Anthenoides granulosus Fisher 1913

Mah, Christopher L., 2018, New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean, Zootaxa 4539 (1), pp. 1-116 : 7-9

Anthenoides granulosus Fisher 1913

Figure 1 A–F

Fisher 1913: 647; Fisher 1919: 333, Pl. 88, fig. 2,3, pl. 94, figs. 4,4a–b; Döderlein 1924: 65; Fell 1958a: 12, pl. 1, fig. B; Fell 1958b: 135 (included in key); Clark H.E.S. 1970: 3; Baker & H.E.S. Clark 1970: 4; Rowe 1989: 289; A.M. Clark 1993: 242; Rowe & Gates 1995: 63; H.E.S. Clark & McKnight 2001: 20 (as A. granulosus )

Fisher 1913: 648; 1919: 338, pls 85(4), 88(4), 90(1), 94(5a,5b); Rowe 1989: 289; Rowe & Gates 1996: 63; A.M. Clark 1993: 242. (as A. rugulosus )

Diagnosis. This species is distinguished by the absence of spines on the inferomarginal plates ( Fig. 1D ) and the presence of variably fine to coarse/small to large granulation on superomarginals. Furrow spines, thick six to eight

( Figs 1 E–F). A duck- bill or straight pedicellariae present abactinally ( Fig. 1C ) and on the adambulacral plates ( Fig. 1E ) on the proximal side of the adambulacral plate.

Comments. Following the synonymy outlined by Clark & McKnight (2001: 23), Anthenoides rugulosus is accepted as a synonym of A. granulosus . Superomarginal width and length are variable and do not provide the taxonomic boundary established by Fisher (1919).

Occurrence: Philippine Islands, Moluccas, New South Wales, Australia, Norfolk Island (near Raoul Island, Kermadec Islands and New Zealand, Madagascar. 143– 731 m.

Material Examined. IE-2007-1040 Between Nosy-bé et Banc du Leven, Madagascar 12°44’S 48°12’E, 442– 491 m, Coll. Bouchet et al . MIRIKY. 2 wet specs. R=6.9, r=2.6; R=7.1, r=2.9; IE-2007-1228 Madagascar, 12°38’S 48°14’E, 436 m, Coll MIRIKY Bouchet et al. 1 wet spec. R=7.6, r=3.3 ; IE-2013-6625 . Madagascar 12°39.8S 48°14.2’E, 395–405 m Coll. A. Crosnier, 14 April 1971. 1 dry spec. R=~12.1, r=5.3, arms upturned ; IE-2007-1052 Between Nosy-bé et Banc du LevenMadagascar 12°4’S 48°11’E 488 m. Coll. Bouchet et al . MIRIKY. 1 wet spec. R=8.9, r=4.3.

Figures

FIGURE 1. Anthenoides granulosus Specimen IE-2013-6625. R=12.1, r=5.3 cm. A. Abactinal surface. Scale=3.0 cm. B. Closeup of abactinal surface and superomarginal plates. Scale=1.0 cm. C. Pedicellariae on abactinal surface. Scale=0.2 cm. D. Actinal surface. Scale=3.0 cm E. Furrow spines and adambulacral pedicellariae. Scale=1.0 cm. F. Oral region and furrow spines. Scale=1.0 cm.