Mutovia intercalaria, Vršanský, Peter & Aristov, Danil, 2012

Vršanský, Peter & Aristov, Danil, 2012, Enigmatic Late Permian cockroaches from Isady, Russia (Blattida: Mutoviidae fam. n.), Zootaxa 3247, pp. 19-31 : 24-25

publication ID	https://doi.org/ 10.5281/zenodo.213150
DOI	https://doi.org/10.5281/zenodo.5674732
persistent identifier	https://treatment.plazi.org/id/1B41B46C-501B-7669-DE88-FB19FEE3A3DA
treatment provided by	Plazi
scientific name	Mutovia intercalaria
status	sp. nov.

Treatment

Mutovia intercalaria sp. n.

( Figs. 2–7 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 )

Type Material. Holotype: Kitchuga Member, Poldars Formation (uppermost Severodvinian stage, Upper Permian), Chroniosaurus levis Subzone of the Proelginia permiana Zone. Isady, Vologda territory, northwestern Russia. PIN 3840/57± (left forewing, male?). Paratypes: The same as for holotype: PIN 3840/54±L, 55±R, 56±L, 66L, 67R, 358±L, 362R, 355, 356, 433, 434, 435, 501, 513, 519, 520, 524, 526, 528, 529=533 (hind wing), 531±, 532±, 534, 535±, 537, 539, 362, 540±, 656, 798 (?males); 8R, 52±L, 53±L, 355R, 356R, 359+-L, 360L, 500, 511, 516, 517, 518+-, 522, 523, 525 527, 539 (?females); PIN 3840/538, 521 (sex undetermined).

Description. Forewing of medium size (15–19 mm x 5.1–6.5 mm). Sc (8–12 veins at margin), R1 (0–3 veins at margin) and RS (4–7 veins at margin) usually separated. M with 2–8 branches at margin, CuA with 6–8 veins. Clavus very short (less than one-fourth of wing length), CuP sometimes branched; anal veins simple or branched (up to 8–10 at margin). Colouration characteristically with pale main veins and dark intercalaries and reticulations, and with distinct coloured reticulation along clavus ( Fig. 2 View FIGURE 2 C).

Hind wing ( Figs. 3 View FIGURE 3 C, 5C) with long, simple, ascending branchelets, R1 with at least 5 veins at margin; RS simple or branched terminally. M expanded, with at least 5 veins at margin, with changed polarity from convex into concave near center of wing. CuA with about 5 veins (one secondarily branched), CuP simple.

Character of preservation. Of forty-nine similar wing fossils, 48 are forewings, with 30 of the wings probably corresponding to males, 17 to females, and 1 wing indeterminable. One wing is a male hindwing. Nine of the identified wings are left wings and six are right wings. The apical posterior margin is often folded, suggesting that the wing was fragile at that area (similar to Santanmantis Grimaldi, 2003 ). The lengths of completely preserved specimens are 13 mm (2 wings), 14 mm (2), 16 mm (7) and 19 mm (3). All other forewings are approximately 16 or 19–21mm long.

Derivation of name. The species name refers to the intercalary veins of the wings, which are well marked and colourful.