Eoclivina basilewskyi, Balkenohl, 2023

Balkenohl, Michael, 2023, The genus Eoclivina (KULT, 1959) from Africa with the description of two new species (Coleoptera, Carabidae, Clivinini), Belgian Journal of Entomology 138, pp. 1-17 : 10-12

Eoclivina basilewskyi sp. nov.

( Figs 2 , 7 , 11 )

urn:lsid:zoobank.org:act:5D13B28C-52D3-4338-87FE-B68FCEBCA35C

ETYMOLOGY. The name is a patronym in honour of the Belgian entomologist Pierre Basilewsky, curator of the Musée Royal de l´Afrique Centrale, Tervuren, Belgium, specialist in Carabidae and especially in Scaritinae , who published his results in 404 contributions during 1929 and 1992 ( REIGNIEZ & ANDRÉ, 1994).

TYPE MATERIAL. Holotype: ♂, with white black printed label and data: “Coll. Mus. Tervuren Nigeria: Ile-Ife J.T. Medler ” ( Fig. 2 ) ( MRACT).

Paratypes: 1 ♂, 6 ♀♀, 1 spec., with blue black printed label and data: “ NIGERIA: Ile-Ife 7.VIII.1986 leg.: F.-T. KRELL” / white, black printed “ Clivina SG. Eoclivina dumolinii PUTZ. det. Balkenohl 94” ( CMBB); 6 specs, same data as before ( SMNS); 1 ♂, 4 ♀♀, 4 specs, “Coll. Mus. Tervuren Nigeria: Ile-Ife J.T. Medler” ( MRACT).

REMARK. The majority of the paratypes from MRACT are more or less rotten inside.

DIAGNOSIS.A small sized fuscous species with long-oval elytra with straight and slightly diverging part of the lateral margins, all intervals of the elytron smooth, and with the apex of the elytron more or less pale fulvous coloured. Distinguished from the most similar species E. dumolinii by the completely reticulated abdominal sternite (in both of the sexes), the less convex eye, the different shape of the male aedeagus and the different shape of the female gonocoxite.

DESCRIPTION. Measurements. Holotype: Body length 4.15 mm, width 1.21 mm; ratio L/W of pronotum 0.96; ratio L/W of elytra 1.86. Paratypes: Body length 3.84–4.18 mm (x = 4.04 mm *), width 1.12–1.21 mm (x = 1.16 mm *), ratio L/W of pronotum 0.95–1.01 (x = 0.97*), ratio L/W of elytra 1.85–1.9 (x = 1.88*); (*n = 10).

Colour: Glossy. Dark fuscous. Mouthparts, antenna, intermediate and hind leg fuscous. Supraantennal plate translucent-fuscous. Elytron with the tip of apex pale fuscous.

Head: About a quarter narrower than pronotum. Clypeus with central part bilobed anteriorly; lateral lobe slightly projecting, wing small, separated from clypeus by obtuse notch, separated from supraantennal plate by sharp notch. Supraantennal plate wide, laterally regularly convex and somewhat less projecting as eye; clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate overlapping eye latero-anteriorly less than a fifth (dorsal view), extended posteriorly into a keel, separated from wing by a slight furrow. Clypeus transverse, convex in frontal view, concave in lateral view, separated from frons by distinct sharp carina in form of a flat inverted V. Frons convex, with a triangle-like impressions at middle, with supraorbital carina extending from anterior to posterior eye-level. Clypeus and frons smooth, separated from supraantennal plate and supraorbital carina by deep moderately wide furrow, with clypeal seta in front; with two supraorbital setigerous punctures at middle and posterior end of the eye. Neck constriction missing. Eye moderately convex in dorsal view. Main part of eye situated ventrally where it is globose. Due to the overlapping supraantennal plate and the gena, the eye resembles in lateral view contracted at middle. Gena distinct, moderately convex, covering a quarter of posterior eye in ventral view. Antenna moderately short, reaching middle of pronotum. Labrum nearly straight anteriorly, slightly trilobed, with transverse to irregular reticulation, six setose. Mentum with base isodiametrically reticulated, with lateral lobe moderately hollowed out, shape of lobe asymmetric, regularly convex laterally, with irregular surface. Median tooth shorter than lateral lobe, wide and hollowed out at apex, with short carina at base.

Pronotum: Disk moderately convex in lateral view, distinctly convex in frontal view with increasing convexity laterally. Indistinctly wider than long, peltate. Reflexed lateral margin smooth, attenuating in anterior half with nearly straight part, widest behind middle; anterior angle distinct, slightly projecting, rounded off, posterior angle missing; lateral channel moderately narrow, completely and regularly rounded off at posterior setigerous puncture and continuing up to base. Median line distinctly engraved, somewhat punctured, complete. Anterior transverse line consisting of longitudinal connected punctures, joining median line, not joining anterior margin. Surface almost smooth, with few micro punctures, with circular group of medium sized punctures baso-laterally, with paramedian longitudinal group of punctures in basal half, with a few transverse wrinkles. Base narrow but distinct, three times as wide as lateral channel. Proepisternum not visible in dorsal view.

Elytron: Disk slightly to moderately convex in lateral view, distinctly and regularly convex in frontal view. Shape long-oval with nearly straight and diverging part in anterior half, with maximum width slightly behind middle. Humerus distinct. Reflexed lateral margin smooth. The row of setigerous punctures in the lateral channel is interrupted posterior of the humerus by a small gap and at middle by a wider gap. Scutellar striole missing; setigerous tubercle at base of first stria, with small tubercle at base of third interval, basal declivity with isodiametric reticulation. Humeral tooth small, situated at base of fourth interval. Striae moderately deep, distinctly and regularly punctuate-striate, one to three free at base, four to six reaching humerus, one and two indistinctly joining at apex, three and four, and five and six distinctly joining apically. Intervals convex, more convex laterally, interval eight narrow, carinate at apex. Surface of intervals glossy, interval eight smooth, intervals smooth at apex.

Hind wing: Fully developed.

Lower surface: Proepisternum with lateral two-thirds distinctly isodiametrically reticulated, with dense transverse wrinkles. Sternites of abdomen in both sexes completely covered with isodiametric reticulation, opaque. The two marginal setigerous punctures are widely separated.

Legs: Protibia covered with distinct longitudinal reticulation, with three spines of moderate length, not sulcate dorsally, movable spur as long as apical spine, slightly arcuate in apical third. Intermediate tibia with five tubercles at the inner side and seven at the lateral side, tubercles furnished with strong setae.

Male genitalia ( Fig. 7 ): Median lobe relatively long, slender, moderately arcuate, flattened laterally, with asymmetric spatula. Endophallus with two groups of bristles.

Female gonocoxite and epipleurite ( Fig. 11 ): Gonocoxite foliform, elongated, completely flattened, with five robust nematiform setae positioned at the median side, apex asetose. Epipleurite long quadrangular, asetose.

Sexual dimorphism at the integument: Not observed.

VARIATION. In the eighteen specimens investigated, the clypeal wing varies in size. On the elytron, stria one and two are joining at apex in some specimens, in others they are ending free. The pale coloration at the apex of the elytron varies considerably in colour and extend.

DISTRIBUTION. Known from Ile-Ifé in Nigeria.

Figures

Figs 1-5. Eoclivina species, habitus, dorsal view. 1, E. dumolinii (Putzeys, 1846), ♂, specimen from Velingara, Guinea. 2, E. basilewskyi sp. nov., holotype. 3, E. machadoi (Basilewsky, 1955), holotype. 4, E. burgeoni (Kult, 1959), holotype. 5, E. schmidli sp. nov., holotype.

Figs 6-9. Eoclivina species, male genitalia. A-B, aedeagus. A, dorsal view. B, lateral view. C-D, parameres, dorsal view. 6, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 7, E. basilewskyi sp. nov., holotype. 8, E. machadoi (Basilewsky, 1955), paratype from Riv. Chiumbe, Angola. 9, E. schmidli sp. nov., holotype.

Figs 10-14. Eoclivina species, females, gonocoxite and epipleurite, ventro-lateral view. 10, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 11, E. basilewskyi sp. nov., paratype. 12, E. machadoi (Basilewsky, 1955), paratype. 13, E. burgeoni (Kult, 1959), holotype. 14, E. schmidli sp. nov., paratype.