Philonthus orientalis, Motschulsky, 1858 a: 212

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2493-2506

orientalis Motschulsky, 1858a: 212 ( Platyprosopus ; Type locality: Indes orientales).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; synonym of fuliginosus ).

— Cameron, 1932: 51 ( Platyprosopus ; synonym of fuliginosus ).

fulvicollis Motschulsky, 1858a: 212 ( Platyprosopus ; Type locality: Indes orientales).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; catalog).

— Cameron, 1932: 51 ( Platyprosopus ; characters; Bihar).

— Scheerpeltz, 1933: 1325 ( Platyprosopus ; catalog).

— Coiffait, 1982a: 232 ( Platyprosopus ; India).

- DISTRIBUTION: India, “East Indies”.

linearis Kraatz, 1859: 115 ( Platyprosopus ; Type locality: India orientalis).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; synonym of fulvicollis ).

— Cameron, 1932: 51 ( Platyprosopus ; synonym of fulvicollis ).

ghidinii Cerruti, 1950: 49 ( Platyprosopus ; Type locality: Gambela, Galla-Sidamo ).

- DISTRIBUTION: Ethiopia.

grandiceps Irmler, 1977: 65 ( Platyprosopus ; Type locality: Estancia la Noria , Rio San Javier, Santa Fé, Argentine).

- DISTRIBUTION: Argentina.

gridellii Cerruti, 1950: 50 ( Platyprosopus ; Type locality: Eritrea: Tessenei ).

- DISTRIBUTION: Eritrea.

hierichonticus Reiche and Saulcy, 1856: 361 ( Platyprosopus ; Type locality: Jourdain ; Chypre).

— Kraatz, 1858a: 65 ( Platyprosopus ; characters; Greece).

— Peyron, 1858: 420 ( Platyprosopus ; Turkey).

— Reitter, 1908: 104 ( Platyprosopus ; cited as hierochonticus; characters; Syria; Palestine; Cyprus; Greece).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; catalog).

— Scheerpeltz, 1933: 1325 ( Platyprosopus ; catalog).

— Coiffait, 1972: 378 ( Platyprosopus ; cited as hierochonticus; characters; references; Palestine; Syria; Asia Minor; Cyprus; Greece).

— Assing and Wunderle, 2001: 36 ( Platyprosopus ; Cyprus).

- DISTRIBUTION: Israel, Jordan, Syria, Cyprus, Turkey, Greece.

indicus Motschulsky, 1858a: 212 ( Platyprosopus ; Type locality: Indes orientales).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; catalog).

— Bernhauer, 1928: 15 ( Platyprosopus ; Sumatra).

— Cameron, 1932: 49 ( Platyprosopus ; characters; India).

— Scheerpeltz, 1933: 1325 ( Platyprosopus ; catalog).

— Cameron, 1933d: 386 ( Platyprosopus ; Sumatra).

— Paulian, 1941: 217 ( Platyprosopus ; larval characters; India).

- DISTRIBUTION: India, Indonesia.

laevicollis Kraatz, 1859: 114 ( Platyprosopus ; Type locality: India borealis).

— Fauvel, 1895: 239 ( Platyprosopus ; synonym of indicus ).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; synonym of indicus ).

— Cameron, 1932: 49 ( Platyprosopus ; synonym of indicus ).

indicus Kraatz, 1859 , see: elatus Fauvel, 1895 .

keiseri Scheerpeltz, 1961 g: 238, 257 ( Platyprosopus ; subgenus Platyprosopus ; Type locality: West-Madagaskar: Tsaramandroso).

- DISTRIBUTION: Madagascar.

kraatzi Bernhauer, 1908d: 326 ( Platyprosopus ; Type locality: Paraguay).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; catalog).

— Irmler, 1977: 60 ( Platyprosopus ; characters; Paraguay; Argentina).

- DISTRIBUTION: Paraguay, Argentina.

laevicollis Kraatz, 1859 , see: indicus Motschulsky, 1858 .

laticeps Sharp, 1876: 103 ( Platyprosopus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; catalog).

- DISTRIBUTION: Brazil.

linearis Kraatz, 1859 , see: fulvicollis Motschulsky, 1858 .

longicollis Eppelsheim, 1885: 113 ( Platyprosopus ; Type locality: Goldküste: Adda).

— Bernhauer and Schubert, 1914: 320 ( Platyprosopus ; catalog).

- DISTRIBUTION: Ghana.

major Sharp, 1876: 102 ( Platyprosopus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

— Irmler, 1977: 58 ( Platyprosopus ; characters; Brazil; Peru).

- DISTRIBUTION: Brazil, Peru.

marshalli Bernhauer, 1937: 300 ( Platyprosopus ; Type locality: Kenya: Homa ,

South Kavirondo).

- DISTRIBUTION: Kenya.

mateui Coiffait, 1966b: 341 ( Platyprosopus ; Type locality: Mauritanie, Tamour Tartafet , Tamchakett).

— Coiffait, 1972: 379 ( Platyprosopus ; characters; Mauritania).

- DISTRIBUTION: Mauritania.

mexicanus Sharp, 1887: 783 ( Platyprosopus ; Type locality: Mexico, San Juan Bautista in Tabasco, Tlacotalpam; [ Note : See lectotype designation by Smetana, 1982]).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

— Irmler, 1977: 60 ( Platyprosopus ; characters; Mexico).

— Smetana, 1982: 24 ( Platyprosopus ; lectotype designation: San Juan Bautista ; characters; Mexico; Texas) .

- DISTRIBUTION: Mexico, USA.

texanus Moore, 1963a: 96 ( Platyprosopus ; Type locality: Texas: Brownsville ).

— Smetana, 1982: 24 ( Platyprosopus ; synonym of mexicanus ).

minor Sharp, 1876: 106 ( Platyprosopus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

— Irmler, 1977: 62 ( Platyprosopus ; characters; Brazil).

- DISTRIBUTION: Brazil.

nigriventris Cameron, 1947a: 55 ( Platyprosopus ; Type locality: Abyssinia: Nargi ).

- DISTRIBUTION: Ethiopia.

niloticus Fauvel, 1904c: 275 ( Platyprosopus ; Type locality: Haute-Egypte : Assouan. Abyssinie).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

— C. Koch, 1936b: 163 ( Platyprosopus ; characters; Egypt).

— Coiffait, 1972: 378 ( Platyprosopus ; characters; Egypt; Ethiopia).

- DISTRIBUTION: Ethiopia, Egypt.

ochraceus Fauvel, 1900b: 72 ( Platyprosopus ; Type locality: Congo: Kinchassa ).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

- DISTRIBUTION: Congo.

opacifrons Sharp, 1876: 107 ( Platyprosopus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

— Irmler, 1977: 62 ( Platyprosopus ; characters; Brazil).

- DISTRIBUTION: Brazil.

orientalis Motschulsky, 1858 , see: fuliginosus Erichson, 1840 .

pallescens Tottenham, 1961: 206 ( Platyprosopus ; Type locality: Senegal: Parc National du Niokolo-Koba, Badi).

- DISTRIBUTION: Senegal.

parallelus Sharp, 1876: 104 ( Platyprosopus ; Type locality: Ega).

— Bernhauer and Schubert, 1914: 321 ( Platyprosopus ; catalog).

— Scheerpeltz, 1940: 42 (key to Palaearctic genera).

— Blackwelder, 1943: 394 (characters; key to genera of West Indies).

— Blackwelder, 1944: 131 (checklist of species from Mexico, Central and South America, and

the West Indies).

— V. Hansen, 1952: 27 (characters; key to Danish genera).

— Coiffait, 1956e: 183, 186 (key to genera).

— Hatch, 1957: 173 (key to genera of Pacific Northwest).

— Smetana, 1958: 131 (characters; key to Czechoslovakian genera and species).

— Arnett, 1963: 275 (characters; list of North American genera).

— Palm, 1963b: 57 (characters; key to genera and species of Sweden).

— Lohse, 1964: 169 (characters; key to central European species).

— Pototskaia, 1967: 63 (larval characters).

— Coiffait and Saiz, 1968: 347, 363 (characters; Chilean species).

— Scheerpeltz, 1968a: 79 (catalog of Austrian species).

— Kasule, 1970: 56 (larval characters).

— Coiffait, 1974: 353 (characters; key to subtribes, genera, and species of western Palaearctic region).

— Muona, 1979: 15 (list of Scandinavian species).

— Burakowski, Mroczkowski, and Stefańska, 1980: 29 (catalog; Poland).

— Newton, 1985: 209 (notes on austral distribution and taxonomic notes).

— Outerelo and Gamarra, 1985: 53 (key to genera of Iberian Peninsula).

— Smetana, 1995e: 47, 48 (notes; key to subtribes).

— Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of

New Zealand species).

— M. Hansen, 1996: 104 (list of Danish species).

— Smetana and Davies, 2000: 4, 12, 37 (discussion of characters; characters and key to subtribes of north temperate region; checklist of genera and species).

— Smetana and Davies, 2000: 18 (key to genera and subgenera associated with Staphylinus

sensu lato).

— Newton, Thayer, Ashe, and Chandler, 2000: 330, 392 (key to Nearctic subtribes; characters; notes).

Subtribe Amblyopinina

Amblyopinina Seevers, 1944 .

— Newton and Thayer, 1992: 65 (subtribe of Staphylinini ).

— Smetana, 1995e: 48 (characters).

Amblyopininae Seevers, 1944: 157 (genera included: Amblyopinus ). TYPE GENUS: Amblyopinus (see Newton and Thayer, 1992: 65).

— Seevers, 1955 (revision; ecology; distribution; host relationships and list; evolution; key to genera).

— Coiffait and Saiz, 1968: 365 (notes; key to Chilean genera).

— Lawrence and Newton, 1995: 830 (included in Staphylininae ).

Amblyopinini Seever, 1944 .

— Machado-Allison, 1963: 380 (key to genera).

— Machado-Allison and Barrera, 1964: 182 (discussion of characters of some genera).

— Barrera, 1972: 79, 82 (key to genera of Peru; discussion).

— Newton, 1985: 209 (notes on austral distribution and taxonomic notes).

— Ashe and Timm, 1988: 48, 54–56 (key to genera; notes on interaction with hosts and on phylogenetic relationships).

— Timm and Ashe, 1989: 379 (comments on systematics of group).

Amblyopinodes

[12 species; Neotropical Region]

Amblyopinodes Seevers, 1955: 247 (species included: gahani , piceus , travassosi , claviger , peruvianus , similis ). TYPE SPECIES: Amblyopinus gahani Fauvel , fixed by original

designation.

— Machado-Allison, 1962: 313 (key to species).

— Machado-Allison, 1962a (notes; key to species; checklist of species with hosts).

— Machado-Allison, 1963 (characters; revision and classification of and key to species; list rodent hosts; distribution).

— Machado-Allison and Barrera, 1964: 181 (characters).

— Martinez, Barrera, and Machado-Allison, 1970: 132 (characters).

— Ashe and Timm, 1988: 49 (characters).

Streptopinodes Martinez, Barrera, and Machado-Allison, 1970: 133 (subgenus of Amblyopinodes ; species included: longus ). TYPE SPECIES: Amblyopinus longus E. Franz , fixed by original designation and monotypy.

adae Machado-Allison, 1962a: 83 ( Amblyopinodes ; Type locality: Brasil: Estado de São Paulo, Estación Biolólgica de Boraceia ).

— Machado-Allison, 1963: 386 ( Amblyopinodes ; characters; notes; host rodent).

- DISTRIBUTION: Brazil.

amazonicus Guerrero, 1996: 660 ( Amblyopinodes ; Type locality: Peru: Madre de Dios, Pakitza ) .

- DISTRIBUTION: Peru.

barrerai Machado-Allison, 1962 , see: caviae Martínez and Prosen, 1958 .

caviae Martínez and Prosen, 1958: 27 ( Amblyopinodes ; Type locality: Paraguay: Dpto. Boquerón, Gran Chaco, Guarnición Oruro ).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 389 ( Amblyopinodes ; characters; notes; host rodent).

— Martinez, Barrera, and Machado-Allison, 1970: 136 ( Amblyopinodes ; subgenus Amblyopinodes ; characters; Argentina; Bolivia; Brazil; Paraguay).

- DISTRIBUTION: Argentina, Paraguay, Bolivia, Brazil.

barrerai Machado-Allison, 1962: 324 ( Amblyopinodes ; Type locality: Brasil: Viçosa , Estado de Ceará).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 386 ( Amblyopinodes ; characters; notes; host rodent).

— Martinez, Barrera, and Machado-Allison, 1970: 136 ( Amblyopinodes ; synonym of caviae ).

claviger E. Franz, 1930: 71 ( Amblyopinus ; Type locality: Hansa , Sta. Catharina, Brasilien).

— Scheerpeltz, 1933: 1469 ( Amblyopinus ; catalog).

— Costa Lima, 1936: 58, 59 ( Amblyopinus ; synonym of gahani ).

— Seevers, 1955: 249 ( Amblyopinodes ; characters; notes; Brazil).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 389 ( Amblyopinodes ; characters; notes; host rodent).

- DISTRIBUTION: Brazil.

distinctus Machado-Allison, 1962 , see: piceus Brèthes, 1926 .

gahani Fauvel, 1901b: 6 ( Amblyopinus ; Type locality: La Plata).

— Bernhauer and Schubert, 1916: 441 ( Amblyopinus ; catalog).

— Scheerpeltz, 1933: 1469 ( Amblyopinus ; catalog).

— Costa Lima, 1936: 59 ( Amblyopinus ; characters; notes).

— Seevers, 1955: 248 ( Amblyopinodes ; characters; notes; Argentina).

— Machado-Allison, 1962: 320 ( Amblyopinodes ; characters; Argentina; Brazil).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 392 ( Amblyopinodes ; characters; notes; host rodent).

— Machado-Allison and Barrera, 1964: 176, 178 ( Amblyopinodes ; characters; notes).

— Martinez, Barrera, and Machado-Allison, 1970: 139 ( Amblyopinodes ; subgenus Amblyopinodes ; notes; Argentina).

- DISTRIBUTION: Argentina, Uruguay, Brazil, Peru.

similis Seevers, 1955: 251 ( Amblyopinodes ; Type locality: Peru: Hacienda Cadena, Marcapata , Cuzco, 1000 meters).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 402 ( Amblyopinodes ; characters; notes; host rodent).

— Machado-Allison and Antequera, 1968: 129 ( Amblyopinodes ; subspecies of gahani ; characters; Peru).

— Barrera, 1972: 81 ( Amblyopinodes ; subspecies of gahani ; notes; Peru).

munoai Mañé Garzón and San Martín, 1961: 103 ( Amblyopinodes ; Type locality: Uruguay: Bañado Tropa Vieja , Dto. Canelones).

— San Martin, 1963 ( Amblyopinodes ; characters of male genitalia).

— Machado-Allison, 1963: 396 ( Amblyopinodes ; characters; notes; host rodent).

— Machado-Allison and Barrera, 1964: 177, 178 ( Amblyopinodes ; subspecies of gahani ; characters; notes; Uruguay).

nordestinus Machado-Allison, 1962: 327 ( Amblyopinodes ; Type locality: Brasil: São Benedito , Estado de Ceará).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 398 ( Amblyopinodes ; characters; notes).

— Machado-Allison and Barrera, 1964: 178 ( Amblyopinodes ; subspecies of gahani ; characters; notes).

guimaraesi Machado-Allison, 1962a: 86 ( Amblyopinodes ; Type locality: Brasil: Estado de

São Paulo, Estación Biológica de Boraceia).

— Machado-Allison, 1963: 390 ( Amblyopinodes ; characters; notes; host rodent).

— Martinez, Barrera, and Machado-Allison, 1970: 139 ( Amblyopinodes ; subgenus Amblyopinodes ; characters; notes; Brazil).

- DISTRIBUTION: Brazil.

longus E. Franz, 1930: 74 ( Amblyopinus ; Type locality: Hansa , Sta. Catharina, Brasilien).

— Scheerpeltz, 1933: 1469 ( Amblyopinus ; catalog).

— Costa Lima, 1936: 62 ( Amblyopinus ; characters).

— Seevers, 1955: 234 ( Amblyopinus ; characters; notes; Brazil).

— Martinez, Barrera, and Machado-Allison, 1970: 133 ( Amblyopinodes ; subgenus Streptopinodes ; characters; Brazil).

- DISTRIBUTION: Brazil.

major Machado-Allison and Barrera, 1972: 11 ( Amblyopinodes ; Type locality: Bolivar State, 85 Km SSE El Dorado, 1032 m).

- DISTRIBUTION: Venezuela.

munoai Mañé Garzón and San Martín, 1961, see: gahani Fauvel, 1901 .

nordestinus Machado-Allison, 1962 , see: gahani Fauvel, 1901 .

peruvianus Seevers, 1955: 250 ( Amblyopinodes ; Type locality: Peru: Quince Mil , Cuzco,

680 meters).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 399 ( Amblyopinodes ; characters; notes; host rodent).

— Barrera, 1972: 81 ( Amblyopinodes ; notes; Peru).

- DISTRIBUTION: Peru.

piceus Brèthes, 1926: 18 ( Omaloxenus ; Type locality: Misiones, par Isidoro Escalera ,

de San Ignacio, Argentina).

— Scheerpeltz, 1933: 1469 ( Omaloxenus ; catalog).

— Costa Lima, 1936: 58 ( Amblyopinus ; synonym of gahani ).

— Seevers, 1955: 248 ( Amblyopinodes ; characters; notes; Argentina).

— Machado-Allison, 1962: 313 ( Amblyopinodes ; characters; Brazil; Argentina).

— Machado-Allison, 1962a: 82 ( Amblyopinodes ; characters).

— Machado-Allison, 1963: 399 ( Amblyopinodes ; characters; notes; host rodent; Brazil; Argentina).

- DISTRIBUTION: Argentina, Brazil.

distinctus Machado-Allison, 1962a: 88 ( Amblyopinodes ; subspecies of piceus ; Type locality: Brasil: Estado de Pernambuco, Municipio de Garanhuns).

— Machado-Allison, 1963: 400 ( Amblyopinodes ; subspecies of piceus ; characters; notes; host

rodent).

barrerai Zaragoza Caballero and Hernández, 1993: 28 ( Amblyopinus ; Type locality: Oaxaca: 4.5 km NO Campamento a Santa María Totontepec , 2480 m).

— Ashe and Timm, 1995: 382 ( Amblyopinus ; characters; host cited; Mexico).

- DISTRIBUTION: Mexico.

bequaerti Notman, 1923: 2 ( Omaloxenus ; Type locality: Alto Itatiaya, Setto do Itatiaya, Brazil, on Monodelphis ).

— Scheerpeltz, 1933: 1469 ( Omaloxenus ; catalog).

— Costa Lima, 1936: 58, 64 ( Amblyopinus ; synonym of henseli ).

— Seevers, 1944: 168 ( Amblyopinus ; valid species; characters).

— Seevers, 1955: 247 ( Amblyopinus ).

- DISTRIBUTION: Brazil.

bolivari Barrera, Machado-Allison, and Muñiz, 1960 , see: schmidti Seevers, 1944 .

brandesi Kraatz, 1900: 212 ( Amblyopinus ; Type locality: Peru, Callanga ).

— Fauvel, 1900a: 64 ( Amblyopinus ; synonym of jelskii).

— Bernhauer and Schubert, 1916: 441 ( Amblyopinus ; catalog).

— Costa Lima, 1936: 57, 62 ( Amblyopinus ; synonym of jelskii).

— Seevers, 1955: 227 ( Amblyopinus ; valid species; characters; notes; Peru).

— Barrera, 1972: 79 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

chilomysi Seevers, 1955: 239 ( Amblyopinus ; Type locality: Colombia: Rio Balcones, Guasca , Cundinamarca, 2750 meters).

- DISTRIBUTION: Colombia.

claviger E. Franz, 1930 , see: Amblyopinodes .

colombiae Seevers, 1955: 231 ( Amblyopinus ; Type locality: Colombia: Las Frias, 10 km. east of Sonsón , Antioquia, 2400 meters).

- DISTRIBUTION: Colombia.

delicatus Barrera and Machado-Allison, 1968: 90 ( Amblyopinus ; Type locality: Colombia: departamento del Valle, Calle, Pichinde, La Flora, 1900 m).

- DISTRIBUTION: Colombia.

emarginatus Seevers, 1955: 239 ( Amblyopinus ; Type locality: Colombia: San Antonio ,

San Agustin, Huila).

— Machado-Allison and Barrera, 1964: 175 ( Amblyopinus ; collecting notes; Panama; Venezuela). — Barrera, 1966: 284 ( Amblyopinus ; Panama).

— Barrera, and Machado-Allison, 1968: 96 ( Amblyopinus ; characters; Colombia).

— Machado-Allison and Barrera, 1972: 8 ( Amblyopinus ; Venezuela).

— Ashe and Timm, 1987 ( Amblyopinus ; interaction with rodent host; Costa Rica).

— Ashe and Timm, 1987a ( Amblyopinus ; interaction with rodent host; Costa Rica).

— Timm and Ashe, 1987 ( Amblyopinus ; comments on host and elevational specificity; Panama). — Ashe and Timm, 1995: 376, 394 ( Amblyopinus ; characters; host cited; notes; Colombia; Costa Rica; Panama).

- DISTRIBUTION: Venezuela, Colombia, Panama, Costa Rica.

fuegensis Arrow, 1907: 126 ( Amblyopinus ; Type locality: Tierra del Fuego, Useless Bay).

— Bernhauer and Schubert, 1916: 441 ( Amblyopinus ; cited as fulgensis; catalog).

— Scheerpeltz, 1933: 1469 ( Amblyopinus ; catalog).

— Bruch, 1936: 92 ( Amblyopinus ; probably synonym of jelskii).

— Costa Lima, 1936: 58, 63 ( Amblyopinus ; probably synonym of jelskii).

— Seevers, 1944: 163 ( Amblyopinus ; characters; Chile).

— Seevers, 1955: 237 ( Amblyopinus ; characters; Chile).

— Coiffait and Saiz, 1968: 367 ( Amblyopinus ; Chile).

— Martinez, Barrera, and Machado-Allison, 1970: 128 ( Amblyopinus ; characters; Chile; Argentina).

- DISTRIBUTION: Argentina, Chile.

gahani Fauvel, 1901 , see: Amblyopinodes .

monticolus Seevers, 1955: 228 ( Amblyopinus ; Type locality: Peru: Lago Suche , Moquegua, 14,600 feet).

— Barrera, 1972: 79 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

montivagans Seevers, 1955: 240 ( Amblyopinus ; Type locality: Colombia: San Cristobal, Bogota , Cundinamarca).

- DISTRIBUTION: Colombia.

oryzomysi Seevers, 1955: 240 ( Amblyopinus ; Type locality: Colombia: Las Frias, 10 km. east of Sonson , Antioquia, 2400 meters).

- DISTRIBUTION: Colombia.

pacae Seevers, 1955: 244 ( Amblyopinus ; Type locality: Peru: Marcapata, Cuzco, 3100 meters). — Barrera, 1972: 81 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

piceus Brèthes, 1926 , see: Amblyopinodes .

piurae Seevers, 1955: 232 ( Amblyopinus ; Type locality: Peru: Tambo , Huancabamba, Pirua, 2880 meters).

— Barrera, 1972: 79 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

proximus Machado-Allison and Barrera, 1972: 1 ( Amblyopinus ; Type locality: Amazonas, Cerro Duida, Cabecera del Caño Culebra, 1400 m).

- DISTRIBUTION: Venezuela.

punae Seevers, 1955: 227 ( Amblyopinus ; Type locality: Cailloma , Arequipa, Peru).

— Barrera, 1972: 79 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

sanborni Seevers, 1944: 166 ( Amblyopinus ; Type locality: Peru: Segrario , Puno).

— Seevers, 1955: 238 ( Amblyopinus ; characters; notes; Peru).

— Barrera, 1972: 81 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

schmidti Seevers, 1944: 164 ( Amblyopinus ; Type locality: Guatemala: Santa Elena, Chimal). — Seevers, 1955: 231 ( Amblyopinus ; characters; Guatemala).

— Barrera, Machado-Allison, and Muñiz, 1960: 128 ( Amblyopinus ; characters).

— Machado-Allison and Barrera, 1972: 8 ( Amblyopinus ; notes).

— Ashe and Timm, 1995: 385, 395 ( Amblyopinus ; characters; host cited; notes; Mexico; Guatemala).

- DISTRIBUTION: Guatemala, Mexico.

bolivari Barrera, Machado-Allison, and Muñiz, 1960: 127 ( Amblyopinus ; Type locality: México: D.F.: El Zarco, a 3100 m).

— Machado-Allison and Barrera, 1964: 174 ( Amblyopinus ; collecting notes; Mexico).

— Machado-Allison and Barrera, 1972: 8 ( Amblyopinus ; subspecies of schmidti ).

— Ashe and Timm, 1995: 388, 394 ( Amblyopinus ; subspecies of schmidti ; characters; host cited; Mexico).

seeversi Machado-Allison and Barrera, 1964: 179 ( Megamblyopinus ; Type locality: Venezuela: Serranía del Avila , 2200 m alt.).

— Machado-Allison and Barrera, 1972: 6 ( Amblyopinus ; characters; Venezuela).

- DISTRIBUTION: Venezuela.

spinipennis Seevers, 1955: 234 ( Amblyopinus ; Type locality: Peru: Maracapata, Limapunco , Cuzco).

— Barrera, 1972: 79 ( Amblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

tiptoni Barrera, 1966: 282 ( Amblyopinus ; Type locality: Panama: Bambito , Chiriqui,

5800 feet).

— Barrera, 1966a ( Amblyopinus ; characters; Costa Rica).

— Ashe and Timm, 1987a ( Amblyopinus ; interaction with rodent host; Costa Rica).

— Timm and Ashe, 1987 ( Amblyopinus ; comments on host and elevational specificity; Panama). — Ashe and Timm, 1995: 389, 395 ( Amblyopinus ; characters; host cited; Costa Rica; Panama).

- DISTRIBUTION: Panama, Costa Rica.

trapidoi Barrera and Machado-Allison, 1968: 93 ( Amblyopinus ; Type locality: Colombia: Departamento del Cauca, vecindad del Cerro Munchique, Cordillera Occidental ,

60 Km W Popayán, 2500 m).

- DISTRIBUTION: Colombia.

travassosi Costa Lima, 1927, see: Amblyopinodes .

waterhousei Fauvel, 1900a: 64 ( Amblyopinus ; Type locality: Equateur).

— Bernhauer and Schubert, 1916: 441 ( Amblyopinus ; catalog).

— Costa Lima, 1936: 63 ( Amblyopinus ; characters).

— Seevers, 1944: 165 ( Amblyopinus ; characters; Colombia).

— Seevers, 1955: 243 ( Amblyopinus ; characters; Colombia).

— Machado-Allison and Barrera, 1964: 176 ( Amblyopinus ; Ecuador; Colombia; Venezuela; collecting notes).

— Barrera, and Machado-Allison, 1968: 98 ( Amblyopinus ; notes; Colombia).

— Machado-Allison and Barrera, 1972: 2 ( Amblyopinus ; Venezuela).

- DISTRIBUTION: Venezuela, Colombia, Ecuador.

wenzeli Martínez, Barrera, and Machado-Allison, 1970: 130 ( Amblyopinus ; Type locality: Brasil: Santa Catarina , Xixim).

- DISTRIBUTION: Brazil.

Chilamblyopinus

[1 species; Neotropical Region]

Chilamblyopinus Ashe and Timm, 1988: 49 (species included: piceus ). TYPE SPECIES: Chilamblyopinus piceus Ashe and Timm , fixed by monotypy and original designation.

piceus Ashe and Timm, 1988: 52 ( Chilamblyopinus ; Type locality: Chile: Osorno Prov. , Valle de La Picada).

- DISTRIBUTION: Chile.

Edrabius

[11 species; Neotropical Region]

Edrabius Fauvel, 1900a: 63 , 65 (species included: philippianus ). TYPE SPECIES: Edrabius philippianus Fauvel , fixed by monotypy.

— Bernhauer and Schubert, 1916: 441 (world catalog; 1 species).

— Bruch, 1928a: 429 (catalog; Argentina).

— Scheerpeltz, 1933: 1469 (world catalog supplement).

— Bruch, 1936: 94 (characters).

— Blackwelder, 1944: 150 (checklist of species from Mexico, Central and South America, and

the West Indies).

— Blackwelder, 1952: 145 (type species: philippianus ).

— Seevers, 1955: 254 (characters; notes; type species: philippianus ).

— Coiffait and Saiz, 1968: 365, 366 (characters; key to Chilean species).

— Ashe and Timm, 1988: 48 (characters).

— Timm and Ashe, 1989 (larval characters for an undescribed species; comments on systematic position).

— Ashe, Timm, and Gallardo, 1996 (summary of distribution of species; host associations). alticolus Seevers, 1955: 259 ( Edrabius ; Type locality: Peru: Pampa de Titire, 29 km. northeast of Tarata, Tacna, 14,600 feet).

— Barrera, 1972: 82 ( Edrabius ; notes; Peru).

— Ashe, Timm and Gallardo, 1996: 738 ( Edrabius ; host; Bolivia).

- DISTRIBUTION: Peru, Bolivia.

argentinus Seevers, 1955: 258 ( Edrabius ; Type locality: Argentina: Hualfin , Catamarca, near Tucuman).

— Coiffait and Saiz, 1968: 366 ( Edrabius ; characters; Chile).

— Ashe, Timm and Gallardo, 1996: 739 ( Edrabius ; host; notes on variation; Argentina).

- DISTRIBUTION: Argentina, Chile.

australis Ashe, Timm and Gallardo, 1996: 735 ( Edrabius ; Type locality: Chile: Bío-Bío Prov.: Laguna Laja ; also host cited).

- DISTRIBUTION: Chile.

chilensis Scheerpeltz, 1957a: 222 ( Edrabius ; Type locality: Chile: Hoch-Anden , Curico,

Las Tablas).

— Ashe, Timm and Gallardo, 1996: 739 ( Edrabius ; host; notes on variation; Chile).

- DISTRIBUTION: Chile.

chilensiformis Ashe, Timm and Gallardo, 1996: 737 ( Edrabius ; Type locality: Chile: coquimob Prov.: 10 km N Puente Los Molles , 32 ° 09 ̍ S, 71 ° 31 ̍ W; also host cited).

- DISTRIBUTION: Chile.

grandis Ashe, Timm and Gallardo, 1996: 733 ( Edrabius ; Type locality: Argentina: Prov. Río Negro : 13 km WSW Comallo, el. 1150 m. Also cited from Chile; also host cited).

- DISTRIBUTION: Argentina, Chile.

kuscheli Scheerpeltz, 1957a: 220 ( Edrabius ; Type locality: Chile: Hoch-Anden , Arica-Lipiche, 4400 m).

— Coiffait and Saiz, 1968: 366, 367 ( Edrabius ; characters; Chile).

- DISTRIBUTION: Chile.

pearsoni Seevers, 1955: 258 ( Edrabius ; Type locality: Peru: Rio Santa Rosa , Puno,

13,300 feet).

— Barrera, 1972: 82 ( Edrabius ; notes; Peru).

- DISTRIBUTION: Peru.

peruanus Seevers, 1955: 259 ( Edrabius ; Type locality: Peru: Pichupichuni, 8 km. north of Huacullani , Puno, 12,600 feet).

— Barrera, 1972: 82 ( Edrabius ; notes; Peru).

— Ashe, Timm and Gallardo, 1996: 739 ( Edrabius ; host; Chile).

- DISTRIBUTION: Peru, Chile.

philippianus Fauvel, 1900a: 65 ( Edrabius ; Type locality: Chili: desert d’Atacama, Antofagasta).

— H. Kolbe, 1911: 117 ( Edrabius ; notes).

— Bernhauer and Schubert, 1916: 441 ( Edrabius ; catalog).

— Scheerpeltz, 1933: 1469 ( Edrabius ; catalog).

— Bruch, 1936: 93, 94 ( Edrabius ; charactes; illustrations).

— Costa Lima, 1936: 65, 66 ( Edrabius ; characters).

— Seevers, 1955: 255 ( Edrabius ; characters; notes; Chile).

— Scheerpeltz, 1957a: 219 ( Edrabius ; characters; Chile; Argentina).

— Coiffait and Saiz, 1968: 366, 367 ( Edrabius ; characters; Chile).

— Ashe, Timm and Gallardo, 1996: 739 ( Edrabius ; type host identification corrected).

- DISTRIBUTION: Chile, Argentina.

weiseri Seevers, 1955: 255 ( Edrabius ; Type locality: Argentina: Troquero , Jujuy).

— Ashe, Timm and Gallardo, 1996: 739 ( Edrabius ; host; Bolivia).

- DISTRIBUTION: Argentina, Bolivia.

Megamblyopinus

[3 species; Neotropical Region]

Megamblyopinus Seevers, 1955: 251 (species included: mniszechi , germaini ). TYPE SPECIES: Amblyopinus mniszechi Solsky , fixed by original designation.

— Machado-Allison and Barrera, 1964: 181 (characters).

— Coiffait and Saiz, 1968: 366, 367 (characters).

— Ashe and Timm, 1988: 48 (characters).

chilensis Coiffait and Saiz, 1968: 367 ( Megamblyopinus ; Type locality: Chili: Camarones ).

- DISTRIBUTION: Chile.

germaini Fauvel, 1900a: 63 ( Amblyopinus ; Type locality: Pérou).

— Bernhauer and Schubert, 1916: 441 ( Amblyopinus ; catalog).

— Bruch, 1936: 57 ( Amblyopinus ).

— Bruch, 1936: 58 ( Amblyopinus ).

— Seevers, 1955: 254 ( Megamblyopinus ; characters; notes; Peru).

— Barrera, 1972: 82 ( Megamblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

mniszechi Solsky, 1875: 11 ( Amblyopinus ; Type locality: Montana de Chanchamago, haienda Amable Maria).

— Fauvel, 1900a: 63 ( Amblyopinus ; Peru).

— H. Kolbe, 1911: 117 ( Amblyopinus ; notes).

— Bernhauer and Schubert, 1916: 441 ( Amblyopinus ; catalog).

— Scheerpeltz, 1933: 1469 ( Amblyopinus ; catalog).

— Bruch, 1936: 58 ( Amblyopinus ; characters).

— Seevers, 1955: 251 ( Megamblyopinus ; characters; notes; Peru).

— Barrera, 1972: 82 ( Megamblyopinus ; notes; Peru).

- DISTRIBUTION: Peru.

seeversi Machado-Allison and Barrera, 1964 , see: Amblyopinus .

Myotyphlus

[1 species; Australian Region]

Myotyphlus Fauvel, 1883: 40 (species included: jansoni ). TYPE SPECIES: Amblyopinus jansoni A. Matthews , fixed by monotypy.

— Duvivier, 1883: 137 (catalog).

— Bernhauer and Schubert, 1916: 440 (world catalog; 1 species).

— Scheerpeltz, 1933: 1468 (world catalog supplement).

— Blackwelder, 1952: 253 (type species: jansoni ).

— Seevers, 1955: 259 (type species: jansoni ).

— Hamilton-Smith and Adams, 1966 (may be in Quediini ).

— Ashe and Timm, 1988: 48 (characters).

Cryptommatus A. Matthews, 1884: 88 (species included: jansoni ). TYPE SPECIES: Amblyopinus jansoni A. Matthews , fixed by monotypy.

— Bernhauer and Schubert, 1916: 440 (synonym of Myotyphlus ).

— Blackwelder, 1952: 113 (synonym of Myotyphlus ; type species: jansoni ).

jansoni A. Matthews, 1878: 278 ( Amblyopinus ; Type locality: Tasmania).

— Fauvel, 1883: 40 ( Myotyphlus ).

— A. Matthews, 1884: 91 ( Cryptommatus ; characters; Tasmania) .

— H. Kolbe, 1911: 117 ( Myotyphlus ; notes).

— Bernhauer and Schubert, 1916: 440 ( Myotyphlus ; catalog).

— Lea, 1925: 253 ( Cryptommatus ; characters; Australia).

— Seevers, 1955: 260 ( Myotyphlus ; characters; notes; Australia).

— Hamilton-Smith and Adams, 1966 ( Myotyphlus ; not an obligate ectoparasite; found in guano

in bat cave; Victoria).

- DISTRIBUTION: Australia.

Subtribe Anisolinina

Anisolinina Hayashi, 1993b: 284 (genera included: Anisolinus , Amichrotus , Tympanophorus , Hesperosoma ). TYPE GENUS: Anisolinus , by original designation.

— Smetana and Davies, 2000: 12, 15 (characters).

Amichrotus

[11 species; Palaearctic and Oriental Regions]

Amichrotus Sharp, 1889: 114 (species included: apicipennis ). TYPE SPECIES: Amichrotus apicipennis Sharp , fixed by monotypy.

— Bernhauer and Schubert, 1914: 367 (world catalog; 4 species).

— Winkler, 1925: 381 (catalog for Palaearctic region).

— Cameron, 1925: 64 (catalog of Indian species).

— Scheerpeltz, 1933: 1376 (world catalog supplement).

— Blackwelder, 1952: 49 (type species: apicipennis ).

— Sawada, 1961: 4 (characters in key).

— Shibata, 1973c: 63 (checklist of species of Taiwan).

— Tikhomirova, 1973a: 186 (checklist of species of USSR).

— Naomi, 1982: 128 (characters).

— Naomi, 1982a: 37 (characters; key to subgenera).

— Naomi, 1983: 49 (characters; species of Japan).

— Hammond, 1984: 205 (checklist of species of Borneo).

— Shibata, 1984: 109 (checklist of species of Japan).

— Shibata, 1986a: 125 (supplemental list of species from Taiwan).

— Hayashi, 2000: 24 (notes on empodial setae).

apicipennis Sharp, 1889: 115 ( Amichrotus ; Type locality: Miyanoshita to Junsai).

— Bernhauer and Schubert, 1914: 367 ( Amichrotus ; catalog).

— Cameron, 1949b: 468 ( Amichrotus ; China, Japan, Formosa).

— Nakane, 1963a: 92 ( Amichrotus ; characters; habitus photograph; Japan).

— Frank, 1982: 33 ( Amichrotus ; list of parasites).

— Naomi, 1983: 51 ( Amichrotus ; characters; Japan).

— J. Li, 1993: 41 ( Amichrotus ; China).

- DISTRIBUTION: Taiwan, China, Japan.

bryanti Cameron, 1937: 23 ( Thoracostrongylus ; Type locality: Java ; Borneo: Quop, W. Sarawak).

— Hammond, 1984: 194, 195 ( Amichrotus ; Borneo).

- DISTRIBUTION: Borneo, Indonesia.

costatus Fauvel, 1895 , see: Thoracostrongylus .

distinctus Cameron, 1932: 215 ( Thoracostrongylus ; Type locality: Burma: Ruby Mines ).

— Scheerpeltz, 1933: 1407 ( Thoracostrongylus ; catalog).

— Hammond, 1984: 194, 195 ( Amichrotus ; Borneo).

- DISTRIBUTION: Myanmar, Borneo.

doriae Gridelli, 1924f: 207 ( Ontholestes ; subgenus Thoracostrongylus ; Type locality: Borneo, Sarawak).

— Scheerpeltz, 1933: 1407 ( Thoracostrongylus ; catalog).

— Hammond, 1984: 195 ( Amichrotus ; Borneo).

- DISTRIBUTION: Borneo.

elegans Cameron, 1920a: 218 ( Amichrotus ; Type locality: Sidapur, Coorg).

— Cameron, 1932: 214 ( Thoracostrongylus ; characters; India).

— Scheerpeltz, 1933: 1407 ( Thoracostrongylus ; catalog).

— Cameron, 1933d: 391 ( Thoracostrongylus ; Sumatra).

— Hammond, 1984: 195 ( Amichrotus ; Borneo).

— Biswas and Biswas, 1985: 146 ( Thoracostrongylus ; India).

- DISTRIBUTION: India, Indonesia, Borneo.

excellens Bernhauer, 1939d: 100 ( Amichrotus ; Type locality: Japan: Kamikochi ).

— Naomi, 1983: 52 ( Amichrotus ; notes).

— Hayashi, 1995: 47 ( Amichrotus ; characters; review of type).

- DISTRIBUTION: Japan.