Aname mellosa, Harvey, Frances S. B., Framenau, Volker W., Wojcieszek, Janine M., Rix, Michael G. & Harvey, Mark S., 2012

Harvey, Frances S. B., Framenau, Volker W., Wojcieszek, Janine M., Rix, Michael G. & Harvey, Mark S., 2012, Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia, Zootaxa 3383, pp. 15-38 : 22-30

Aname mellosa n. sp.

Figs 1-5, 9- 24

Type material. AUSTRALIA: Western Australia: holotype male, Jinayri, ca. 60 km NW. of Newman , 22°58 ’05.7” S, 119°17 ’06.1” E, 3 February 2009, pitfall trap, J. Gollan, N. Sullivan, M. Semeniuk, M. Beatson ( WAM T94870 ). Paratypes: 1 male, same data as holotype except 22°57 ’46.1” S, 119°13 ’36.3” E, 6 February 2009 ( WAM T96116 ) ; 2 males, same data except 22°57 ’46.5” S, 119°13 ’36.5” E, 7 February 2009 ( WAM T96121 , T94864 ) ; 1 male, same data except 22°57 ’47.4” S, 119°13 ’37.1” E, 6 February 2009 ( WAM T96122 ) ; 1 male, same data except 22°57 ’47.4” S, 119°13 ’37.1” E, 3 February 2009 ( QM S91581 ) ; 1 male, same data except 22°57 ’48.7” S, 119°13 ’38.1” E, 5 February 2009 ( WAM T94881 ) ; 1 male, same data except 22°58 ’03.2” S, 119°17 ’06.2” E, 5 February 2009 ( WAM T96113 ) ; 1 male, same data except 22°58 ’04.0” S, 119°17 ’06.4” E, 5 February 2009 ( WAM T94871 DNA ) ; 1 male, same data except 22°58 ’05.0” S, 119°17 ’06.2” E, 5 February 2009 ( WAM T96117 ) ; 2 males, same data except 22°58 ’09.5” S, 119°15 ’35.1” E, 3 February 2009 ( WAM T94885, T94872 ) ; 1 male, same data except 22°58 ’09.5” S, 119°15 ’36.2” E, 7 February 2009 ( WAM T96128 ) ; 2 males, same data except 22°58 ’09.5” S, 119°15 ’36.2’Έ, 1 February 2009 ( WAM T94884 DNA , T94882 ) ; 1 male, same data except 22°58 ’09.5” S, 119°15 ’36.4” E, 5 February 2009 ( WAM T94857 ) ; 1 male, same data except 22°58 ’09.5” S, 119°15 ’36.4” E, 6 February 2009 ( WAM T94862 ) ; 2 males, same data except 22°58 ’09.8” S, 119°15 ’36.5” E, 7 February 2009 ( WAM T96144 , T96126 ) ; 1 male, same data except 22°58 ’10.2” S, 119°15 ’37.1” E, 6 February 2009 ( WAM T94863 ) ; 1 male, same data except 22°58 ’26.4” S, 119°18 ’00.8” E, 6 February 2009 ( WAM T94874 ) ; 1 female, same data except 22°58 ’26.9” S, 119°18 ’03.1” E, 1 February 2009 ( WAM T96145 ) ; 2 males, same data except 22°58 ’26.9” S, 119°18 ’03.1” E, 6 February 2009 ( WAM T94856 , T94860 ) ; 1 male, same data except 22°58 ’27.0” S, 119°18 ’04.0” E, 7 February 2009 ( WAM T94886 ) ; 1 male, same data except 22°58 ’42.6” S, 119°15 ’39.8” E, 5 February 2009 ( WAM T96129 ) ; 1 male, same data except 22°58 ’42.9” S, 119°15 ’41.2” E, 7 February 2009 ( WAM T94859 ) ; 1 male, same data except 22°58 ’43.5” S, 119°15 ’40.5” E, 7 February 2009 ( WAM T96123 ) ; 1 male, same data except 22°59 ’03.3” S, 119°16 ’03.3” E, 5 February 2009 ( WAM T96114 ) ; 1 male, same data except 22°59 ’03.7” S, 119°18 ’03.8” E, 2 February 2009 ( WAM T94879 DNA ) ; 1 male, same data except 22°59 ’04.5” S, 119°16 ’02.3” E, 7 February 2009 ( WAM T94866 ) ; 1 male, same data except 22°59 ’04.5” S, 119°16 ’02.3” E, 6 February 2009 ( WAM T94868 ) ; 2 males, same data except 22°59 ’08.6” S, 119°15 ’02.8” E, 7 February 2009 ( WAM T94854 , T94861 ) ; 1 male, same data except 22°59 ’08.8” S, 119°15 ’03.1” E, 7 February 2009 ( WAM T94869 ) ; 2 males, same data except 22°59 ’09.2” S, 119°15 ’04.4” E, 7 February 2009 ( WAM T94853 , T96118 ) ; 1 male, same data except 22°59 ’09.5” S, 119°15 ’35.1” E, 7 February 2009 ( WAM T94867 ) ; 1 male, same data except 22°59 ’14.1” S, 119°16 ’09.6” E, 6 February 2009 ( WAM T94858 ) ; 1 male, same data except 22°59 ’14.9” S, 119°16 ’10.3” E, 7 February 2009 ( WAM T96119 ) ; 1 male, same data except 22°59 ’15.2” S, 119°16 ’11.3” E, 6 February 2009 ( WAM T96125 ) ; 1 male, same data except 22°59 ’19.9” S, 119°14 ’11.3” E, 8 February 2009 ( WAM T96127 ) ; 1 male, same data except 22°59 ’21.3” S, 119°14 ’10.7” E, 6 February 2009 ( WAM T94865 ) ; 1 male, same data except 22°59 ’23.1” S, 119°14 ’11.1” E, 8 February 2009 ( WAM T94851 ) ; 1 male, same data except 22°59 ’23.8” S, 119°14 ’11.5” E, 7 February 2009 ( WAM T96124 ) ; 1 male, same data except 22°59 ’23.8” S, 119°14 ’11.5” E, 6 February 2009 ( WAM T96120 ) ; 1 male, same data except 22°59 ’24.5” S, 119°14 ’12.0” E, 5 February 2009 ( WAM T94883 ) ; 1 male, same data except 23°00 ’23.3” S, 119°14 ’42.6” E, 7 February 2009 ( WAM T94876 ) ; 1 male, same data except 23°00 ’23.9” S, 119°14 ’43.6” E, 5 February 2009 ( WAM T96151 ) ; 1 male, same data except 23°00 ’39.8” S, 119°14 ’09.0” E, 7 February 2009 ( WAM T96115 ) ; 1 male, same data except 23°00 ’49.2” S, 119°14 ’13.9” E, 6 February 2009 ( WAM T94852 ) ; 1 male, same data except 23°01 ’29.0” S, 119°14 ’37.8” E, 3 February 2009 ( WAM T96140 DNA ) ; 1 male, same data except 23°01 ’29.4” S, 119°14 ’37.0” E, 4 February 2009 ( WAM T94875 ) ; 1 male, same data except 23°02 ’26.0” S, 119°16 ’45.2” E, 2 February 2009 ( WAM T94880 DNA ) ; 2 males, same data except 23°03 ’08.8” S, 119°16 ’14.9” E, 3 February 2009 ( WAM T94873 , T94878 ) ; 1 male, same data except 23°03 ’09.8” S, 119°16 ’13.9” E, 5 February 2009 ( WAM T94850 ) ; 1 female, same data as holotype except 22°59 ’24.5” S, 119°14 ’12.0” E, 8 February 2009 ( WAM T96150 ) .

Other material examined. AUSTRALIA: Western Australia: 1 male, Abydos mine lease, ca. 64 km W. of Marble Bar , 21°07 ’39.7” S, 119°08 ’03.2” E, 15 May 2010, dry pitfall trap, P. Bolton ( WAM T 102291 ) ; 1 juvenile, same data except 21°07 ’50.2” S, 119°07 ’32.7” E, 14 May 2010 ( WAM T 102290 ) ; 1 male, same data except 21°07 ’53” S, 119°08 ’07.2” E, 4-10 May 2010 ( WAM T 102287 ) ; 1 male, same data except 5-10 May 2010 ( WAM T 102288 ) ; 1 male, same data except 21°07 ’54.9” S, 119°07 ’54” E, 28-5 May 2010, wet pitfall trap ( WAM T 102284 ) ; 1 male, API Hardey, 96.9 km WNW. of Paraburdoo , 22°49 ’04” S, 116°48 ’53” E, 24 March 2010, dug from burrow, P. Runham ( WAM T 110361 ) ; 1 male, Aquila, 12.75 km SE. of Karratha , 20°48 ’34” S, 116°56 ’29” E, 8 April 2009, dug from burrow, P. Runham ( WAM T100079 ) ; 1 male, same data except 5 April 2009 ( WAM T100080 ) ; 1 male, Aquila, 20.6 km E. of Karratha , 20°45 ’45” S, 117°02 ’31” E, 5 April 2009, dug from burrow, P. Runham ( WAM T100077 ) ; 1 male, same data except 7 April 2009 ( WAM T100078 ) ; 1 male, Area C, 101 km NW. of Newman , 23°00 ’20” S, 118°49 ’38” E, 25 February 2010, dug from burrow, open mulga woodland on clayey loam plain, M. Greenham ( WAM T 101158 ) ; 1 female, same data except D. Kamien, T. Sachse ( WAM T 101162 ) ; 1 male, Area C, 61.8 km NW. of Newman , 22°59 ’55” S, 118°52 ’07” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105930 ) ; 1 male, same data ( WAM T105931 ) ; 1 male, Area C, 74.1 km NW. of Newman , 22°55 ’04” S, 119°11 ’21” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105957 ) ; 1 female, same data except 22°55 ’05” S, 119°11 ’21” E, 18 February 2010, dug from burrow, drainage in valley/flats, D. Kamien ( WAM T 101163 ) ; 1 female, Area C, 75.7 km NW. of Newman, 22°54 ’21” S, 119°10 ’48” E, 19 February 2010, dug from burrow, loamy plain, D. Kamien, T. Sachse ( WAM T 101165 ) ; 1 male, Area C, 75.8 km NW. of Newman , 22°54 ’21” S, 119°10 ’48” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105943 ) ; 1 male, Area C, 78.9 km NW. of Newman , 22°54 ’10” S, 119°08 ’38” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105958 ) ; 1 male, Area C, 81.4 km NW. of Newman , 22°57 ’29” S, 119°04 ’09” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105914 ) ; 1 male, Area C, 81.8 km NW. of Newman , 23°00 ’20” S, 119°01 ’58” E, 21 February 2010, dug from burrow, plateau/top of rocky hill, M. Greenham ( WAM T 101156 ) ; 1 female, Area C, 82.6 km NW. of Newman , 22°58 ’54” S, 119°00 ’49” E, 20 February 2010, dug from burrow, Gully, Z. Hamilton ( WAM T 101161 ) ; 1 male, same data except 22°58 ’55” S, 119°00 ’50” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105921 ) ; 1 male, Area C, 82.7 km NW. of Newman , 22°53 ’57” S, 119°05 ’58” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105954 ) ; 1 male, Area C, 82.7 km NW. of Newman , 22°57 ’49” S, 119°02 ’50” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105910 ) ; 1 female, Area C, 82.7 km NW. of Newman , 22°57 ’53” S, 119°02 ’53” E, 20 February 2010, dug from burrow, mulga plain, M. Greenham ( WAM T 101164 ) ; 1 male, Area C, 82.7 km NW. of Newman , 23°00 ’18” S, 119°01 ’26” E, 1 May 2010, wet pit trap, S. Ford, G. O’Con- ( WAM T105960 ) ; 1 male, Area C, 83.1 km NW. of Newman , 22°58 ’30” S, 119°02 ’16” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105956 ) ; 1 male, same data ( WAM T105967 ) ; 1 male, Area C, 83.9 km NW. of Newman , 22°59 ’35” S, 119°00 ’57” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105932 ) ; 1 male, Area C, 84.7 km NW. of Newman , 22°54 ’21” S, 119°01 ’44” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105950 ) ; 1 male, same data ( WAM T105974 ) ; 1 male, Area C, 85.2 km NW. of Newman , 23°00 ’14” S, 118°59 ’41” E, 22 February 2010, dug from burrow, Z. Hamilton ( WAM T 101154 ) ; 1 male, Area C, 86.2 km NW. of Newman , 23°00 ’05” S, 118°58 ’54” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105928 ) ; 1 male, Area C, 87.2 km NW. of Newman , 22°59 ’22” S, 118°58 ’58” E, 22 February 2010, dug from burrow, Loamy clay plain, D. Kamien ( WAM T 101155 ) ; 1 male, Area C, 89 km NW. of Newman , 22°57 ’23” S, 118°59 ’00” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105911 ) ; 1 male, same data ( WAM T105912 ) ; 1 male, same data ( WAM T105975 ) ; 1 male, Area C, 89.2 km NW. of Newman, 22°57 ’56” S, 118°58 ’32” E, 23 February 2010, dug from burrow, rocky loam plain, D. Kamien, T. Sachse ( WAM T 101157 ) ; 1 male, Area C, 89.4 km NW. of Newman , 22°58 ’49” S, 118°57 ’44” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105925 ) ; 1 male, same data ( WAM T105944 ) ; 1 male, same data ( WAM T105959 ) ; 1 male, Area C, 89.6 km NW. of Newman , 22°54 ’31” S, 119°00 ’26” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105908 ) ; 1 male, same data ( WAM T105976 ) ; 1 female, Area C, 89.6 km NW. of Newman , 23°00 ’03” S, 118°56 ’49” E, 24 February 2001, dug from burrow, Rocky gully, Z. Hamilton ( WAM T 101160 ) ; 1 male, Area C, 90.6 km NW. of Newman , 22°53 ’56” S, 119°00 ’07” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105934 ) ; 1 male, Area C, 90.6 km NW. of Newman , 22°59 ’52” S, 119°01 ’47” E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T105915 ) ; 1 male, Area C, 92.7 km NW. of Newman , 23°01 ’04” S, 118°54 ’24” E, 24 February 2010, dug from burrow, open mulga woodland with scattered eucalypts, M. Greenham ( WAM T 101153 ) ; 1 female, Area C, 98.2 km NW. of Newman , 23°00 ’28” S, 118°51 ’12” E, 21 February 2010, dug from burrow, open mulga woodland on clayey loam plain, M. Greenham ( WAM T 101166 ) ; 1 male, 36 km NW. of Balfour Downs Homestead, Pilbara Biological Survey site BDRN11 , 22°30 ’59” S, 120°39 ’34” E, 11 September 2005 - 14 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T109337 ) ; 1 male, Bungaroo, 17.1 km S. of Pannawonica , 21°47 ’04” S, 116°15 ’31” E, 14 March 2010, dug from burrow, J. Alexander ( WAM T 102531 ) ; 1 male, Bungaroo , same data except dry pit trap, J. Alexander ( WAM T 102533 ) ; 1 juvenile, same data except dug from burrow, D. Keirle ( WAM T 102536 ) ; 1 male, Bungaroo, 19.5 km S. of Pannawonica , 21°51 ’00” S, 116°22 ’12” E, 15 March 2010, dry pit trap, J. Alexander ( WAM T 102530 ) ; 1 male, same data except 13 March 2010, D. Keirle ( WAM T 102532 ) ; 1 male, same data ( WAM T 102535 ) ; 1 male, Bungaroo, 22.1 km S. of Pannawonica , 21°50 ’30” S, 116°21 ’31” E, 13 March 2010, Dry pit trap, D. Keirle ( WAM T 102534 ) ; 1 male, Bungaroo, 26.9 km S. of Pannawonica, site BUN8 , 21°49 ’08” S, 116°17 ’59” E, 14-20 March 2005, dry pitfall traps, Z. Hamilton ( WAM T72405 ) ; 1 male, Charles Darwin Reserve, 2.5 km N. of White Wells Homestead , 29°33 ’29” S, 116°57 ’52” E, 4-5 March 2008, gas lantern/pitfall, mixed shrubland with emergent eucalypts and native pine, T.F. Houston ( WAM T88785 ) ; 2 males, 9 km NE. of Cowra Line Camp, Pilbara Biological Survey site RHNW11 , 22°17 ’38.8” S, 119°03 ’40.9” E, 27 August 2003 - 20 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109348 ) ; 1 male, 16 km N. of Cowra Line Camp, Pilbara Biological Survey site RHNW07 , 22°13 ’18.2” S, 119°01 ’29.5” E, 14 August 2003 - 20 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109353 ) ; 1 male, Davidson Creek, ~ 75 km E. of Newman, vert site 1 , 23°31 ’29.81” S, 120°38 ’28.26” E, 8 April 2010, dry pitfall, J. Clark ( WAM T 102154 ) ; 1 male, Davidson Creek, ~ 75 km E. of Newman, vert site 3 , 23°28 ’27.25” S, 120°34 ’15.86” E, 8 April 2010, Dry pitfall, J. Clark ( WAM T 102156 ) ; 1 male, Davidson Creek ~ 75 km E. of Newman, vert site 5 , 23°25 ’15.91” S, 120°28 ’59.56” E, 9 April 2010, dry pitfall, J. Clark ( WAM T 102155 ) ; 1 male, Fortescue Marsh , 22°17 ’32” S, 119°04 ’10” E, 2-5 June 2010, dry pitfall, leaf litter, P. Roberts, L. Quinn ( WAM T107179 ) ; 1 male, same data ( WAM T107180 ) ; 1 male, Fortescue Marsh , 22°18 ’26” S, 119 °12 ’57” E, 2-5 June 2010, dry pitfall, leaf litter, P. Roberts, L. Quinn ( WAM T107182 ) ; 1 male, 67.5 km S. of Fortescue River crossing on Dampier Paraburdoo Railway, Pilbara Biological Survey site PE12 , 22°03 ’33.2” S, 117°41 ’36.5” E, 5 September 2005 - 11 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T106274 ) ; 1 male, 7.4 km SW. of Fortescue Roadhouse , 21°20 ’58” S, 116°05 ’46” E, 24 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T88702 ) ; 1 male, 13.1 km SW. of Fortescue Roadhouse , 21°23 ’15” S, 116°03 ’05” E, 19 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T88698 ) ; 1 male, same data except 20 March 2004 ( WAM T88700 ) ; 1 male, same data except 22 March 2004 ( WAM T88705 ) ; 1 male, same data ( WAM T88708 ) ; 1 male, same data ( WAM T88713 ) ; 1 male, 17.9 km SW. of Fortescue Roadhouse , 21°25 ’00” S, 116°02 ’24” E, 19 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T88703 ) ; 1 male, same data ( WAM T88706 ) ; 1 male, same data except 24 March 2004 ( WAM T88707 ) ; 1 male, 18.4 km NE. of Fortescue Roadhouse , 21°09 ’14” S, 116°13 ’59” E, 21 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T88701 ) ; 1 male, same data ( WAM T88710 ) ; 1 male, same data ( WAM T88712 ) ; 1 male, same data except 22 March 2004 ( WAM T88714 ) ; 1 male, 19.3 km SW. of Fortescue Roadhouse , 21°26 ’07” S, 116°01 ’55” E, 24 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T88699 ) ; 1 male, 42 km N. of Fortescue Roadhouse , 20°54 ’53” S, 116°14 ’03” E, 20 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T88697 ) ; 1 male, same data ( WAM T88709 ) ; 1 male, same data ( WAM T88711 ) ; 2 males, Jimblebar minesite, 35 km E. of Newman , 23°22 ’09” S, 120°11 ’44” E, 9 February 2009, pitfall trap, P. Bolton, C. Weston ( WAM T95399 DNA ) ; 1 male, same data except 23°22 ’29” S, 120°12 ’37” E, 12 February 2009, active search ( WAM T95402 DNA ) ; 1 male, same data except 23°22 ’45” S, 120°06 ’59” E, 9 February 2009, pitfall trap ( WAM T95400 DNA ) ; 1 male, same data except 23°23 ’27” S, 120°05 ’43” E, 6 February 2009, active search ( WAM T95401 DNA ) ; 1 male, Jinayri, ca. 65 km NW. of Newman , 22°51 ’10” S, 119°16 ’34” E, 31 March 2009, pitfall trap, J. Gollan, et al. ( WAM T97013 DNA ) ; 1 male, same data except 22°54 ’37” S, 119°12 ’30” E, 1 April 2009 ( WAM T97033 DNA ) ; 7 males, 3.5 km S. of Karratha, Pilbara Biological Survey, site DRC01 , 20°46 ’4.7” S, 116°50 ’31.3” E, 1 July 2003 - 2 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109322 ) ; 1 male, 6 km S. of Karratha, Pilbara Biological Survey site DRC03 , 20°47 ’40.1” S, 116°51 ’24.1” E, 3 July 2003 - 3 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109354 ) ; 1 male, 5 km N. of Lake Poongkaliyarra, Pilbara Biological Survey site DRC07 , 20°56 ’04.4” S, 117°06 ’54.3” E, 4 July 2003 - 3 October 2004, wet pitfall, DEC Pilbara survey ( WAM T109323 ) ; 1 male, 9 km NW. of Lake Poongkaliyarra, Pilbara Biological Survey site DRC05 , 20°56 ’23.8” S, 117°02 ’05.3” E, 3 July 2003, 3 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109330 ) ; 1 male, ca. 21 km S. of Laverton, site 1 , 28°47 ’34” S, 122°25 ’53” E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110258 ) ; 1 male, same data ( WAM T 110263 ) ; 1 male, same data ( WAM T 110269 ) ; 1 male, same data ( WAM T 110272 ) ; 1 male, ca. 21 km S. of Laverton, site 1 , 31°20 ’52” S, 119°23 ’26” E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110264 ) ; 1 female, ca. 21 km S. of Laverton, site 11 , 28°49 ’35” S, 122°23 ’20” E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110273 ) ; 1 male, ca. 21 km S. of Laverton, site 13 , 28°49 ’11” S, 122°23 ’14” E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110276 ) ; 1 male, ca. 21 km S. of Laverton, site 2 , 28°47 ’55” S, 122°26 ’01” E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110265 ) ; 1 male, same data except 8 January 2011 ( WAM T 110270 ) ; 1 male, ca. 21 km S. of Laverton, site 5 , 28°49 ’07” S, 122°26 ’02” E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110271 ) ; 1 male, ca. 21 km S. of Laverton, site 7 , 28°49 ’25” S, 122°26 ’31” E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110262 ) ; 1 male, same data except 6 January 2011 ( WAM T 110267 ) ; 1 male, ca. 21 km S. of Laverton, site 9 , 28°49 ’55” S, 122°25 ’34” E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110257 ) ; 1 male, Lorna Glen Biological Survey , 26°06 ’46” S, 121°30 ’14” E, 17 March 2004, dry pit, Cowan, M. ( WAM T88663 ) ; 1 male, 3.8 km E. of Lorna Glen Station , 26°13 ’35” S, 121°31 ’08” E, 16 March 2004, dry pit, M. Cowan ( WAM T88653 ) ; 1 male, same data except 17 March 2004 ( WAM T88658 ) ; 1 male, 11.9 km SW. of Lorna Glen Station , 26°18 ’11” S, 121°28 ’31” E, 17 March 2004, dry pit, M. Cowan ( WAM T88659 ) ; 1 male, 25.2 km N. of Lorna Glen Station , 26°00 ’05” S, 121°33 ’48” E, 16 March 2004, dry pit, M. Cowan ( WAM T88656 ) ; 1 male, Mallina Station, SW. of homestead, Pilbara Biological Survey site DRE12 , 20°54 ’36” S, 117°58 ’58” E, 14 November 2003 - 12 May 2004, wet pitfall, CALM Pilbara Survey ( WAM T109343 ) ; 5 males, 10 km S. of Mallina Homestead, Pilbara Biological Survey site DRE13 , 20°58 ’10.4” S, 118°02 ’54.0” E, 10 July 2003 - 3 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109327 ) ; 1 male, 1.2 km SSE. of Millstream, Pilbara Biological Survey site PW08 , 21°36 ’15” S, 117°04 ’39” E, 14 July 2003 - 12 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109336 ) ; 1 juvenile, Mesa A, 43.9 km W. of Pannawonica, site MAT21 , 21°41 ’18” S, 115°54 ’06” E, 5-11 August 2005, dry pitfall traps, R. Teale ( WAM T72402 ) ; 1 male, Mesa J, 16.6 km SW. of Pannawonica, site MEJ1 , 21°45 ’34” S, 116°13 ’39” E, 14-20 March 2005, dry pitfall traps, D. Kamien ( WAM T72400 ) ; 2 males, same data ( WAM T72401 ) ; 3 males, Mesa J, 17 km SW. of Pannawonica, site MEJ2 , 21°45 ’37” S, 116°13 ’16” E, 14-20 March 2005, dry pitfall traps, D. Kamien ( WAM T72399 ) ; 3 males, 2.5 km N. of Mile Camp, Pilbara Biological Survey site RHNE02 , 22°45 ’57.8” S, 119°37 ’53.1” E, 10 August 2003 - 19 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109321 ) ; 1 male, Millstream-Chichester National Park, Pilbara Survey site PW10 , 21°32 ’48” S, 117°03 ’33” E, 23 November 2003 - 8 May 2004, wet pitfall trap, CALM Pilbara Survey ( WAM T109320 ) ; 1 male, 24 km ENE. of Moorimoordinina Native Well, Pilbara Biological Survey site RHNE08 , 22°27 ’07” S, 119°58 ’34” E, 9 August 2003 - 20 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109335 ) ; 4 males, 10.5 km NW. of Mt Berry, Pilbara Biological Survey site WYE12 , 22°24 ’38.6” S, 116°23 ’30.8” E, 10 September 2003 - 9 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109341 ) ; 2 males, 19.7 km WNW of Mt Berry, Pilbara Biological Survey site WYE09 , 22°26 ’15.3” S, 116°16 ’27.3” E, 8 September 2003 - 10 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109344 ) ; 2 males, 8 km NNE. of Mt Edith, Pilbara Biological Survey, site WYE05 , 22°34 ’16.9” S, 116°08 ’53.8” E, 11 September 2003 - 7 October 2004, Ethylene Glycol Pit, CALM Pilbara Survey ( WAM T109345 ) ; 1 male, 14.5 km SE. of Mt Florance Homestead, Pilbara Biological Survey site PE06 , 21°51 ’22.2” S, 117°58 ’47.3” E, 3 September 2003 - 11 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109362 ) ; 6 males, 24 km WSW. of Mt Marsh, Pilbara Biological Survey site RHNW02 , 22°32 ’09” S, 118°59 ’51” E, 12 August 2003 - 19 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109329 ) ; 1 male, 26 km WSW. of Mt Marsh, Pilbara Biological Survey site RHNW01 , 22°32 ’06” S, 118°58 ’39” E, 12 August 2003 - 19 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109350 ) ; 1 male, 7 km SSE. of Mt Minnie, Pilbara Biological Survey site WYW03 , 22°10 ’10.8” S, 115°33 ’39” E, 27 September 2003 - 30 September 2004, wet pitfall, CALM Pilbara Survey ( WAM T109325 ) ; 1 male, Mount Webber , 20°31 ’21” S, 119°18 ’41” E, 31 March 2010 - 7 May 2010, wet pitfall trap, gorge, P. Bolton ( WAM T 103131 ) ; 1 male, Mount Webber , 21°31 ’57” S, 119°17 ’44” E, 30 March 2010 - 7 May 2010, wet pitfall trap, east facing ridge, P. Bolton ( WAM T 103128 ) ; 1 male, Mount Webber , 21°32 ’00” S, 119°17 ’33” E, 29 March 2010 - 7 May 2010, wet pitfall trap, gorge, P. Bolton ( WAM T 103129 ) ; 2 males, Mt Stuart Station, NE. of homestead, Pilbara Survey site WYE13 , 22°25 ’30” S, 116°25 ’57” E, 1 May 2004 - 10 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109349 ) ; 1 male, North Star, Pilbara , 21°16 ’23.81” S, 119°3 ’24.48” E, 11-18 February 2011, leaf litter, Thompson ( WAM T 112591 ) ; 1 male, 12.5 km N. of Nullagine, Pilbara Biological Survey site NW03 , 21°46 ’13.1” S, 120°05 ’30.7” E, 5 August 2003 - 18 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109331 ) ; 1 male, 30 km NE. of Nullagine , 21°48 ’16.08” S, 120°23 ’18.00” E, 29 April 2010, C. Jackson ( WAM T 102065 ) ; 2 males, 56.3 km N. of Nullagine, Pilbara Biological Survey site NW04 , 21°40 ’42.7” S, 120°05 ’18.2” E, 19 May 2004 - 18 May 2005, ethylene glycol pits, CALM Pilbara Survey ( WAM T104454 ) ; 1 male, Orebody 35, ca. 8 km W. of Newman, Site10-P6 , 23°23 ’59.65” S, 119°39 ’07.47” E, 12 March 2010 - 18 May 2010, pitfall trap, open floodplain, J. Gollan ( WAM T104794 ) ; 1 male, Orebody 35, ca. 8 km W. of Newman, Site3-HC1 , 23°24 ’17.00” S, 119°36 ’18.90” E, 15 March 2010, hand collected, drainage gully, south facing ridge, deeply vegetated, J. Gollan, M. Bulbert, M. Semeniuk, J. Bevan ( WAM T104784 ) ; 1 male, Orebody 35, ca. 8 km W. of Newman, Site 6-P8 , 23°24 ’07.00” S, 119°34 ’27.88” E, 12 March-18 May 2010, pitfall trap, slope with spinifex, J. Gollan ( WAM T104810 ) ; 1 male, Orebody 35, ca. 8 km W. of Newman, Site9-P10 , 23°23 ’22.32” S, 119°38 ’36.06” E, 12 March-18 May 2010, pitfall trap, open floodplain, J. Gollan ( WAM T104807 ) ; 1 male, 2.3 km ESE. of Python Pool, Pilbara Biological Survey site PW01 , 21°20 ’22.8” S, 117°15 ’36.3” E, 16 July 2003 - 11 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T109356 ) ; 2 males, 5 km WSW. of Python Pool, Pilbara Biological Survey site PW03 , 21°20 ’28.7” S, 117°11 ’18.9” E, 8 May 2004 - 12 May 2005, wet pitfall, CALM Pilbara Survey ( WAM T109339 ) ; 5 males, 1 juvenile, 11.5 km SW. of Rhodes Ridge, Pilbara Biological Survey site RHNC07 , 23°08 ’45.7” S, 119°15 ’56.9” E, 25 May 2004 - 11 May 2005, wet pitfall, CALM Pilbara Survey ( WAM T109324 ) ; 1 male, Roy Hill , 22°25 ’35.77” S, 119°56 ’32.69” E, 11-17 March 2010, wet pitfall trap, N. Thompson, L. Roque-Albelo ( WAM T 102618 ) ; 1 female, South Parmelia, 66 km NW. of Newman , 23°02 ’08” S, 119°11 ’17” E, 12 April 2011, dug from burrow, R. Teale, M. Greenham ( WAM T113574 ) ; 1 female, same data except 23°02 ’04” S, 119°11 ’32” E ( WAM T113576 ) ; 1 female, South Parmelia, 52 km NW. of Newman , 23°05 ’11” S, 119°19 ’07” E, 16 April 2011, dug from burrow, R. Teale, M. Greenham ( WAM T113582 ) ; 1 female, Southern Flank, 71 km NW. of Newman 22°59 ’40” S, 119°09 ’55” E, 17 April 2011, dug from burrow, R. Teale, M. Greenham ( WAM T113612 ) ; 1 male, Tom Price , 22°42'S, 117°47'E, 3 January 1994, M. Hyams ( WAM T30075, formerly 94/486 ) ; 1 female, Tom Price Airport, 14 km NW. of Tom Price , 22°34 ’48” S, 117°43 ’51” E, 1-30 November 2007, D. Kamien, J. Adcroft ( WAM T 102911 ) ; 2 males, Warramboo, 50.5 km W. of Pannawonica, site WA3 , 21°40 ’00” S, 115 °50 ’06” E, 14 -20 March 2005, dry pitfall traps, D. Kamien ( WAM T72404 ) ; 1 male, Warramboo, 52.1 km W. of Pannawonica, site WA2 , 21°39 ’49” S, 115°49 ’10” E, 14-20 March 2005, dry pitfall traps, D. Kamien ( WAM T72403 ) ; 1 male, 14 km SE. of Wheelarra Hill, Pilbara Biological Survey site BDRS10 , 23°28 ’16” S, 120°12 ’54” E, 6 September 2005 - 10 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T109352 ) ; 1 male, 34 km E. of Wheelarra Hill, Pilbara Biological Survey site BDRS03 , 23°21 ’31” S, 120°27 ’33” E, 4 September 2005 - 11 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T109347 ) ; 1 male, 11 km SSE. of Whim Creek Hotel, Pilbara Biological Survey site DRE10 , 20°55 ’11.4” S, 117°51 ’40.6” E, 11 July 2003 - 3 October 2004, ethylene glycol pits CALM Pilbara Survey ( WAM T109333 ) ; 2 males, 1 female, 2 juveniles, 5 km E. of Whim Creek Hotel, Pilbara Biological Survey site DRE08 , 20°50 ’54.6” S, 117°51 ’16.2” E, 11 July 2003 - 3 October 2004, ethylene glycol pits CALM Pilbara Survey ( WAM T109319 ) ; 1 male, ca. 7 km NE. of Wodgina mine site , 21°09 ’10.04” S, 118°43 ’10.03” E, 1 April-4 May 2010, wet pitfall trap, minor drainage line, P. Bolton ( WAM T107002 ) ; 1 male, ca. 70 km SE. of Wodgina mine , 21°35 ’22” S, 119°11 ’10” E, 31 March-6 April 2010, wet pitfall trap, gully, P. Bolton ( WAM T105227 ) ; 1 male, same data ( WAM T105228 ) ; 1 male, 8.5 km WSW. of Yanyare River Mouth, Pilbara Biological Survey site DRW02 , 20°50 ’40.6” S, 116°22 ’02.9” E, 23 September 2003 - 2 October 2004, wet pitfall CALM Pilbara Survey ( WAM T109326 ) .

Etymology. The specific epithet refers to the yellow-brown colouration of parts of the carapace (mellosa, Latin, of honey, honey-coloured) ( Brown 1956).

Diagnosis. Males of A. mellosa differ from all other named species of the genus by the colouration of the carapace which is overall light yellowish-brown with the cephalic region honey-coloured to very dark reddish-brown (Fig. 10). Females also have a somewhat darker cephalic region but the colour difference between the cephalic and thoracic regions is much less pronounced (Fig. 21). Tibia I of males has a large spur and megaspine located medially, and metatarsus I is incrassate with the ‘heel’ of the thickening about half-way; this conformation most closely resembles A. pallida from eastern Queensland from which it differs by the less pronounced metatarsal process and the smaller tibial spur and megaspine. Female genitalia resemble those of A. longitheca Raven, 1984 from central Queensland in possessing long lateral tubes (Figs 23, 24), but they differ in the shape of the median spermathecae which are coiled in A. longitheca and S-shaped in A. mellosa . See Remarks for further clarification.

Description. Adult male (based on holotype WAM T 94870): medium-sized nemesiid spider (total length 18.4).

Colour, carapace (Fig. 10) light yellow-brown in thoracic region turning darker towards anterior parts, cephalic region dark reddish-brown, eye region black (Fig. 13); sternum light yellow-brown, sigilla and labium largely brown (Fig. 12); abdomen dorsally dark brownish-grey, mottled with small light-brown spots (Fig. 11), ventrally light yellowish; chelicerae very dark reddish-brown; pedipalp segments reddish-brown, tibia somewhat lighter (Figs 17, 18); legs light yellowish-brown (Figs 15, 16).

Carapace: 7.0 long, 5.6 wide; eye group (Fig. 13) 1.4 wide, 0.7 long; fovea procurved (Fig. 10).

Sternum and labium: 4.4 long, 2.8 wide; 2 pairs of elongated sigilla in posterior half (Fig. 12); labium wider than long, slightly indented anteriorly (Fig. 14).

Maxillae: with> 50 pin-like cuspules, extending to heel (Fig. 14).

Chelicerae: without rastellum (Fig. 12), promargin with 9 large teeth, retromargin with 4 smaller teeth proximally.

Abdomen: 7.4 long, 5.0 wide (Fig. 11). Four spinnerets, terminal segment of posterior lateral spinnerets digitiform.

Pedipalp: length of femur 4.0, patella 2.2, tibia 3.1, tarsus 3.1. Femur with 1 disto-prolateral spine; patella with 3 prolateral spines; tibia with 2 baso-ventral spines and 7-8 prolateral spines in apical half, tibia with slight disto-ventral depression without setae (accommodates bulb and embolus) (Figs 17, 18); tarsus terminally blunt; bulb globular, embolus ca. 0.8 long, slightly curved and tapering (Figs 17-20).

Legs: femur I with row of 7 dorsal spines in basal half and 2 disto-prolateral spines, patella with 2 prolateral spines, metatarsi and tarsi without spines. Tibia I with large spur and megaspine, metatarsi incrassate (Fig. 16). Tarsi and apical half of metatarsi of legs I and II ventrally with entire dense scopulae, tarsi of legs III and IV with divided weak scopulae. Tarsi, metatarsi and tibiae with numerous dorsal trichobothria of variable length. Paired tarsal claws of legs with two rows of 7-10 teeth each; third tarsal claw reduced and very small.

Leg measurements: length of legs I>IV>II>III. leg I: femur 6.0, patella 3.5, tibia 4.6, metatarsus 4.2, tarsus 2.5, total = 20.8. Dorsal proximal width of patella I = 1.4, tibial index = 0.12, leg I formula = 2.97. Leg II: femur 6.0, patella 3.4, tibia 4.5, metatarsus 4.9, tarsus 3.1, total = 21.9. Dorsal proximal width of patella II = 1.2, tibial index = 0.15; leg II formula = 3.13. Leg III: femur 5.2, patella 3.0, tibia 3.2, metatarsus 6.6, tarsus 2.9, total = 20.9. Dorsal proximal width of patella III = 1.3, tibial index = 0.19; leg III formula = 2.99. Leg IV: femur 7.0, patella 3.5, tibia 5.9, metatarsus 6.3, tarsus 2.9, total = 25.6. Dorsal proximal width of patella IV = 1.3, tibial index = 0.14; leg III formula = 3.66.

Variation: total length 5.1-6.2, carapace 6.1-7.9 long, 16.9-20 wide (n = 12).

Adult female (based on WAM T 96150): medium-sized nemesiid spider (total length 19.5).

Colour, light yellowish-brown with indistinctly darker cephalic region (Fig. 21), eye region brown; sternum, sigilla and labium uniformly light yellowish-brown; abdomen dorsally dark greyish-brown with purple tinge and mottled with small lighter small spots, ventrally light yellowish-brown; chelicerae orange-brown; pedipalps and legs uniformly light yellow-brown.

Carapace: 7.5 long, 6.4 wide; eye group 1.4 wide, 0.8 long; fovea slightly procurved.

Sternum and labium: 5.0 long, 3.2 wide (Fig. 22); 2 pairs of indistinct elongated sigilla in posterior half, 1 pair of round sigilla at base of leg II; labium wider than long, slightly indented anteriorly.

Maxillae: with> 100 pin-like cuspules, extending to heel and covering in low density almost entire retrolateral half of maxillae.

Chelicerae: without rastellum, promargin with 9 large teeth, retromargin with 4 smaller teeth proximally.

Abdomen: 8.9 long, 5.9 wide. Four spinnerets, terminal segment of posterior lateral spinnerets digitiform.

Genitalia: median spermathecal stalks S-shaped, spermathecae slightly wider than spermathecal stalks; separate straight and elongated tube accompanies spermatheca (Figs 23, 24).

Pedipalp: length of femur 3.6, patella 2.3, tibia 2.5, tarsus 2.7. Femur without spine, patella with 2 prolateral spines; tibia with 11-13 irregularly placed spines in mainly apical half (not dorsal); tarsus with scopulae ventrally and 2 baso-ventral spines and curved terminal claw.

Legs: femur I with 1 dorsal spines proximally and 1 disto-prolateral spine, patella with 2 (left leg 1) prolateral spine, tibia with a row of three ventral spines (5 on right leg of which 2 are paired) and 2 disto-ventral spines, metatarsi 1 basal and 1 apical pair of ventral spines (left leg) (3 additional medio-ventral spines on left leg). Tarsi and apical half of metatarsi of leg I and II ventrally with entire dense scopulae, tarsi of legs III and IV with divided very weak scopulae. Tarsi, metatarsi and tibiae with numerous dorsal trichobothria of variable length. Paired tarsal claws of legs with two rows of 10-13 (legs I-II) or 5-7 (legs III-IV) teeth each; third tarsal claw reduced and very small.

Leg measurements: length of legs IV>I>II>III. leg I: femur 6.0, patella 3.1, tibia 4.0, metatarsus 3.5, tarsus 2.3, total = 18.9. Dorsal proximal width of patella I = 1.0, tibial index = 0.14, leg I formula = 2.12. Leg II: femur 5.0, patella 3.0, tibia 3.3, metatarsus 3.1, tarsus 2.1, total = 16.5. Dorsal proximal width of patella II = 0.9, tibial index = 0.14; leg II formula = 1.85. Leg III: femur 4.0, patella 2.5, tibia 2.0, metatarsus 2.9, tarsus 1.7, total = 13.1. Dorsal proximal width of patella III = 1.1, tibial index = 0.24; leg III formula = 1.47. Leg IV: femur 5.5, patella 3.2, tibia 4.0, metatarsus 4.4, tarsus 2.2, total = 19.3. Dorsal proximal width of patella IV = 1.1, tibial index = 0.15; leg III formula = 2.17.

Variation: total length 19.3-25.0, carapace 5.4-6.6 long, 6.6-8.7 wide (n = 8).

Distribution. Aname mellosa has been found throughout the central Pilbara, from Jimblebar minesite near the township of Newman in the east to Mount Minnie in the west, and south into the Midwest region of Western Australia (Fig. 5).

Remarks. Care must be taken when identifying A. mellosa as a number of undescribed species from northern Western Australia have a very similar somatic appearance, but have a different genital morphology. The embolus of male A. mellosa is long and slightly curved, but it is often much shorter and/or much more curved in the undescribed species. An unambiguous identification of females or juveniles in this group is currently not possible, however, we have listed females and juveniles when they have been collected with or in close proximity to males. As stated in the Diagnosis, females of A. mellosa have an unusual genital morphology with long, straight lateral tubes in addition to the median pair of spermathecae. This morphology is similar to that of A. longitheca ( Raven 1985a), but both species differ in the shape of the median spermathecae.

All records of mature males were collected between January and early June suggesting this species to be reproductively active towards the end of the wet season. Many mature males of A. mellosa were removed from their burrows ( R. Teale, pers. comm.) where it is likely that they had matured and were waiting for suitable conditions to emerge and search for adult females. The silk-lined open burrow of A. mellosa is dug at an angle into the substrate and lacks a lid (Figs 2-4) ( R. Teale, pers. comm.).