Crossopriza johncloudsleyi Deeleman-Reinhold & van Harten, 2001

Bernhard A. Huber & Charles M. Warui, 2012, East African pholcid spiders: an overview, with descriptions of eight new species (Araneae, Pholcidae), European Journal of Taxonomy 29 (29), pp. 1-44: 18-22

publication ID

http://dx.doi.org/10.5852/ejt.2012.29

publication LSID

lsid:zoobank.org:pub:28503B8D-9B2C-484D-A7A2-B44D4DC04F6D

persistent identifier

http://treatment.plazi.org/id/1C1A87FF-1048-FFAF-F7C6-121DFB05FD19

treatment provided by

Jeremy

scientific name

Crossopriza johncloudsleyi Deeleman-Reinhold & van Harten, 2001
status

sp. nov.

Crossopriza johncloudsleyi Deeleman-Reinhold & van Harten, 2001 

Figs 28-31View Figs 28 - 49. — 28 - 31, 62-63View Figs 50 - 69. — 50 - 52, 70-74View Figs 70 - 74

Crossopriza johncloudsleyi Deeleman-Reinhold & van Harten, 2001: 197  -199; figs 11-16, ♂, ♀♀ (Yemen).

Diagnosis

Easily distinguished from known congeners by male clypeus modification (shapes of pair of apophyses; Fig. 29View Figs 28 - 49. — 28 - 31), male palp (shapes of procursus and bulbal process; Figs 70, 71View Figs 70 - 74), and distinctive epigynum (large pits and lateral processes; Figs 30View Figs 28 - 49. — 28 - 31, 62View Figs 50 - 69. — 50 - 52, 73View Figs 70 - 74).

Type material

Holotype

♂, examinedGoogleMaps  .

Paratypes

5 ♀♀, 1 juv. (3 ♀♀ examined, the other two ♀♀ paratypes could not be located)GoogleMaps  .

Type locality

YEMEN, Ja’ar (13°13’N, 45°18’E), 50 m a.s.l., 15-16 May 2000 (A. van Harten), in RMNH ( 15173).

Other material examined

KENYA: Rift Valley Province: 2 ♂♂, 2 ♀♀, in CJFM (1 ♂, 1 ♀, 3841)GoogleMaps  and ZFMK (1 ♂, 1 ♀, Ar 8736), Lake Hannington [= Lake Bogoria , ~ 0°15’N, 36°05’E], 1000 m a.s.l., lakeside scrub, 2 Aug. 1974 (Murphy)GoogleMaps  ; 2 ♀♀, in CJFM ( 2275), Lake Baringo [~ 0°38’N, 36°01’E, 980 m a.s.l.], 28 Aug. 1972 (Murphy)GoogleMaps  ; 1 ♀, in CJFM ( 3805), Baringo , near hot springs, 1100 m a.s.l., 31 Jul. 1974 (Murphy)  ; 1 ♀, 2 juvs., in MCZ ( 34026), Lokori [~ 1°58’N, 36°06’E, 600 m a.s.l.], 16 Aug. 1965 (B. Patterson Expedition)GoogleMaps  ; 1 ♀, in MCZ ( 34012), 15 miles ENE Lokori [~ 2°02’N, 36°18’E, 600 m a.s.l.], 28-29 Jun. 1966 (collector not given)GoogleMaps  ; 1 ♂, in MCZ ( 34022), Lothagam (2°56’N, 36°04’E) [~ 400 m a.s.l.], 2-5 Aug. 1967 (B. Patterson)GoogleMaps  ; 1 ♀, in MCZ ( 34018), Kangatotha (“ 40 km E of Lodwar, W of Lake Rudolf”) [~ 3°07’N, 36°00’E, 400 m a.s.l.], 27 Aug. – 7 Sep. 1963 (B. Patterson)GoogleMaps  ; 1 ♂, in NMH, Kajiado [1°51’S, 36°47’E, 1700 m a.s.l.], road culverts, May 1973 (collector not given)GoogleMaps  .

Description

Male (Lake Bogoria)

MEASUREMENTS. Total body length 2.8, carapace width 1.2. Leg 1: 21.0 (6.0 + 0.5 + 5.8 + 7.2 + 1.5), tibia 2: 3.7, tibia 3: 2.7, tibia 4: 3.3; tibia 1 L/d: 50. Distance PME-PME 70 µm, diameter PME 85 µm, distance PME-ALE 35 µm, distance AME-AME 25 µm, diameter AME 85 µm.

COLOR. Carapace ochre-yellow, medially slightly darker including ocular area and clypeus, sternum light brown with slightly darker radiating marks, legs ochre-yellow with many small black marks, mostly on femora ( Fig. 28View Figs 28 - 49. — 28 - 31) and tibiae, not on coxae, few on metatarsi, abdomen ochre-grey with indistinct small dark and white internal marks dorsally and laterally, ventrally with darker pattern, wider anterior of gonopore.

BODY. Habitus as in Figs 28 and 29View Figs 28 - 49. — 28 - 31; ocular area elevated, compact, no ‘accessory eyes’ (cf. Huber 2009); carapace with thoracic pit; clypeus with pair of distally slightly hooked apophyses on rim, ~140 µm long. Chelicerae as in Fig. 72View Figs 70 - 74, with large frontal apophyses provided with one thick modified hair each and pair of smaller apophyses laterally, some hair bases accompanied by small cuticular elevations, fine stridulatory ridges. Sternum wider than long (0.78/0.50), unmodified.

PALPS. As in Figs 70 and 71View Figs 70 - 74, coxa with prominent joint (arrow) and retrolateral hump, trochanter simple, femur with distinct ventral apophysis, tibia very large, procursus curved towards femur, with strong.

distal apophysis, bulb with only one process (bulbal apophysis) with distinctive prolatero-ventral ridges, sperm duct apparently opening at basis of bulbal apophysis.

LEGS. Without curved hairs, few vertical hairs, with spines ventrally on femur 1 (single row on distal half, ~17 spines); retrolateral trichobothrium on tibia 1 at 4%; prolateral trichobothrium absent on tibia 1, present on other tibiae; pseudosegments not visible in dissecting microscope.

VARIATION. Tibia 1 in two other males from Kenya: 5.6, 5.7. The holotype is slightly smaller (tibia 1: 4.5), lacks the small proximal apophyses on the chelicerae (arrow in Fig. 72View Figs 70 - 74), and has a slightly differently shaped palpal femur (ventral projection; see fig. 15 in Deeleman-Reinhold & van Harten 2001).

Female

In general similar to male but femur 1 without spines, clypeus unmodified, chelicerae without stridulatory ridges; tibia 1 in 6 females from Kenya: 4.6-5.2 (mean 4.8); in 3 females from Yemen: 3.4-3.5. Without stridulatory apparatus between prosoma and abdomen. Epigynum with large and complex frontal plate with pair of long lateral apophyses, pair of deep pits, and posterior elevated transversal ridge; simple posterior plate; internal genitalia as in Figs 63View Figs 50 - 69. — 50 - 52 and 74View Figs 70 - 74. Paratypes with slightly smaller epigyna with slightly shorter projections, otherwise identical (also in dorsal view). ALS with only two spigots each.

Distribution

Known from Yemen and from several localities in Kenya, Rift Valley Province ( Fig. 19View Figs 16 - 19).

RMNH

Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]

CJFM

CJFM

ZFMK

Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"

MCZ

USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology

NMH

NMH

AME

USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Pholcidae

Genus

Crossopriza