Orsunius undetermined, ASSING 2011, ASSING, 2011

Assing, V., 2014, On Orsunius II. Eight new species and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina), Linzer biologische Beiträge 46 (1), pp. 461-479 : 476-478

Updated key to species

1 Pronotum with dense, finely granulose punctation, matt ( Figs 27 , 37, 41 ). Median excision of anterior margin of labrum broadly V-shaped. The O. granulosus group..........2

- Punctation of pronotum not granulose, less dense. Median excision of anterior margin of labrum in most species deeper, narrower, and U-shaped................................................4

2 Aedeagus with apically very acute ventral process in ventral view; sclerotized internal structures not sickle-shaped in ventral view ( Figs 38-39 ). Posterior excision of male sternite VIII small and semi-circular ( Figs 45-46 ). Laos ( Map 2 )......... O. cuneatus nov.sp.

- Apex of ventral process of aedeagus less acute in ventral view; sclerotized internal structures sickle-shaped in ventral view. Posterior excision of male sternite VIII larger, either of trapezoid shape or broadly semi-circular..............................................................3

3 Sclerotized internal structures of the aedeagus broad-based and apically slender ( Figs 42-43 ). Posterior excision of male sternite VIII broadly semi-circular ( Figs 47-48 ). Malaysia: Kedah ( Map 2 )........................................................................... O. heissi nov.sp.

- Sclerotized internal structures of the aedeagus of different shape, not distinctly broadbased ( Figs 31-32 ). Posterior excision of male sternite VIII trapezoid ( Figs 29-30 ). Laos, Thailand, North Vietnam ( Map 2 ) ........................................... O. granulosus nov.sp.

4 Protarsomeres I-IV not, or only weakly, dilated.................................................................5

- Protarsomeres I-IV distinctly dilated..................................................................................7

5 Head with conspicuously coarse and dense punctation ( Fig. 19 ). Male sternite VIII with broad and shallow posterior excision ( Fig. 33 ). Aedeagus as in Figs 20-21 . Thailand ( Map 2 ) ................................................................................ O. latexcisus nov.sp.

- Head with finer punctation. Male sexual characters different.............................................6

6 Eyes shorter than postocular region in dorsal view ( Fig. 2 ). Head with uniformly fine punctation ( Fig. 2 ). Male sternite VIII with very shallow posterior excision ( Figs 3-4 ). Aedeagus as in Figs 5-6 . Phippines: Mindanao ( Map 2 ).................. O. spathulatus nov.sp.

- Eyes approximately as long as postocular region in dorsal view ( ASSING 2011: figure 41). Head with double punctation, i.e., with micropunctation and with interspersed somewhat larger punctures ( ASSING 2011: figure 42). Male sternite VIII as in ASSING (2011: figure 45). Aedeagus as in ASSING (2011: figures 47-49). India, Nepal ( Map 1 ) ...... ......................................................................................................... O. parvexcisus ASSING

7 Head with conspicuously dense punctation everywhere, without shiny patch in median or posterior dorsal portion ( Fig. 11 ; ASSING 2011: figures 20-21)).....................................8

- Head with sparse punctation, or densely punctate with a shiny (impunctate or less densely punctate) patch in median or posterior dorsal portion............................................9

8 Head dark-brown, distinctly darker than the reddish pronotum; elytra more or less distinctly bicoloured ( Fig. 11 ). Male sternite VIII with rather narrow, nearly V-shaped posterior excision ( Fig. 8 ). Aedeagus as in Figs 12-13 . North Vietnam, Thailand ( Map 2 ) .......................................................................................................... O. stimulans nov.sp.

- Head reddish, or of similar coloration as pronotum; elytra reddish, not distinctly bicoloured ( ASSING 2011: figure 20). Male sternite VIII broadly V-shaped ( ASSING 2011: figure 24). Aedeagus as illustrated in ASSING (2011: figures 25-26). Taiwan ( Map 1 ).................................................................................................... O. excisus ASSING

9 Anterior and lateral dorsal portions of head with dense and relatively coarse punctation, with the interstices distinctly narrower than the diameter of the punctures...10

- Lateral and anterior dorsal portion with sparser and/or finer punctation...........................12

10 Punctation of head and pronotum slightly less dense ( Fig. 15 ). Body slightly smaller and more slender ( Fig. 14 ). Male sternite VIII with very broadly and shallowly concave posterior margin, without distinct posterior excision ( Fig. 9 ). Aedeagus as in Figs 16-17 . Thailand ( Map 2 ) ............................................................... O. gladiator nov.sp.

- Punctation of head and pronotum somewhat denser ( Assing 2011: figures 2-3, 9, 16). Body somewhat larger and more robust ( Assing 2011: figures 1, 15). Male sexual characters different............................................................................................................11

11 Head and pronotum reddish to dark-brown; elytra dark-brown, with the anterior margins, the suture, and the posterior margins reddish, rarely uniformly reddish; abdomen reddish to reddish-brown, with the middle of tergite VII infuscate. Male sternite VIII as in ASSING (2011: figure 17). Aedeagus as in ASSING (2011: figures 18- 19). China: Yunnan ( Map 1 ) ............................................................... O. yunnanus ASSING

- Head and pronotum reddish; elytra reddish-yellow to reddish, with the anterior half often slightly and diffusely darker; abdomen reddish to dark-reddish, tergite VII not infuscate. Male sternite VIII as in ASSING (2011: figure 14). Aedeagus as in ASSING (2011: figures 10-11). Thailand ( Map 1 )............................................. O. wunderlei ASSING

12 Head, pronotum, and abdomen dark-brown; elytra distinctly bicoloured, with dark-brown middle and with yellowish anterior and posterior margins ( Figs 22-23 ). Male sternite VIII with small and shallow posterior excision ( Figs 34-35 ). Aedeagus as in Figs 24-25 . Thailand ( Map 2 ) ........................................................ O. flavovirgatus nov.sp.

- Body, except for the sometimes bicoloured elytra, reddish. Male sexual characters different.............................................................................................................................13

13. Pronotum small in relation to head ( ASSING 2011: figure 28), at most as broad as head. Punctation of head and pronotum very fine ( ASSING 2011: figures 29, 35). Gular sutures very narrowly separated ( ASSING 2011: figure 34). Male sternite VIII with very shallow posterior excision ( ASSING 2011: figure 39). Aedeagus as in ASSING (2011: figures 36-37). North India, Malaysia, Thailand, Laos ( Map 1 ) ..... O. immsi (BERNHAUER)

- Pronotum larger in relation to head. Punctation of head and pronotum coarser. Gular sutures more broadly separated. Male primary and secondary sexual characters different................................................................................................................................. 14

14 Eyes larger, approximately as long as postocular region of head in dorsal view. Protarsomeres I-IV moderately dilated. Male sternite VIII less oblong and posteriorly with pubescence ( ASSING 2011: figure 52). Aedeagus as in ASSING (2011: figures 55- 56). Thailand ( Map 1 ) .............................................................................. O. pravus ASSING

- Eyes smaller, distinctly shorter than postocular region in dorsal view. Protarsomeres I-IV strongly dilated. Male sternite VIII more oblong and posteriorly without pubescence ( ASSING 2011: figure 66). Ventral process of aedeagus of completely different shape, not distinctly asymmetric in dorsal view ( ASSING 2011: figures 67-69); internal sac with distinctly sclerotized apical structures of characteristic shape. Indonesia: Bali ( Map 1 ) ................................................................... O. prolongatus ASSING

Figures

Map 1: Distributions of Orsunius immsi (white circles), O. parvexcisus (black diamonds), O. yunnanus (white triangles), O. pravus (black triangles), O. wunderlei (white star), O. excisus (black square), and O. prolongatus (black star).

Map 2: Distributions of Orsunius gladiator (white and black circles), O. latexcisus (black circle), O. flavovirgatus (black circle), O. heissi (black star), O. stimulans (black and grey triangles), O. granulosus (white and grey triangles), O. cuneatus (white diamond), and O. spathulatus (black square).

Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm.

Figs 36-43: Orsunius cuneatus nov.sp. (36-39) and O. heissi nov.sp. (40-43): (36, 40) habitus; (37, 41) forebody; (38, 42) aedeagus in lateral view; (39, 43) aedeagus in ventral view. Scale bars: 36- 37, 40-41: 1.0 mm; 38-39, 42-43: 0.2 mm.

Figs 44-48: Orsunius cuneatus nov.sp. (44-46) and O. heissi nov.sp. (47-48): (44) male sternite VII; (45, 47) male sternite VIII; (46, 48) posterior portion of male sternite VIII. Scale bars: 44-45, 47: 0.2 mm; 46, 48: 0.1 mm.

Figs 18-25: Orsunius latexcisus nov.sp. (18-21) and O. flavovirgatus nov.sp. (22-25): (18, 22) habitus; (19, 23) forebody; (20, 24) aedeagus in lateral view; (21, 25) aedeagus in ventral view. Scale bars: 18-19, 22-23: 1.0 mm; 20-21, 24-25: 0.2 mm.

Figs 1-9: Orsunius spathulatus nov.sp. (1-6), O. stimulans nov.sp. (7-8), and O. gladiator nov.sp. (9): (1) habitus; (2) forebody; (3, 8, 9) male sternite VIII; (4) posterior portion of male sternite VIII; (5) aedeagus in lateral view; (6) ventral process in ventral view; (7) male sternite VII. Scale bars: 1- 2: 1.0 mm; 3-9: 0.2 mm.

Figs 10-17: Orsunius stimulans nov.sp. (10-13) and O. gladiator nov.sp. (14-17): (10, 14) habitus; (11, 15) forebody; (12, 16) aedeagus in lateral view; (13) aedeagus in ventral view; (17) ventral process in ventral view. Scale bars: 10-11, 14-15: 1.0 mm; 12-13, 16-17: 0.2 mm.