Deltochilum Eschscholtz, 1822

Rosa, Cecilia Lozano De La, Cupello, Mario & Vaz-De-Mello, Fernando Z., 2024, The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution, European Journal of Taxonomy 959 (1), pp. 1-272 : 70-72

Genus Deltochilum Eschscholtz, 1822

Deltochilum Eschscholtz, 1822:37 (original description). Type species: Deltochilum dentipes Eschscholtz, 1822 , by original monotypy. Note: Bouchard et al. (2024) recently stated that D. dentipes was fixed as the type species of Deltochilum by the subsequent designation of Thon (1832), but this is incorrect. Deltochilum dentipes was the sole nominal species that Eschscholtz originally included in the genus without hesitation. It is true that he also said that Ateuchus gibbosus ( Fabricius, 1775) (currently, Deltochilum gibbosum ) was “closely related” to D. dentipes and “seems to also belong to the genus” (“[...] welcher ebenfalls zu dieser Gattung zu gehören scheint”). However, because this was at most a doubtful assignment of the species to Deltochilum , gibbosus cannot be considered a second originally included nominal species per Article 67.2.5 of the Code ( ICZN 1999) and has, therefore, never been eligible for type fixation. Deltochilum dentipes , having been the only originally included nominal species, was automatically fixed as the type species by original monotypy (Art. 68.3). After we brought this to his attention, Patrice Bouchard (pers. com. to MC, 12 th June 2024) now agrees with us.

Anamnesis Vigors, 1826: 510 (original description). Type species: Anamnesis macleayii Vigors, 1826 , by original monotypy.

Meghyboma Kolbe, 1893: 192 (original description). Type species: Deltochilum dentipes Eschscholtz, 1822 , by subsequent designation of Shipp (1897).

Telhyboma Kolbe, 1893: 192 (original description). Type species: Deltochilum orbiculare Van Lansberge, 1874 , by original monotypy.

Deltochilum – Agassiz 1846: 341 (catalogue). — Burmeister 1848: 134 (redescription); 1873: 408 (redescription). — Lacordaire 1855: 79 (redescription). — LeConte 1861: 125 (key). — Harold 1869d: 995 (catalogue, distribution). — Van Lansberge 1874: 188 (characteristics). — Kolbe 1893: 191 (redescription). — Shipp 1897: 194 (comments). — Gillet 1911b: 35 (catalogue). — Lucas 1920: 228 (catalogue, distribution). — Paulian 1938: 235, 237 (key, redescription). — Pessôa & Lane 1941: 411, 426 (key, redescription). — Blackwelder 1944: 202 (list). — Lane 1946: 171 (comments). — Roze 1955: 43 (checklist for Venezuela). — Pereira & Martínez 1956a: 96 (key). — Martínez 1959: 50 (catalogue for Argentina). — Halffter 1961: 231 (key); 2003: 26 (redescription). — Vulcano & Pereira 1964: 639 (catalogue). — Halffter & Matthews 1966: 261 (catalogue, distribution). — Vulcano & Pereira 1967: 549 (key). — Halffter & Martínez 1977: 36 (key). — Howden & Young 1981: 14, 36 (key, redescription). — Halffter & Edmonds 1982: 139 (catalogue, distribution). — Medina& Lopera-Toro 2000: 311 (key). — Vaz-de-Mello 2000: 192 (checklist for Brazil). — Medina et al. 2001: 136 (checklist for Colombia). — Ratcliffe et al. 2002: 49 (key); 2015: 195 (checklist for Peru). — Ratcliffe 2002: 13 (checklist for Panama). — Medina et al. 2003: 64 (distribution). — Hamel-Leigue et al. 2006: 14 (list). — Vaz-de-Mello et al. 2011a: 25 (key). — Carvajal et al. 2011: 123, 316 (diagnosis, list). — Krajcik 2012: 88 (list). — Solís & Kohlmann 2012: 4 (checklist for Costa Rica). — Boilly & Vaz-de-Mello 2013: 107 (key, characteristics). — Chamorro et al. 2018: 76, 93–94 (list for Ecuador); 2019: 9, 82 (catalogue). — Hielkema & Hielkema 2019: 74 (catalogue for the Guianas).

Anamnesi – Agassiz 1846: 58 (catalogue). — Harold 1869d: 995 (cited as synonym, cited as Annamesis). — Shipp 1897: 194 (comments, cited as synonym of Deltochilum ). — Gillet 1911b: 35 (cited as synonym of Deltochilum , cited as Annamesis). — Paulian 1938: 237 (cited as synonym of Deltochilum , cited as Annamesis). — Blackwelder 1944: 202 (cited as synonym of Deltochilum , cited as Annamesis). — Lane 1946: 171 (comments). — Pereira & Martínez 1956a: 120 (cited as synonym of Deltochilum ). — Martínez 1959: 50 (cited as synonym of Deltochilum ). — Vulcano & Pereira 1964: 639 (cited as synonym of Deltochilum ). — Ratcliffe 2002: 13 (cited as synonym of Deltochilum ). — Solís & Kohlmann 2012: 4 (cited as synonym of Deltochilum , cited as Annamesis).

Meghyboma – Shipp 1897: 194 (comments, cited as synonym of Deltochilum ). — Paulian 1938: 243, 246 (key, review). — Blackwelder 1944: 202 (cited synonym of Deltochilum ). — Lane 1946: 174 (cited synonym of Deltochilum ). — Halffter & Edmonds 1982: 139 (cited as subgenus of Deltochilum ). — Solís & Kohlmann 2012: 4 (cited as synonym of Deltochilum ).

Telhyboma – Paulian 1938: 219–220, 224 (cited, comments). — Génier 2012: 25–26, 30–31 (revision, new synonymy). — Gonzalez-Alvarado & Vaz-de-Mello 2021a: 2‒3 (revision).