880) 

Dasynotus aerarius Broun 1880: 94  . Type locality: Tairua; Whangarei Harbour. Type depository: BMNH LT, 1 PLT (designated below). Note: DN in Nunn (2003); not Calodera  but not placed, record ex Auckland, Assing (2003). Distribution: New Zealand (AK,CL,ND,WA,WN; DN).

Remarks. Broun indicated that 6- 7 specimens were the basis for naming this species, two of which were located in

the BMNH. In order to stabilize this name a lectotype and one paralectotype are here designated for a specimen of

Dasynotus aerarius  .

Type material examined. Lectotype ( BMNH): card-mounted, “ 173. [green label] // Tairua // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Calodera  aeraria [in Broun’s hand]”. Paralectotype ( BMNH): cardmounted, “ Type [round label with red border] // 173. [green label] // Manaia. Whangarei. [in Broun’s hand] // aeraria [in unknown hand]”.

Sytus bifossuta ( Cameron 1945) 

Dasynotus bifossuta Cameron 1945: 170  (attributed to Broun in litt. (in Calodera  )). Type locality: New Zealand: Greymouth. Type depository: BMNH HT. Distribution: New Zealand (BR).

Remarks. Cameron based his description on a “unique” specimen that we confirmed as the holotype.

Type material examined. Holotype ( BMNH): card-mounted (most of the left antenna missing), “Holo-type [round

label with red border] // Greymouth. - Lewis - [in Broun’s hand] // Calodera  biffossuta [in Broun’s hand] //

Dasynotus bifossuta Cam  TYPE [in Cameron’s hand, “ type ” in red ink]”.

Sytus (Bunandrogaster) curiosus (Bernhauer 1941) 

Leptusa (Leptusa) curiosa Bernhauer 1941: 37  (attributed to in litt. (in Oxypoda  )). Type locality: Mt. Hutt. Type depository: FMNHAbout FMNH LT, BMNH 2 PLT (designated below). Note: to Sytus, Pace (1982)  . Distribution: New Zealand (MC).

Remarks. Bernhauer did not indicate the number of specimens he examined for naming this species, but one

syntype was located in the FMNHAbout FMNH and two in the BMNH were curated under the genus “ Brunandrogaster ”. In

order to stabilize this name, a lectotype ( FMNHAbout FMNH) and two paralectotypes ( BMNH) are here designated for these

specimens of Leptusa curiosa  . The FMNHAbout FMNH syntype was erroneously cited as "tipo unico" by Pace (1982).

Type material examined. Lectotype ( FMNHAbout FMNH): card-mounted (genitalia in balsam on acetate mount and

mouthparts on microslide), “ New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Mt. Hutt. 12.4. 1912

[in Bernhauer’s hand] // ♂ … [illegible, in Bernhauer’s hand] // Oxypoda curiosa Brh. Typ.  [in Bernhauer’s hand].

] Bnhauer det. // BRITAbout BRIT. MUS. DON. ARROW // curiosa Brnh.  Typus Leptusa  [in Bernhauer’s hand, red label] //

Chicago NHMus M.Bernhauer Collection // Dasynotus curiosus  ( BERNH.) det. R. Pace 1980 [in Pace’s hand,

partly printed] // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label] // Photographed Kelsey

Keaton 2014 Emu Catalog [blue label] // FMNHINS 2819290 FIELD MUSEUM [barcode label]”. Paralectotypes

( BMNH): 1, card-mounted, “ Type [round label with red border] // New Zealand. [red underline] Broun Coll. Brit.

Mus. 1922 - 482. // M t. Hutt. 12.4. 1912. [in Broun’s hand] // Oxypoda curiosa Bernh. Bernhauer  det. Typ. [in

Bernhauer’s hand, partly printed] // Homalota curiosa  – near – [in Broun’s hand, partly illegible]”; 1, card-

mounted, “M t. Hutt. 12.4. 1912. [in Broun’s hand] // Ischno. sp. [in Broun’s hand, red marked] // New Zealand. [red

underline] Broun Coll. Brit. Mus. 1922 - 482.”.

Sytus flavescens (Broun 1880) 

Dasynotus flavescens Broun 1880: 93  . Type locality: Parua [ LT, published as "Whangarei Harbour"]. Type depository: BMNH LT, 1 PLT (designated below). Note: DN in Nunn (2003). Distribution: New Zealand (AK,ND,TO; DN).

Remarks. Broun based this species on two specimens. In order to stabilize this name a lectotype and one

paralectotype are here designated for these syntypes of Dasynotus flavescens  .

Type material examined. Lectotype ( BMNH): card-mounted (part of the right antenna is missing), “ Type [round

label with red border] // 171. [green label] // Parua. [in Broun’s hand] // New Zealand. [red underline] Broun Coll.

Brit. Mus. 1922 - 482. // Calodera flavescens  . [in Broun’s hand]”. Paralectotype ( BMNH): card-mounted (some

appendages missing), “ 171. [green label] // Auck. [in Broun’s hand] // Calodera  [in Broun’s hand] // New Zealand.

[red underline] Broun Coll. Brit. Mus. 1922 - 482.”.

Sytus fulgens (Broun 1880) 

Dasynotus fulgens Broun 1880: 95  . Type locality: Parua, near Whangarei Harbour. Type depository: BMNH HT. Distribution: New Zealand (ND).

Remarks. Broun based his description on a single specimen that we confirmed as the holotype.

Type material examined. Holotype ( BMNH): Lectotype ( BMNH): card-mounted (head is mounted dorsally), “ Type [round label with red border] // 175. [green label] // Parua. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Calodera fulgens  . [in Broun’s hand]”.

Sytus granifer (Broun 1894) 

Calodera granifer Broun 1894: 381  . Type locality: Waitakerei Range. Type depository: BMNH HT. Note: as Sytus, Kuschel (1990)  . Distribution: New Zealand (AK,ND).

Remarks. Broun based his description on a single specimen that we confirmed as the holotype.

Type material examined. Holotype ( BMNH): card-mounted (left hind leg missing), “ Type [round label with red

border] // 2689. [in Broun’s hand] // Waitakerei // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. /

/ Calodera granifer  . [in Broun’s hand]”.

Sytus optabilis (Broun 1880) 

Dasynotus optabilis Broun 1880: 94  . Type locality: Manaia, Whangarei [ LT, originally published as near

Whangarei Harbour]. Type depository: BMNH HT. Distribution: New Zealand (ND). Remarks. Broun based his description on a single specimen that we confirmed as the holotype. Type material examined. Holotype ( BMNH): Lectotype ( BMNH): card-mounted, “ Type [round label with red border] // 172. [green label] // Manaia. Whangarei. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922 - 482. // Calodera optabilis  . [in Broun’s hand]”.

NZAC

New Zealand Arthropod Collection

BPBM

Bishop Museum

FMNH

Field Museum of Natural History

NEW

University of Newcastle

AMNZ

Auckland Institute and Museum

BRIT

Botanical Research Institute of Texas

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Loc

OXYPODINI

Leschen, Richard A. B. & Newton, Alfred F. 2015

2015
Loc

Dasydera

Frank & Ahn 2011: 28

2011
Loc

Neodoxa giachinoi

Pace 2008

2008
Loc

Neodoxa giachinoi

Pace 2008: 86

2008
Loc

giachinoi

Pace 2008: 89

2008
Loc

Neodoxa

Klimaszewski, Marris & Thorpe 2003: 119

2003
Loc

Makara

Klimaszewski et al. 1996: 149

1996
Loc

Gyronotus

Kuschel 1990: 46

1990
Loc

Sytus

Kuschel 1990

1990
Loc

Gastrolamprusa

Pace 1982

1982
Loc

Gastrolamprusa

Pace 1982: 174

1982
Loc

Gastrolamprusa

Pace 1982

1982
Loc

Bunandrogaster

Pace 1982: 176

1982
Loc

Sytus

Pace 1982

1982
Loc

Colle

Steel 1964: 373

1964
Loc

Colle campbellensis

Steel 1964

1964
Loc

Colle campbellensis

Steel 1964

1964
Loc

Colle campbellensis

Steel 1964: 373

1964
Loc

Brouniana

Blackwelder 1952: 86

1952
Loc

Brounia

Blackwelder 1952: 85

1952
Loc

Marecon

Blackwelder 1952: 230

1952
Loc

Mecrona

Blackwelder 1952: 232

1952
Loc

Sytus

Blackwelder 1952: 368

1952
Loc

Austrocalea brookesi

Cameron 1950

1950
Loc

Austrocalea brookesi

Cameron 1950: 25

1950
Loc

Heterodoxa

Cameron 1950

1950
Loc

Heterodoxa secreta

Cameron 1950

1950
Loc

Heterodoxa

Cameron 1950: 25

1950
Loc

Heterodoxa secreta

Cameron 1950

1950
Loc

Neodoxa secreta (

Cameron 1950

1950
Loc

Heterodoxa secreta

Cameron 1950: 25

1950
Loc

Austrocalea

Cameron 1948

1948
Loc

Austrocalea

Cameron 1948: 728

1948
Loc

Austrocalea lewisi

Cameron 1948

1948
Loc

Austrocalea lewisi

Cameron 1948: 729

1948
Loc

Makara

Cameron 1948: 731

1948
Loc

Gyronotus

Cameron 1948: 731

1948
Loc

Dasydera

Cameron 1948: 731

1948
Loc

Paraphytopus

Cameron 1948

1948
Loc

Paraphytopus

Cameron 1948: 730

1948
Loc

Paraphytopus brookesi

Cameron 1948

1948
Loc

Paraphytopus brookesi

Cameron 1948

1948
Loc

Paraphytopus brookesi

Cameron 1948: 730

1948
Loc

Paraphytopus minutus

Cameron 1948: 731

1948
Loc

Brounia vulcanica

Cameron 1945

1945
Loc

Brounia

Cameron 1945: 176

1945
Loc

Brounia vulcanica

Cameron 1945

1945
Loc

Brouniana lucida (

Cameron 1945

1945
Loc

Brounia lucida

Cameron 1945: 178

1945
Loc

Brounia vulcanica

Cameron 1945: 177

1945
Loc

Maoria hudsoni

Cameron 1945

1945
Loc

Maoria

Cameron 1945: 171

1945
Loc

Maoria hudsoni

Cameron 1945

1945
Loc

Maoria hudsoni

Cameron 1945

1945
Loc

Makara hudsoni (

Cameron 1945

1945
Loc

Maoria hudsoni

Cameron 1945: 172

1945
Loc

Eurynotus

Cameron 1945: 170

1945
Loc

Calodera

Cameron 1945

1945
Loc

Calonotus

Cameron 1945: 171

1945
Loc

Dasynotus bifossuta

Cameron 1945: 170

1945
Loc

Leptusa helmsi

Bernhauer 1941

1941
Loc

Gastrolamprusa helmsi

Bernhauer 1941

1941
Loc

Leptusa (Pasilia)

Helmsi Bernhauer 1941: 38

1941
Loc

Leptusa curiosa

Bernhauer 1941

1941
Loc

Leptusa (Leptusa) curiosa

Bernhauer 1941: 37

1941
Loc

Anocalea

Fenyes 1921: 27

1921
Loc

Anocalea thaxteri

Fenyes 1921

1921
Loc

Calodera granifer

Broun 1894: 381

1894
Loc

Calodera algophila

Broun 1886

1886
Loc

Calodera algophila

Broun 1886

1886
Loc

Calodera algophila

Broun 1886

1886
Loc

Mecrona algophila

Broun 1886

1886
Loc

Calodera algophila

Broun 1886: 941

1886
Loc

Gyrophaena rufipenne

Broun 1880

1880
Loc

Gyrophaena rufipenne

Broun 1880

1880
Loc

Gyrophaena rufipenne

Broun 1880

1880
Loc

Marecon rufipennis

Broun 1880

1880
Loc

Gyrophaena rufipenne

Broun 1880: 92

1880
Loc

Polylobus sternalis

Broun 1880

1880
Loc

Gyrophaena sternale

Broun 1880: 88

1880
Loc

Dasynotus fulgens

Broun 1880

1880
Loc

Dasynotus

Broun 1880: 93

1880
Loc

Dasynotus fulgens

Broun 1880

1880
Loc

Sytus aerarius

Broun 1880

1880
Loc

Dasynotus aerarius

Broun 1880: 94

1880
Loc

Dasynotus flavescens

Broun 1880: 93

1880
Loc

Dasynotus fulgens

Broun 1880: 95

1880
Loc

Dasynotus optabilis

Broun 1880: 94

1880
Loc

Polylobus

Solier 1849: 354

1849
Loc

Polylobus maculipennis

Solier 1849

1849