Maechidiini, Telnov, 2020

Telnov, Dmitry, 2020, A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line, European Journal of Taxonomy 721, pp. 1-210 : 130-137

publication ID

https://doi.org/ 10.5852/ejt.2020.721.1127

publication LSID

lsid:zoobank.org:pub:89E62EF8-2E45-4C59-94B7-6A5603E8939B

persistent identifier

https://treatment.plazi.org/id/1A3787F6-9D9D-92D3-FF2E-F9DBCD212F8B

treatment provided by

Valdenar

scientific name

Maechidiini
status

trib. nov.

Key to the Maechidiini View in CoL from the Indo-Australian transition zone and Bali

This key is generally adapted for males and is mainly based on male features, including genital organs

.

1. Punctures of elytral disc ordinary: circular, variably ovoid or hexagonal, annular or not but not elongate linear or sinusoid incision-shaped..................................................................................... 31

– Punctures of elytral disc at least in part conspicuous: elongate linear or sinusoid, incision-shaped (cf. Figs 288, 290–291, 296 View Figs 286–301 , 303, 314 View Figs 302–316 , 317–320, 323, 328, 331 View Figs 317–332 , 339–340, 343, 348 View Figs 333–348 , 351, 353, 355 View Figs 349–358 , etc.) .................................................................................................................................................... 2

2. Lateral margins of pronotum in dorsal view distinctly rounded (more or less distinctly constricted anteriad and posteriad), without or with prebasal emargination or constriction ............................... 6

– Lateral margins of pronotum in dorsal view slightly sinuous, somewhat widened from anterolateral angles towards base or nearly straight in basal half, not distinctly constricted both anteriad and posteriad ( Figs 218 View Figs 208–225 , 226, 241 View Figs 226–241 ) ........................................................................................................... 3

3. Dorsal punctures of forebody comparatively large, distinctly annular; aedeagus not as in Figs 646– 648 View Figs 632–651 ..................................................................................................................................................... 4

– Dorsal punctures of forebody comparatively less large, not annular ................................................ 5

4. Lateral margin of pronotum with about 20 flat crenulae; elytron without tracks of longitudinal carinae; intervening spaces of elytra not transversely wrinkled; aedeagus unknown ............................................................................................. M. parallelicollis Moser, 1920 View in CoL

– Lateral margin of pronotum with about 19–20 flat crenulae; elytron with vague tracks of longitudinal carinae; intervening spaces of elytra transversely wrinkled; aedeagus as in Figs 646– 648 View Figs 632–651 ............................................................................................................ M. humeralis Heller, 1914 View in CoL

– Lateral margin of pronotum with about 24–25 crenulae; elytron with vague tracks of three longitudinal carinae; intervening spaces of elytra transversely wrinkled; aedeagus as in Narakusumo & Balke (2019: fig. 1e–f) ................................................ M. acutus ( Narakusumo & Balke, 2019) View in CoL comb. nov.

5. Anterolateral angles of labroclypeus broadly obtuse, not or slightly produced anteriad ( Fig. 135 View Figs 133–144 ); elytral setae very inconspicuous, strongly appressed; elytra without metallic shine; aedeagus as in Figs 646–648 View Figs 632–651 .................................................................................... M. lineatopunctatus Frey, 1969 View in CoL

– Anterolateral angles of labroclypeus subacute, strongly produced anteriad ( Fig. 120 View Figs 109–120 ); some elytral setae are longer and stouter than others, suberect, arranged in longidutinal rows; elytra with vague metallic shine; aedeagus not studied................................................................ M. gressitti Frey, 1969 View in CoL

6. Lateral margins of pronotum crenulate (rarely crenulate-denticulate); male labroclypeus broadly emarginate to subtruncate on anterior margin ................................................................................... 7

– Lateral margins of pronotum distinctly denticulate ( Fig. 199 View Figs 193–207 ); anterior margin of male labroclypeus deeply U-shaped emarginate on anterior margin, its anterolateral angles strongly protruding anteriad ( Fig. 99 View Figs 97–108 ) .......................................................................................................... M. alesbezdeki View in CoL sp. nov.

7. At least male metatibia distinctly thickened distally, not long and slender; all tarsi with short to moderately long tarsomeres; aedeagus not as in Figs 652–654 View Figs 652–666 ......................................................... 8

– Male metatibia slender and hardly widened distally ( Fig. 55 View Figs 54–57 ); all tarsomeres slender and lengthened ( Fig. 55 View Figs 54–57 ); aedeagus as in Figs 652–654 View Figs 652–666 ............................................................... M. longipes View in CoL sp. nov.

8. Setae on lateral margins of pronotum and elytra shorter than length of compound eye (in older specimens setae often become broken and look shorter); aedeagus not as in Figs 614–616 View Figs 601–616 or 703–705 ........................................................................................................................................... 10

– Setae on lateral margins of pronotum and elytra in fresh specimens as long or longer than length of compound eye; aedeagus as in Figs 614–616 View Figs 601–616 or 703–705 ................................................................ 9

9. Hypomeron separated from prosternum by low straight carina; aedeagus as in Figs 614–616 View Figs 601–616 .......... ............................................................................................................................ M. esau Heller, 1914 View in CoL

– Hypomeron almost fused to prosternum, carina at place of fusion not indicated or inconspicuous; aedeagus as in Figs 703–705 View Figs 697–711 ..................................................................... M. rugicollis Moser, 1920 View in CoL

10. Lateral margin of pronotum distinctly emarginate or strongly constricted prebasally.................... 11

– Lateral margin of pronotum not or hardly emarginate prebasally, evenly rounded all along or slightly constricted prebasally ..................................................................................................................... 16

11. Lateral margin of pronotum moderately strongly emarginate or constricted, upper angle of constriction not produced posteriad, either obtuse or completely rounded; aedeagus not as in Figs 601–603 View Figs 601–616 ... 12

– Lateral margin of pronotum very deeply emarginate prebasally, upper angle of emargination produced posteriad and acute ( Fig. 206 View Figs 193–207 ); lateral margin of pronotum in emargination area almost smooth, markedly less strongly crenulate than anterior part of it; aedeagus as in Figs 601– 603 View Figs 601–616 .......................................................................................................................... M. brocki View in CoL sp. nov.

12. Underside /lateral margins of male basal metatarsomere with ordinary short or long setae, which are not arranged brush-like .................................................................................................................... 14

– Ventrolateral margins of male basal metatarsomere with a brush of long (sub)erect setae (cf. Figs 436–437 View Figs 436–450 ) .................................................................................................................................. 13

13. Abdominal ventrites medially with golden (sub)erect setae ( Fig. 435 View Figs 424–435 ); aedeagus as in Figs 730– 732 View Figs 726–738 ................................................................................................................. M. tarsalis Arrow, 1941 View in CoL

– Abdominal ventrites without golden (sub)erect setae; aedeagus as in Figs 620–622 View Figs 617–631 ........................ ...................................................................................................................... M. hirtipes Arrow, 1941 View in CoL

14. Lateral margin of pronotum markedly emarginate prebasally; intervening spaces of elytra densely microreticulate, opaque; basal protarsomere without tooth; aedeagus not as in Figs 745–747 View Figs 739–750 ...... 15

– Lateral margin of pronotum rather constricted prebasally, not markedly emarginate; intervening spaces of elytra smooth or microscopically wrinkled, glossy; basal protarsomere in both sexes with central tooth ( Fig. 447 View Figs 436–450 ); aedeagus as in Figs 745–747 View Figs 739–750 ....................................... M. yamdena View in CoL sp. nov.

15. Underside of male basal metatarsomere with brush of long erect setae; aedeagus as in Figs 724– 725 View Figs 712–725 .......................................................................................................... M. subcostatus Heller, 1895 View in CoL

– Underside of male basal metatarsomere with a long setae which are not brush-like arranged and rather suberect; aedeagus as in Figs 709–711 View Figs 697–711 ......................................................... M. similis View in CoL sp. nov.

16. Lateral margin of pronotum with 20 or more delicate crenulae; lateral margin of male labroclypeus variably strongly sinuous; aedeagus not as in Figs 706–708 View Figs 697–711 .......................................................... 18

– Lateral margin of pronotum with 13–16 rough crenulae; lateral margin of male labroclypeus strongly sinuous ( Figs 117 View Figs 109–120 , 171 View Figs 169–180 )..................................................................17 [two possibly conspecific taxa]

17. Intervening spaces on pronotal dorsum completely flat; pronotal disc without impressions; dorsal punctures of pronotum comparatively stronger ovoid ( Figs 171 View Figs 169–180 , 250 View Figs 242–258 ); each elytron with vague tracks of three longitudinal carinae (including sutural one); aedeagus as in Figs 706– 708 View Figs 697–711 ...................................................................................................... M. seriegranosus Heller, 1914 View in CoL

– Intervening spaces on pronotal dorsum in part slightly globose, not completely flat; pronotal disc with vague circular impression on either side of hardly longitudinally impressed midline; dorsal punctures of pronotum rather circular to irregularly hexagonal, distinctly ovoid along lateral sides only ( Figs 38 View Figs 38–41 , 117 View Figs 109–120 ); each elytron with vague tracks of four longitudinal carinae (including sutural one); aedeagus unknown.......................................................................... M. fraterculus Moser, 1920 View in CoL

18. Intervening spaces of elytra opaque, microreticulate ...................................................................... 19

– Intervening spaces of elytra glossy, smooth or wrinkled, not microreticulate ................................ 23

19. Male and female labroclypeus emarginate or subtruncate (not truncate) on anterior margin, bent up to with regard to axis of frons in lateral view; head dorsally flattened or convex on frons, impressed dorsally at anterolateral angles of labroclypeus; pronotal punctures annular or not; crenulae lateral pronotal margin separated by delicate shallow or broad and distinct spaces ................................. 20

– Female labroclypeus gradually constricted anteriad on lateral margins, its anterior margin truncate ( Fig. 156 View Figs 145–156 ), not or hardly bent up with regard to axis of frons in lateral view; head dorsally flattened, hardly impressed dorsally at anterolateral angles of labroclypeus; pronotal punctures not annular; crenulae of lateral pronotal margin separated by delicate shallow spaces; male is unknown .................................................................................................... M. papuanus Moser, 1926 View in CoL

20. Parameres long and slender, constricted towards apices ................................................................. 21

– Parameres short and broad, widened apically ( Figs 679–680 View Figs 667–681 ) ............... M. nepenthephilus View in CoL sp. nov.

21. Parameres narrowly separated at base ( Figs 655–657 View Figs 652–666 , 748–750 View Figs 739–750 ); crenulae of lateral pronotal margin separated by distinct moderately deep and broad intervening spaces ............................................. 22

– Parameres moderately broadly separated at base ( Figs 736–738 View Figs 726–738 ); crenulae of lateral pronotal margin narrowly separated in anterior half, nearly merged ....................................... M. vicinus Heller, 1914 View in CoL

22. Lateral margin of pronotum evenly broadly rounded ( Fig. 229 View Figs 226–241 ) ........... M. luniceps Fairmaire, 1883 View in CoL

– Lateral margin of pronotum somewhat constricted towards base in basal third ( Fig. 242 View Figs 242–258 ) ................ .................................................................................................................. M. paupianus Heller, 1910 View in CoL

23. Elytral incision-shaped punctures distinctly sinuous....................................................................... 27

– Elytral incision-shaped punctures more or less straight .................................................................. 24

24. Dorsal forebody with greenish metallic lustre; dorsal punctures of pronotum regularly circular to ovoid ................................................................................................................................................ 25

– Forebody without metallic lustre; dorsal punctures of pronotum, especially lateral and laterobasal ones, irregularly ovoid, double annular and large ( Figs 95 View Figs 94–96 , 264 View Figs 259–265 ); aedeagus as in Figs 742– 744 View Figs 739–750 ..................................................................................................... M. woodlarkianus Heller, 1914 View in CoL

25. Intervening spaces of elytra smooth or in part delicately wrinkled; elytral longitudinal carinae vague and inconspicuous; aedeagus is unknown ....................................................................................... 26

– Intervening spaces of elytra irregularly roughly wrinkled; elytral longitudinal carinae rather broad and elevated; aedeagus as in Figs 670–672 View Figs 667–681 .............................................. M. milneanus Heller, 1914 View in CoL

26. Male labroclypeus narrowed anteriad, truncate on anterior margin (female unknown); anterolateral angles of labroclypeus obtuse, not marked ( Fig. 187 View Figs 181–192 ); aedeagus as in Figs 733– 735 View Figs 726–738 ........................................................................................................................ M. trivialis View in CoL sp. nov.

– Female labroclypeus not markedly narrowed anteriad, subtruncate on anterior margin (male unknown); anterolateral angles of labroclypeus obtuse but marked ( Fig. 97 View Figs 97–108 ) .................................... ................................................................................................................. M. aenescens Heller, 1910 View in CoL

27. Total body length less than 8 mm; body brown to castaneous brown ............................................. 29

– Total body length about 9–10 mm; body black to dark brown........................................................ 28

28. Dorsal punctures of pronotum oblong-ovoid, horseshoe-shaped, moderately dense; regular transverse row of punctures present along anterior and basal margin of pronotum; male labroclypeus broadly V-shape emarginate on anterior margin ( Fig. 130 View Figs 121–132 ); aedeagus unknown .... M. jobiensis Moser, 1920 View in CoL

– Dorsal punctures of pronotum circular to ovoid, horseshoe-shaped along lateral margins only, sparse; no row of punctures along either anterior and basal margins of pronotum, male labroclypeus shallowly emarginate on anterior margin ( Fig. 101 View Figs 97–108 ); aedeagus unknown........ M. aroae Heller, 1914 View in CoL

29. Lateral margin of pronotum stronger rounded, constricted anteriad and posteriad; posterolateral angle of pronotum obtuse ................................................................................................................ 30

– Lateral margin of pronotum less rounded, stronger constricted anteriad than posteriad; posterolateral angle of pronotum nearly right-angled ............................... M. moluccanus Moser, 1920 View in CoL comb. rest.

30. Aedeagus as in Figs 649–651 View Figs 632–651 ...................................................................... M. lobaticeps Frey, 1969 View in CoL

– Aedeagus as in Figs 700–702 View Figs 697–711 ................................................................................. M. riedeli View in CoL sp. nov.

31. With conspicuous dense brush-like cluster of setae at least along lateral margins of pronotum or on its postero-lateral angles (cf. Figs 122 View Figs 121–132 , 221 View Figs 208–225 , 238 View Figs 226–241 , 246 View Figs 242–258 , 261 View Figs 259–265 , 358 View Figs 349–358 ).................................................... 32

– Dorsal pubescence ordinary or scale-like, not brush-like clustered ............................................... 36

32. Pronotum slightly narrower than combined width of elytra............................................................ 33

– Pronotum distinctly narrower than combined width of elytra...... M. helleri ( Frey, 1969) View in CoL comb. nov.

33. Lateral margin of pronotum flattened and laterally expanded in anterior half, strongly emarginate postmedially ( Figs 246 View Figs 242–258 , 261 View Figs 259–265 ) ..............................................................................34 [ M. ursus View in CoL group]

– Lateral margin of pronotum not expanded laterally in anterior half, postmedian emargination short, inconspicuous ( Fig. 221 View Figs 208–225 ) ................................................................................. M. kazantsevi View in CoL sp. nov.

34. Elytra without longitudinal rows of glossy glabrous humps ........................................................... 35

– Longitudinal carinae of elytra formed by large irregular glossy glabrous humps ( Figs 74 View Figs 74–77 , 337– 338 View Figs 333–348 ) ............................................................................................ M. perlatus ( Frey, 1969) View in CoL comb. nov.

35. Metatibia widened, flattened and glabrous on inner margin ( Fig. 444 View Figs 436–450 ); sutural carina of elytron present; lateral postmedian emargination of pronotum deep ( Fig. 261 View Figs 259–265 ) ................. M. ursus View in CoL sp. nov.

– Metatibia slender, not flattened and glabrous on inner margin; sutural carina of elytron absent; lateral postmedian emargination of pronotum moderately strong ( Fig. 238 View Figs 226–241 )................................................. .............................................................................................. M. opatroides Arrow, 1941 View in CoL comb. rest.

36. Dorsal sculpture of pronotum and elytra of strongly irregularly arranged punctures forming conspicuous fingerprint-like pattern ( Figs 297 View Figs 286–301 , 347 View Figs 333–348 ).......................................37 [ M. sturnus View in CoL group]

– Punctures of pronotum and elytra not fingerprint-like arranged, not like in Figs 297 View Figs 286–301 , 347 View Figs 333–348 ............ 38

37. Middle of posterior margin of male abdominal sternites 2–3 each with a paired bunch of three long setae on each side ( Fig. 433 View Figs 424–435 ); aedeagus as in Figs 721–723 View Figs 712–725 ........................ M. sturnus Arrow, 1941 View in CoL

– Male abdominal sternites medially with somewhat stronger appressed curved setae but without paired bunches of long setae; aedeagus as in Figs 592–594 View Figs 586–600 .............................. M. caperatus View in CoL sp. nov.

38. Male 3 rd visible abdominal sternite with obtuse ventral median triangular hump ( Figs 429– 432 View Figs 424–435 ) .......................................................................................................................39 [ M. dani View in CoL group]

– Male 3 rd visible abdominal sternite not humped, evenly rounded or flattened ventrally ................ 40

39. Male labroclypeus comparatively shallower emarginate on anterior margin ( Fig. 121 View Figs 121–132 ); ventral hump of visible abdominal ventrite 3 acute in ventral view ( Figs 431–432 View Figs 424–435 ); aedeagus as in Figs 617– 619 View Figs 617–631 ...................................................................................................................... M. hamatus View in CoL sp. nov.

– Male labroclypeus comparatively deeper emarginate on anterior margin ( Fig. 114 View Figs 109–120 ); ventral hump of visible abdominal ventrite 3 rather obtuse in ventral view ( Figs 429–430 View Figs 424–435 ); aedeagus as in Figs 604– 606 View Figs 601–616 ............................................................................................................................. M. dani View in CoL sp. nov.

40. Punctures of elytra circular to ovoid, arranged in more or less regular paired longitudinal rows (cf. Figs 327 View Figs 317–332 , 345 View Figs 333–348 ) .................................................................................................................................. 41

– Punctures of elytra circular, ovoid or of different shape, if arranged in longitudinal rows then those not paired ......................................................................................................................................... 46 41. Lateral margin of pronotum distinctly emarginate prebasally ( Fig. 255 View Figs 242–258 and Narakusumo & Balke 2019: fig. 5a).................................................................................................................................... 42

– Lateral margin of pronotum evenly broadly rounded, not or indistinctly emarginate prebasally ... 43

42. Generally larger species, total body length over 9 mm; male metatibia slender, inner margin straight; aedeagus as in Figs 715–717 View Figs 712–725 ................................................................................... M. sougb View in CoL sp. nov.

– Generally smaller species, total body length under 7 mm; male metatibia thickened and somewhat curved on inner margin (cf. Narakusumo & Balke 2019: fig. 5e); aedeagus as in Narakusumo & Balke (2019: fig. 5f–g)................................... M. obiensis ( Narakusumo & Balke, 2019) View in CoL comb. nov.

43. Generally larger species, total body length over 6 mm; labroclypeus deeply emarginate anteriorly; aedeagus not as in Figs 676–678 View Figs 667–681 ..................................................................................................... 44

– Generally smaller species, total body length under 5 mm; labroclypeus shallowly emarginate anteriorly; aedeagus as in Figs 676–678 View Figs 667–681 ......................................................... M. nanus Arrow, 1941 View in CoL

44. Male metatibia straight, not curved ................................................................................................. 45

– Male metatibia strongly curved (cf. Narakusumo & Balke 2019: fig. 2e); aedeagus as in Narakusumo & Balke (2019: fig. 2f–g).............................. M. arcuatus ( Narakusumo & Balke, 2019) View in CoL comb. nov.

45. Crenulae of lateral margin of pronotum flat, nearly merged, intervening spaces indistinct ( Fig. 244 View Figs 242–258 ); aedeagus as in Figs 688–690 View Figs 682–696 ........................................................................... M. pedarioides Arrow View in CoL

– Crenulae of lateral margin of pronotum strong, intervening spaces rather deep and broad (cf. Narakusumo & Balke 2019: fig. 3a–b); aedeagus as in Narakusumo & Balke (2019: fig. 3e–f)........ ....................................................................... M. cakalele ( Narakusumo & Balke, 2019) View in CoL comb. nov.

46. Male basal metatarsomere ordinary, not conspicuously widened and flattened ............................. 50

– Male basal metatarsomere conspicuously widened and flattened (cf. Fig. 446 View Figs 436–450 ) ............................. 47

47. Dorsal setae of forebody long, surpassing length of corresponding punctures; aedeagus not as in Figs 640–642 View Figs 632–651 ................................................................................................................................... 48

– Dorsal setae of forebody inconspicuous, short, hardly surpassing length of corresponding punctures ( Fig. 315 View Figs 302–316 ); aedeagus as in Figs 640–642 View Figs 632–651 .............................................................. M. legalovi View in CoL sp. nov.

48. Dorsal setae at least on forebody long, pointed or hardly widened apically; male and female labroclypeus subtruncate on anterior margin................................................................................... 49

– Dorsal setae slightly clavate (widened apically), comparatively shorter ( Figs 58 View Figs 58–61 , 144 View Figs 133–144 , 231 View Figs 226–241 ); male labroclypeus comparatively deeper emarginate on anterior margin ( Fig. 144 View Figs 133–144 ); aedeagus as in Figs 661–663 View Figs 652–666 ........................................................................................................... M. maleo View in CoL sp. nov.

49. Aedeagus as in Figs 712–714 View Figs 712–725 .................................................................................. M. skalei View in CoL sp. nov.

– Aedeagus as in Figs 691–693 View Figs 682–696 ............................................................ M. peregrinus Lansberge, 1886 View in CoL

50. Parameres entirely glabrous............................................................................................................. 51

– Parameres on apical margin with delicate cilia-like setae ( Figs 595–596 View Figs 586–600 ) .......... M. ciliatus View in CoL sp. nov.

51. Lateral margins of pronotum in dorsal view distinctly rounded (more or less distinctly constricted anteriad and posteriad)..................................................................................................................... 52

– Lateral margins of pronotum in dorsal view nearly straight, not distinctly constricted anteriad and posteriad ( Fig. 230 View Figs 226–241 ); aedeagus as in Figs 658–660 View Figs 652–666 .................................................. M. mailu View in CoL sp. nov.

52. Hypomeron separated from prosternum by flange-like carina; preapical hump of elytron not Г- shaped........................................................................................................................................... 53

– Hypomeron separated from prosternum by moderately high carina; preapical hump of elytron Г- shaped.......................................................................................................... M. simplex Frey, 1969 View in CoL

53. Dorsal setae at least in part distinctly clavate (distinctly widened apically) ...................................... ..................................................................................................................... 66 [ M. pauxillus group]

– Dorsal setae ordinary or slightly widened apically.......................................................................... 54

54. Species from New Guinea or Sulawesi ........................................................................................... 55

– Species from Lesser Sunda Islands (Bali); aedeagus as in Figs 643–645 View Figs 632–651 ........ M. leucopsar View in CoL sp. nov.

55. Male labroclypeus subtruncate, truncate or shallowly emarginate on anterior margin .................. 56

– Male labroclypeus distinctly emarginate on anterior margin .......................................................... 58

56. Species from Sulawesi; aedeagus as in Figs 589–591 View Figs 586–600 or 726–729 ................................................. 57

– Species from New Guinea; aedeagus as in Figs 673–675 View Figs 667–681 ............................ M. muticus Arrow, 1941 View in CoL

57. Aedeagus as in Figs 589–591 View Figs 586–600 ; lateral margin of each elytron with a row of short setae; elytron on disc with track of single longitudinal carina.................................................... M. boessnecki View in CoL sp. nov.

– Aedeagus as in Figs 726–729 View Figs 726–738 ; lateral margin of each elytron with a row of long setae; elytron on disc with tracks of two longitudinal carinae ............................................................... M. suwawa View in CoL sp. nov.

58. Dorsal setae of forebody inconspicuous, appressed to suberect; setae short, generally not or slightly surpassing length of corresponding punctures (some setae can be longer than corresponding punctures, but most of them are shorter) ......................................................................................... 62

– Dorsal setae of forebody suberect to erect, distinctly longer than length of corresponding punctures.......................................................................................................................................... 59

59. Lateral margin of elytron with a row of short setae......................................................................... 60

– Lateral margin of elytron with a row of extraordinarily long setae ( Fig. 24 View Figs 22–24 ) ........... M. awu View in CoL sp. nov.

60. Pronotum dorsally with shallow circular impression on either side of median part; punctures of pronotum comparatively larger and more circular ......................................................................... 61

– Pronotum dorsally without impressions; punctures of pronotum comparatively smaller and stronger ovoid ( Fig. 287 View Figs 286–301 ); aedeagus as in Figs 570–572 View Figs 570–585 ...................................................... M. aiyura View in CoL sp. nov.

61. Aedeagus as in Figs 582–585 View Figs 570–585 ; dorsal punctures of pronotum smaller and generally ovoid ( Fig. 203 View Figs 193–207 ) ......................................................................................................... M. babyrousa View in CoL sp. nov.

– Aedeagus as in Figs 607–610 View Figs 601–616 ; dorsal punctures of pronotum larger and generally circular ( Fig. 211 View Figs 208–225 )............................................................................................................... M. deltouri View in CoL sp. nov.

62. Species from New Guinea; aedeagus not as in Figs 629–631 View Figs 617–631 ........................................................ 63

– Species from Sulawesi; aedeagus as in Figs 629–631 View Figs 617–631 ............................................. M. konjo View in CoL sp. nov.

63. Anterolateral angles of labroclypeus either acute or right-angled................................................... 64

– Anterolateral angles of labroclypeus strongly obtuse...................................................................... 65

64. Aedeagus as in Figs 667–669 View Figs 667–681 ; male upper metatibial terminal spur half the length of male basal metatarsomere, not surpassing distal margin of metatibia in dorsal view ( Fig. 59 View Figs 58–61 ) ................................................................................................ M. miklouhomaclayi View in CoL sp. nov.

– Aedeagus as in Figs 576–578 View Figs 570–585 ; male upper metatibial terminal spur longer than half the length of male basal metatarsomere, surpassing distal margin of metatibia in dorsal view ............................... .............................................................................................................. M. angusticeps Arrow, 1941 View in CoL

65. Elytral longitudinal carinae glabrous, glossy, stronger elevated; sutural carina present; intervening spaces on pygidium opaque, microreticulate; aedeagus unknown ...................................................... ............................................................................................... M. interruptocarinulatus Heller, 1914 View in CoL

– Elytra without elevated tracks of longitudinal carinae; sutural carina not indicated; intervening spaces on pygidium glossy; aedeagus unknown................................ M. seriepunctatus Moser, 1920 View in CoL

66. Elytron with elevated glossy and glabrous longitudinal carina(e) (these carinae are more or less widely interrupted, in some specimens strongly reduced to few humps per carina and can be limited to sutural carina only) ...................................................................................................................... 67

– Longitudinal carina(e) of elytron when present not elevated nor glossy or glabrous...................... 70

67. Dorsal setae on lateral parts of pronotum scale-like but not extraordinarily enlarged and not arranged in two lateral longitudinal stripes; aedeagus not as in Figs 586–588 View Figs 586–600 .............................................. 68

– Dorsal setae on lateral parts of pronotum scale-like and extraordinarily enlarged, as in Figs 26 View Figs 25–27 , 204 View Figs 193–207 , arranged in two lateral longitudinal rows; aedeagus as in Figs 586–588 View Figs 586–600 ............. M. bintang View in CoL sp. nov.

68. Lateral margin of pronotum nearly straight to gradually slightly widened towards postmedium, with shallow prebasal emargination; elytron with less than four longitudinal carinae including sutural one.................................................................................................................................................... 69

– Lateral margin of pronotum sinuous – somewhat emarginate in anterior and posterior half, obtuse angulate medially ( Fig. 216 View Figs 208–225 ); elytron with four in part interrupted elevated longitudinal carinae including sutural one................................................... M. heterosquamosus Heller, 1910 View in CoL comb. rest.

69. Aedeagus as in Figs 611–613 View Figs 601–616 ; male pygidium as in Fig. 543 ...................... M. dendrolagus View in CoL sp. nov.

– Aedeagus as in Figs 632–639 View Figs 632–651 ; male and female pygidium as in Figs 481–482 View Figs 469–486 ... M. lapsus View in CoL sp. nov.

– Aedeagus as in Figs 718–720 View Figs 712–725 ; male and female pygidium as in Figs 519-521 View Figs 504–521 .................................. ................................................................................................ M. speciosus ( Frey, 1969) View in CoL comb. nov.

70. Anterolateral angles of labroclypeus acute in dorsal view .............................................................. 71

– Anterolateral angles of labroclypeus obtuse in dorsal view ( Figs 169–170 View Figs 169–180 ); aedeagus as in Figs 697– 699 View Figs 697–711 .................................................................................................. M. popei ( Frey, 1969) View in CoL comb. nov.

71. Species from New Guinea............................................................................................................... 72

– Species from Central Moluccas (Seram); aedeagus as in Prokofiev (2018: fig. 5) ............................. .......................................................................................... M. agnellus ( Prokofiev, 2018) View in CoL comb. nov.

72. Aedeagus as in Figs 598–600 View Figs 586–600 ; male and female pygidium as in Figs 465–466 View Figs 451–468 .................................. ........................................................................................................................... M. crypticus View in CoL sp. nov.

– Aedeagus as in Figs 664–666 View Figs 652–666 ; male pygidium as in Fig. 492 View Figs487–503 ............................ M. merdeka View in CoL sp. nov.

– Aedeagus as in Figs 682–684 View Figs 682–696 ; male and female pygidium as in Figs 499 View Figs487–503 –5000................................ .................................................................................................................... M. owenstanleyi View in CoL sp. nov.

– Aedeagus as in Figs 685–687 View Figs 682–696 ; male and female pygidium as in Figs 505–506 View Figs 504–521 .................................. ................................................................................................ M. pauxillus Heller, 1910 comb. rest.

– Aedeagus as in Figs 739–741 View Figs 739–750 ; male and female pygidium as in Figs 531–532.... M. weigeli View in CoL sp. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

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