Cosmodela virgula ( Fleutiaux, 1893 )

Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, Zootaxa 4661 (2), pp. 271-308 : 287-288

Cosmodela virgula ( Fleutiaux, 1893)

( Figs 8D , 13C )

Cicindela aurulenta var. virgula Fleutiaux, 1893:491 .

Cicindela aurulenta virgula: Fowler 1912:382 .

Cosmodela aurulenta virgula: Rivalier 1961:128 .

Cicindela (Cosmodela) virgula: Acciavatti & Pearson 1989:132 .

Cosmodela virgula: Naviaux 1991:274 .

Specimens examined. (N = 4) [ Laos] 1♂ (EUMJ), Phu Pan, altitude 1600-1700 m, Xam Neua prefecture, Houapan province, 19. V. 2012, T. Kikuhara leg; 1♂ ( EUMJ), XamNeua., Ph-Pan. Alt. 1750 m, 28. VI. – 2. V. 2002, M. Sato leg; [NE Laos] 1♂ ( EUMJ), Houaphan Prov., Xam Nua District, Xam Nua District, Ban Saleui , 1350 m, 22-24. VI. 2003, S. Nagai leg ; 1♂ ( EUMJ), Sam Neua , 17. VI. 2002, unknown leg .

Description. Internal sac everted widely and dorsally in lateral view. DA largest in bladders; B, VLL and A the second largest; VA, DLL and VLR the third largest; VB and AL smallest in the bladders.

A cylindrically swollen, everted apically. AL and VB slightly bulging at base of A, vested with fine spines, which are forming oblong spot. DA well swollen basally. DLL roundly swollen, vested with fine spines. VA short triangular swollen at base of DA, everted apically. VLR swollen left laterally at base of VA. VLL oval and widely swollen right laterally, with well sclerotized spines, which are forming a wide band. B roundly swollen right ventrolaterally, with well sclerotized spines, which are forming an oblong spot. Flagellum very long, curved and formed S-shaped. Medial tooth extending apically.

Remarks. This species shares the characteristics of the position of each bladder with C. duponti but is distinguished by the following characteristics: A everted apically. DLL small.

Figures

FIGURE 8. Habitus of Cosmodela, Cicindela (Cicindela) and Cylindera.A, Cosmodela sp. (cf. separata); B, C. batesi (Fleutiaux) from Iriomote Is., Japan; C, C. aurulenta (Fabricius) from Penang Is., Malaysia; D, C. virgula (Fleutiaux) from Laos; E, C. duponti duponti (Dejean) from South India; F, Cicindela (Cicindela) japana japana Motschulsky; G, C. (C.) gemmata aino Lewis; H, C. (C.) sachaliensis niohozana Bates; I, C. (C.) lewisii lewisii Bates; J, Cylindera (Eugrapha) elisae elisae (Motschulsky); K, Cicindela (Cicindela) transbaicalica japanensis Chaudoir. Scales: 10 mm.

FIGURE 13. Everted internal sacs of Cosmodela. A, Cosmodela sp. (cf. separata) from Vietnam; B, C. aurulenta (Fabricius) from Penang Is., Malaysia; C, C. virgula (Fleutiaux) from Laos; D, C. duponti duponti (Dejean) from South India. d1, d2, d3, d4, show the connecting of flagellum. Scales: 1.0 mm.