Philonthus Stephens 1829

Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey, 2018, Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae), Zootaxa 4449 (1), pp. 1-95: 58-81

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https://doi.org/10.11646/zootaxa.4449.1.1

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Philonthus Stephens 1829
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Philonthus Stephens 1829 

Philonthus Stephens 1829a: 23  , attributed to Leach. Type species: Staphylinus splendens Fabricius, 1792  , designated by Curtis (1836: pl. 610). Notes: also Stephens (1829b: 279, 1832: 226), Curtis (1829: 25), Dillwyn (1829: 68); Bouchard et al. (2011) use Curtis (1829) as author.

= Spatulonthus Tottenham 1955: 154  (as subgenus of Philonthus  ). Type species: Philonthus longicornis Stephens, 1832  , by original designation. Note: as valid genus, Coiffait & Sáiz (1968: 360); as synonym (not subgenus) of Philonthus, Smetana (1995: 51)  .

= Endeius Coiffait & Sáiz 1968: 355  . Type species: Endeius loensis Coiffait & Sáiz, 1968  , by original designation. Distribution: Chile, Argentina, Juan Fernandez Is., Peru, Galapagos Is. Notes: revised (Chani-Posse de Maus 2008); as synonym of Philonthus  , one species moved to Philonthina  incertae sedis (Chani-Posse 2013: 404).

Notes: The global genus Philonthus  , with 1326 species currently known, is the fourth largest genus within Staphylinidae  and the largest within Staphylininae  (Newton unpublished database). The Neotropical region accounts for 189 species and most of them (160 spp.) are endemic species. The large number of Philonthus  species to be studied has been a challenging factor to achieve a modern concept of the genus, which has been

repeatedly shown to be non-monophyletic ( Chani-Posse et al. 2018). The current concept of the genus is that of Smetana (1995: 51), who did not recognize subgenera.

Philonthus abdominalis Erichson 1840 

Philonthus abdominalis Erichson 1847: 89  . Type locality: [ Peru, east side]  . Type depository: ZMHB 1 ST ♀ [ MCPAbout MCP seen]. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Philonthus accedens Sharp 1885 

Philonthus accedens Sharp 1885: 403  . Type locality: Guatemala: Capetillo. Type depository: BMNH 2 ST ♀. Distribution: Mexico, Guatemala. Note: distribution ( Navarrete-Heredia et al. 2002). Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus acromyrmecis Scheerpeltz 1976 

Philonthus acromyrmecis Scheerpeltz 1976: 135  . Type depository: NMWAbout NMW HT ♂, 3 PT; FMNH 1View Materials PT; DZUP 1View Materials PT. Type locality: Kolonie Nova Teutonia , Brazil  . Distribution: Brazil (Santa Catarina). Notes: with Acromyrmex crassispinus  (originally as hispidus atratus; Navarrete-Heredia 2001). 

Philonthus actinus Bernhauer 1917 

Philonthus actinus Bernhauer 1917c: 103  , attributed to Fauvel in litteris. Type locality: Yuracaris , Bolivien  . Type depository: FMNH 2View Materials ST; ZMUH 1View Materials +ST?. Distribution : Bolivia  .

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus actocharis Sharp 1887 

Philonthus actocharis Sharp 1887: 788  . Type locality: Tehuantepec , Mexico  . Type depository: BMNH 7 ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus aenescens Solsky 1870 

Philonthus aenescens Solsky 1870: 262  . Type locality: " Mexique "  . Type depository: ZINAbout ZIN? #? ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus aeruginosus Nordmann 1837 

Philonthus aeruginosus Nordmann 1837: 95  . Type locality: Brasilia meridionali. Type depository: ZMHB LT ♂, 4 PLT. Distribution: Colombia, Guyana, Brazil, Paraguay, Uruguay, Argentina, Chile. Notes: lectotype designation, redescription, distribution including Chile ( Chani-Posse 2010: 12); distribution within Brazil (Asenjo et al. 2013).

Philonthus aliiseriatus Bruch 1915  , nomen nudum

Philonthus aliiseriatus Bruch 1915: 501  , attributed to Bernhauer in litteris; nomen nudum. Locality: Jujuy. Distribution: Argentina. Note: not in Fernández et al. (2007) or Bachmann et al. (2017).

Philonthus alius Bernhauer & Schubert 1914 

Philonthus alius Bernhauer & Schubert 1914: 328  (new name for Philonthus crassulus Sharp 1885: 396  ). Distribution: Mexico, Guatemala. Notes: Márquez-Luna & Navarrete-Heredia (1995), Blackwelder (1944) and Bernhauer & Schubert (1914) cite only Guatemala, in error; distribution, also within Mexico (Navarrete- Heredia et al. 2002, Márquez 2006, Navarrete-Heredia 2012); ex external refuse piles Atta mexicana  (Márquez-Luna & Navarrete-Heredia 1995, Navarrete-Heredia 2001, 2012).

= Philonthus crassulus Sharp 1885: 396  (preoccupied, junior primary homonym of Philonthus crassulus Horn 1884: 213  ). Type depository: BMNH 3+ ST. Type locality: Mexico: Las Vigas, Guanajuato. Distribution: Mexico. Note: with Atta mexicana ( Kistner 1982)  .

Philonthus alticola Sharp 1885 

Philonthus alticola Sharp 1885: 400  . Type locality: Guatemala: Totonicapam 8500–10500 ft  . Type depository: BMNH HT ♀. Distribution : Guatemala  .

Philonthus alumnus Erichson 1840 

Philonthus alumnus Erichson 1840: 490  . Type depository: ins. Puerto Rico. Type depository: ZMHB LT ♂, 3 PLT. Distribution: Cuba, Puerto Rico, Haiti, Cayman Is., Mona I. Notes: lectotype designation, redescription and distribution ( Smetana 1994: 342); restricted concept, West Indies only (distribution cited) ( Smetana 1995); distribution within Cuba ( Peck 2005), within Cayman Is. (Frank et al. 2011).

Philonthus alutaceus Horn 1884 

Philonthus alutaceus Horn 1884: 183  . Type locality: California [no specific locality]. Type depository: MCZAbout MCZ (Horn) LT ♀, 3 PLT; (LeConte) 7 PLT. Distribution: southern USA, Mexico. Note: lectotype designation, redescription, distribution in Mexico ( Smetana 1995: 106); Note: distribution, also within Mexico (Navarrete- Heredia et al. 2002).

Philonthus ambiguus Erichson 1840 

Philonthus ambiguus Erichson 1840: 441  . Type locality: Columbia. Type depository: ZMHB 4 ST: 2 ♂, 2 ♀ [ MCPAbout MCP seen]. Distribution : Colombia  .

Philonthus apheles Solsky 1868 

Philonthus apheles Solsky 1868: 132  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]  . Type depository: ZINAbout ZIN? ( Solsky ) #? ST. Distribution: Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus apicetestaceus Scheerpeltz 1960 

Philonthus apicetestaceus Scheerpeltz 1960: 106  . Type locality: Bolivien, Yungas de Arepucho , Chacisacha, etwa 1500m  . Type depository: NMWAbout NMW ( Scheerpeltz ) HT ♀. Distribution: Bolivia  .

Philonthus argus Herman 2001 

Philonthus argus Herman 2001a: 46  (new name for Philonthus tucumanensis Bernhauer 1934a: 118  ). Notes: redescription ( Chani-Posse 2010: 13); distribution ( Chani-Posse 2010); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Philonthus tucumanensis Bernhauer 1934a: 118  (preoccupied, junior primary homonym of Philonthus tucumanensis Bernhauer 1927: 245  ). Type locality: Tucuman: Parque Aconquija. Type depository: FMNH LT ♂. Distribution: Argentina. Notes: not in Blackwelder (1944); lectotype designation, one other labeled as type considered not type ( Chani-Posse 2010: 14).

Philonthus arizonensis Horn 1884 

Philonthus arizonensis Horn 1884: 203  . Type locality: Arizona (implied, no locality cited) [no specific locality]. Type depository: MCZAbout MCZ (Horn) LT ♂, 6 PLT; (LeConte) 4 PLT. Distribution: USA, Mexico. Notes: lectotype designation, redescription, distribution in Mexico ( Smetana 1995: 374); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus atramentarius Erichson 1840 

Philonthus atramentarius Erichson 1840: 440  . Type locality: Columbia. Type depository: ZMHB 1 ST ♀ [ MCPAbout MCP seen]. Distribution : Colombia  .

Philonthus audanti Blackwelder 1943 

Philonthus audanti Blackwelder 1943: 406  . Type locality: Haiti: 5 miles south of St. Marc. Type depository: USNMAbout USNM HT ♂. Distribution : Haiti  .

Philonthus auripennis Bernhauer 1916 

Philonthus auripennis Bernhauer 1916a: 37  , attributed to Fauvel in litteris. Type locality: Bolivien. Type depository: FMNHAbout FMNH HT ♀  . Distribution: Bolivia. Note: as valid species in Herman (2001d).

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus avicola Bernhauer 1942 

Philonthus avicola Bernhauer 1942: 25  . Type locality: Reventazon , Ebene Limon (Hamburgfarm) [ Costa Rica]  . Type depository: FMNH 2View Materials ST. Distribution : Costa Rica  .

Comment. Not Philonthus  , to be placed in a further study.

Philonthus belonuchiformis Bruch 1915  , nomen nudum

Philonthus belonuchiformis Bruch 1915: 501  , attributed to Bernhauer in litteris; nomen nudum. Locality: Buenos Aires. Distribution: Argentina. Note: not in Fernández et al. (2007) or Bachmann et al. (2017); not Philonthus belonuchiformis Bernhauer (1915: 145)  from Ethiopia.

Philonthus bicoloristylus Chani-Posse 2004 

Philonthus bicoloristylus Chani-Posse 2004: 219  . Type locality: Argentina, Tucumán, El Cadillal. Type depository: IMLAAbout IMLA HT ♂, 16 PT; FMNHAbout FMNH 5 PT, MACNAbout MACN 1 PT. Distribution: Colombia, Argentina. Notes: redescription, distribution including Colombia ( Chani-Posse 2010: 14); distribution ( Aballay et al. 2014).

Philonthus bolivianus Bernhauer 1916 

Philonthus bolivianus Bernhauer 1916a: 35  . Type locality: Bolivien: Coroico, 1800m. Type depository: FMNHAbout FMNH HT ♂. Distribution: Bolivia, Brazil (based on material in FMNHAbout FMNH identified by M. Bernhauer). Note: Brazil (Rio de Janeiro); not in Asenjo et al. (2013).

Philonthus bonariensis Bernhauer 1909 

Philonthus bonariensis Bernhauer 1909: 245  . Type locality: Buenos Aires, Argentina. Type depository: FMNHAbout FMNH LT ♂; MCSNAbout MCSN 4 PLT?. Distribution: Brazil, Paraguay, Uruguay, Argentina, Chile. Notes: distribution including Uruguay (Chani-Posse 2004: 223); lectotype designation (Chani-Posse de Maus 2009: 38); redescription, distribution including Brazil, Chile and Paraguay ( Chani-Posse 2010: 15); distribution within Brazil (Asenjo et al. 2013); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Philonthus catamarcanus Bernhauer 1916a: 36  . Type locality: Prov. Catamarca, Argentinien [LT]. Type depository: FMNH LT ♀, 1 PLT. Distribution: Argentina. Note: as synonym of P. bonariensis  , lectotype designation (Chani-Posse de Maus 2009: 38); also type specimens labeled as P. catamarcensis ( Chani-Posse 2010: 19)  .

= Philonthus weiseri Bernhauer 1921c: 169  . Type locality: Cueva Iturba, 3700m, Jujuy, Argentina. Type depository: FMNHAbout FMNH LT ♂, 2 PLT. Distribution: Argentina. Note: as synonym of P. bonariensis  , lectotype designation (Chani-Posse de Maus 2009: 38).

Philonthus brachypterus Solsky 1872 

Philonthus brachypterus Solsky 1872: 297  . Type locality: Cayenne [Amérique du Sud]  . Type depository: ZINAbout ZIN? (Solsky) HT. Distribution: French Guiana ( Cayenne ). 

Philonthus brasilianus Bernhauer 1916 

Philonthus brasilianus Bernhauer 1916a: 40  . Type locality: Brasilien: S. Paulo (Ypiranga)  . Type depository: FMNH 2View Materials ST. Distribution : Brazil (São Paulo)  . Note: distribution within Brazil (Asenjo et al. 2013).

Philonthus bruchianus Chani-Posse 2010 

Philonthus bruchianus Chani-Posse 2010: 20  , attributed to Bernhauer MS. Type locality: Tucumán, Siambon. Type depository: FMNHAbout FMNH HT ♂, 1 PT ♀. Distribution : Argentina  . Note: distribution in carcasses in Argentina ( Aballay et al. 2014).

Philonthus caliensis Bernhauer 1916 

Philonthus caliensis Bernhauer 1916a: 33  . Type locality: West-Columbien: Umgebung von Cali im Caucatale. Type depository: FMNH 1View Materials ST. Distribution : Colombia  .

Comment. Not Philonthus  , to be placed in a further study.

Philonthus callosipennis Scheerpeltz 1960 

Philonthus callosipennis Scheerpeltz 1960: 100  . Type locality: Cochabamba ( Bolivien), Yungas de Palmar , 2000m  . Type depository: NMWAbout NMW ( Scheerpeltz ) 3 ST. Distribution: Bolivia. Note: generic affiliation unclear. 

Philonthus caribaeus Bierig 1940 

Philonthus caribaeus Bierig 1940: 141  . Type locality: Provincia de la Habana y de Pinay del Río. Type depository: FMNH 2View Materials ST. Distribution : Cuba  . Note: distribution within Cuba ( Peck 2005).

Philonthus carinulatus Sharp 1885 

Philonthus carinulatus Sharp 1885: 402  . Type locality: Mexico  : Jalapa, Oaxaca, Las Vigas. Type depository: BMNH 4 ST: 2 Oaxaca, 1 Jalapa, 1 Las Vigas ; other 2 specimens only from Mexico [ MCPAbout MCP seen]. Distribution   : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus catamarcensis Bruch 1915  , nomen nudum

Philonthus catamarcensis Bruch 1915: 501  , attributed to Bernhauer in litteris; nomen nudum. Locality: Catamarca. Depository: MACN (nine specimens). Distribution: Argentina. Note: see Bachmann et al. (2017: 44).

Philonthus cavifrons Sharp 1885 

Philonthus cavifrons Sharp 1885: 408  . Type locality: Panama, Bugaba 80 0–1500 ft  . Type depository: BMNH HT ♂. Distribution : Panama  .

Philonthus centralis Sharp 1885 

Philonthus centralis Sharp 1885: 422  . Type locality: Mexico: Jalapa; Guatemala, San Gerónimo; Panama, Volcan de Chiriqui. Type depository: BMNH 9 ST: from all type localities [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala, Panama. Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002); generic affiliation unclear.

Philonthus chiriquensis Sharp 1885 

Philonthus chiriquensis Sharp 1885: 404  . Type locality: Panama, Volcan de Chiriqui 4000–6000 ft  . Type depository: BMNH HT ♀. Distribution : Panama  .

Philonthus chlorocephalus Solsky 1872 

Philonthus chlorocephalus Solsky 1872: 301  . Type locality: Cayenne [Amérique du Sud]  . Type depository: ZINAbout ZIN? (Solsky) #? ST. Distribution: French Guiana ( Cayenne ). 

Philonthus columbianus Baudi di Selve 1848 

Philonthus (Philonthus) columbianus Baudi di Selve 1848: 126  . Type locality: Novae Valenciae in Columbia [ Venezuela, teste Herman 2001c: 1594]. Type depository: MRSNAbout MRSN / MCSNAbout MCSN? #?ST. Distribution: Colombia, Venezuela.

Philonthus convexicollis Lynch-Arribálzaga 1884 

Philonthus convexicollis Lynch-Arribálzaga 1884: 157  , attributed to Fauvel. Type locality: Chacabuco in western Buenos Aires prov. [orig. publication]. Type depository: MACNAbout MACN LT ♀. Distribution: Paraguay, Argentina, Chile. Notes: lectotype designation, redescription, distribution ( Chani-Posse 2010: 21).

Philonthus corallipennis Sharp 1876 

Philonthus corallipennis Sharp 1876a: 166  . Type locality: Pará, Obydos, Ega & Tapajos   . Type depository: BMNH 10 ST: from all type localities except Obydos [ MCPAbout MCP seen]. Distribution: Brazil ( Amazonas , Pará)  . Note: distribution within Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.

Philonthus cordilleranus Bernhauer 1916 

Philonthus cordilleranus Bernhauer 1916a: 34  . Type locality: West-Columbien: Ost-Cordilleren, La Pena bei Carmen, 3500m. Type depository: FMNHAbout FMNH HT ♀. Distribution : Colombia. Note: generic affiliation unclear. 

Philonthus cribrellus Fauvel 1891 

Philonthus cribrellus Fauvel 1891: 112  . Type locality: Caracas & Nouvelle-Grenade [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 2? ST. Distribution: Colombia, Venezuela. Note: generic affiliation unclear.

Philonthus cribriceps Bernhauer 1921 

Philonthus cribriceps Bernhauer 1921c: 105  . Type locality: Petropolis, Brazil  . Type depository: FMNHAbout FMNH HT. Distribution : Brazil (Rio de Janeiro)  . Note: distribution within Brazil (Asenjo et al. 2013).

Philonthus cribripennis Sharp 1885 

Philonthus cribripennis Sharp 1885: 403  . Type locality: Guatemala; Panama, Volcan de Chiriqui 2000–3000 ft. Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution: Guatemala, Panama;? Mexico.

Philonthus cribriventris Bernhauer 1912 

Philonthus cribriventris Bernhauer 1912a: 43  . Type locality: Prov. Tucuman [LT]. Type depository: FMNHAbout FMNH LT ♂, 1 PLT. Distribution: Argentina. Notes: lectotype designation (Chani-Posse de Maus 2009: 39); redescription, distribution ( Chani-Posse 2010: 25).

= Philonthus catamarcanus var. densior Bernhauer 1916a: 36  . Type locality: Provinz Tucuman, Argentinien. Type depository: FMNHAbout FMNH HT ♀  . Distribution: Argentina. Note: as synonym of P. cribriventris (Chani-Posse de Maus 2009: 39)  .

Philonthus crinitus Schubert 1909 

Philonthus crinitus Schubert 1909: 293  . Type locality: Jalapa region  , Veracruz, Mexico  . Type depository: ZMHB ( Schubert ) 1 ST [ MCPAbout MCP seen]. Distribution: Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus cruentatus ( Gmelin 1790)  [adventive in CASA]

Staphylinus cruentatus Gmelin 1790: 2035  . Type locality: Europe. Type depository: ?? #?ST, not seen by Smetana (1995: 233). Distribution: Europe, Russia (European, Siberia, Far East), Georgia, Armenia, Algeria, Morocco, Canary Is., Turkey, Lebanon, Iran, Kazakhstan, Kyrgyzstan, Afghanistan, Tajikistan, Uzbekistan, Mongolia, Japan, India; adventive in Canada, USA, Mexico. Notes: redescription, distribution ( Smetana 1995: 232); distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Navarrete-Heredia 2012).

Philonthus cyaneoaureus Bernhauer 1935 

Philonthus cyaneoaureus Bernhauer 1935: 90  . Type locality: Venezuela  . Type depository: BMNH #?ST; FMNH 1View Materials ST. Distribution : Venezuela  .

Comment. Not Philonthus  , to be placed in a further study.

Philonthus deplanatus Sharp 1885 

Philonthus deplanatus Sharp 1885: 423  . Type locality: Guatemala, San Gerónimo. Type depository: BMNH HT ♀. Distribution : Guatemala  .

Philonthus diamantinus Bernhauer 1917 

Philonthus diamantinus Bernhauer 1917c: 102  , attributed to Fauvel in litteris. Type locality: Yuracaris , Bolivien  . Type depository: FMNH 2View Materials ST; ZMUH 1View Materials +ST?. Distribution : Bolivia  .

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus discedens Schubert 1909 

Philonthus discedens Schubert 1909: 291  . Type locality: Jalapa region  , Veracruz, Mexico  . Type depository: ZMHB ( Schubert ) 1 ST ♀ [ MCPAbout MCP seen]. Distribution: Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus discoideus Gravenhorst 1802  [adventive in CASA]

Staphylinus discoideus Gravenhorst 1802: 38  . Type locality: [Brunswick, Germany]. Type depository: ZMHB LT ♂, 6 PLT. Distribution: Palearctic (Europe to north Africa, Iran, Korea, Japan), China, Oriental; adventive in Afrotropical region, Canada, USA, Mexico to Colombia, Venezuela, Brazil, Argentina, Chile, Cuba, Jamaica, Hispaniola, Puerto Rico, Lesser Antilles, Australia, New Zealand, Fiji, Tahiti, Galapagos Is., Madagascar, Mauritius, Réunion Is.. Notes: West Indies ( Blackwelder 1943, Frank 1983); lectotype designation, redescription and distribution ( Smetana 1995: 186); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution including Bolivia, Brazil, Colombia, Chile, Panamá, Paraguay ( Chani-Posse 2010); distribution, also within Cuba ( Peck 2005), within Galapagos Is. ( Newton & Peck 2006); Lesser Antilles ( Peck 2011a: Northern Leeward Is., Peck et al. 2014: Guadeloupe Archipelago; Peck 2016); redescription, distribution ( Hromádka 2012b); distribution within Brazil (Asenjo et al. 2013); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Philonthus emelinae Coiffait & Sáiz 1968: 354  . Type locality: Curicó : Romeral, Chile  . Type depository: Sáiz (in MNNCAbout MNNC) HT ♂. Note: as synonym of P. discoideus (Chani-Posse de Maus 2009: 39)  . 

Philonthus discretus Sharp 1876 

Philonthus discretus Sharp 1876b: 428  . Type locality: Ciudad de México  . Type depository: BMNH 2 ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus dispersus Fauvel 1891 

Philonthus dispersus Fauvel 1891: 111  . Type locality: Colonie Tovar, Lagonita & Nouvelle-Grenade [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 3 ST [ MCPAbout MCP seen]. Distribution: Colombia, Venezuela.

Comment. Not Philonthus  , to be placed in a further study.

Philonthus duploseriatus Solsky 1868 

Philonthus duploseriatus Solsky 1868: 136  . Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #?ST. Distribution: Mexico. Notes: distribution within Mexico ( Dugès & Villada 1901, Navarrete-Heredia et al. 2002); probably not Philonthus  (AKH pers. obs.).

Philonthus eryx Smetana 1995 

Philonthus eryx Smetana 1995: 306  . Type locality: California, Ventura Co., Fillmore, Santa Clara River. Type depository: AMNHAbout AMNH HT ♂, etc. Distribution: USA, Mexico. Note: distribution, also within Mexico (Navarrete- Heredia et al. 2002).

Philonthus excellens Bernhauer 1916 

Philonthus excellens Bernhauer 1916a: 37  . Type locality: West-Columbien: Umgebung von Cali. Type depository: FMNHAbout FMNH HT. Distribution : Colombia  .

Comment. Not Philonthus  , to be placed in a further study.

Philonthus exemplaris Wendeler 1956 

Philonthus exemplaris Wendeler 1956: 231  . Type locality: [no specific locality]. Type depository: ZMHB 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution : Brazil  . Note: distribution within Brazil unknown (Asenjo et al. 2013).

Philonthus fassli Bernhauer 1916 

Philonthus fassli Bernhauer 1916a: 29  . Type locality: Huancabamba , Peru  . Type depository: FMNHAbout FMNH HT [not seen]. Distribution : Peru  . Notes: Peru ( Asenjo 2004); probably not Philonthus  .

Philonthus fenyesi Bernhauer 1910 

Philonthus fenyesi Bernhauer 1910: 381  . Type locality: Cordoba, Veracruz  . Type depository: FMNH 2View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus feralis Erichson 1840 

Philonthus feralis Erichson 1840: 469  . Type locality: Columbia. Type depository: ZMHB LT ♂, 2 PLT. Distribution: Mexico, Guatemala, Costa Rica, Panama, Colombia, Venezuela, Peru, Brazil; Argentina probably in error. Notes: Argentina in Blackwelder (1944); valid or synonym of P. longicornis? (Navarrete-Heredia 1995)  ; characters, lectotype designation, valid species ( Smetana 1994: 346); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); redescription, distribution not including southern South America (Chani- Posse 2010: 65); distribution including Peru, also within Brazil (Asenjo et al. 2013).

Philonthus ferreipennis ferreipennis Horn 1884  [not in CASA]

Philonthus ferreipennis Horn 1884: 202  . Type locality: [no specific locality; not orig. fixed (Washington Territory to the southern part of California)]. Type depository: MCZAbout MCZ (Horn) LT ♂, 4 PLT; (LeConte) 3 PLT. Distribution: Canada, USA (northwestern, to central California). Notes: ssp. ferreipennis (Horn)  , lectotype designation, redescription, separation and distribution of subspecies, distribution ( Smetana 1995: 348). Represented in CASA ( Mexico) only by the subspecies P. f. stigmatias Smetana 1995  , see below.

Philonthus ferreipennis stigmatias Smetana 1995 

Philonthus ferreipennis stigmatias Smetana 1995: 350  . Type locality: Mt. Laguna, San Diego Co., California. Type depository: CNC HT ♂; 150 PT in many collections. Distribution: USA (southwestern), Mexico. Notes: ssp. stigmatias ( Smetana 1995: 349)  ; distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus fibularius Erichson 1840 

Philonthus fibularius Erichson 1840: 442  . Type locality: Mexico. Type depository: ZMHB 1 ST [ MCPAbout MCP seen]. Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

= Philonthus boucardi Solsky 1868: 127  . Type locality: Mexico, probably the state of Oaxaca. Type depository: ZINAbout ZIN? (Solsky) #?ST. Note: probably P. fibularius ( Sharp 1885: 404)  ; as synonym of P. fibularius ( Bernhauer & Schubert 1914: 338)  .

Philonthus figulus Erichson 1840 

Philonthus figulus Erichson 1840: 464  . Type locality: Brasilia. Type depository: ZMHB LT ♂. Distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Uruguay, Argentina, Cuba, Puerto Rico, St. Vincent, Trinidad. Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002), Cuba ( Peck 2005), Lesser Antilles ( St. Vincent) ( Peck 2010); lectotype designation, redescription, distribution ( Chani-Posse 2010: 28); Peru ( Asenjo 2004), also within Brazil (Asenjo et al. 2013).

= Philonthus amazonicus Sharp 1876a: 165  . Type locality: Pará & Ega [ Brazil]  . Type depository: BMNH 6 ST. Distribution : Brazil. Note: as synonym of P. figulus ( Sharp 1885: 415)  . 

Philonthus fissilis Sharp 1885 

Philonthus fissilis Sharp 1885: 399  . Type locality: Guatemala: Totonicapam 8000–10500 ft  . Type depository: BMNH 2 ST ♂. Distribution : Guatemala  .

Philonthus flavolimbatus Erichson 1840 

Philonthus flavolimbatus Erichson 1840: 471  . Type locality: Caracas [LT]; "Columbia" [orig. publication]. Type depository: ZMHB LT ♂, 2 PLT. Distribution: USA, Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Peru, Bolivia, Brazil, Paraguay, Uruguay, Argentina, Martinique, Grenadines. Notes: lectotype designation Smetana 1995: 201); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); Peru ( Asenjo 2004); distribution including Uruguay (Chani-Posse 2004); redescription, distribution ( Smetana 1995: 200, Chani-Posse 2010: 29); distribution, also within Lesser Antilles ( Martinique) ( Peck 2011b), within Brazil (Asenjo et al. 2013); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Philonthus apicipennis Lynch-Arribálzaga 1884: 155  , attributed to Fauvel. Type locality: "America meridionali et boreali" (after Fauvel in litteris), but seen only from Chacabuco (Buenos Aires prov.) [orig. publication]. Type depository: MACNAbout MACN LT ♀, 3 PLT. Notes: lectotype designation ( Chani-Posse 2010: 30). Distribution: Argentina. Note: as synonym of P. flavolimbatus ( Fauvel 1887: 232)  .

= Philonthus wacoensis Casey 1915: 430  . Type locality: Waco, Texas. Type depository: USNM LT ♂. Notes: lectotype designation, as synonym of P. flavolimbatus ( Smetana 1995: 200)  .

Philonthus flohri Sharp 1876 

Philonthus flohri Sharp 1876b: 429  . Type locality: neighborhood of the city of Mexico. Type depository: BMNH 5 ST. Distribution: Mexico. Notes: as variety of P. furvus ( Blackwelder 1944)  ; distribution ( Scheerpeltz 1971b: 205); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus floresi Chani-Posse 2010 

Philonthus floresi Chani-Posse 2010: 31  . Type locality: Argentina, Tucumán  . Type depository: MACNAbout MACN HT ♂; IADIZA 1 PT; AMNH 2View Materials PT. Distribution : Argentina  .

Philonthus formicarius Bernhauer 1908 

Philonthus formicarius Bernhauer 1908: 334  . Type locality: Mar d'Hespanha , Minas Gerais, Brazil  . Type depository: FMNHAbout FMNH HT. Distribution : Brazil (Minas Gerais)  . Note: with leaf cutter ant ( Bernhauer 1908); distribution within Brazil (Asenjo et al. 2013).

Philonthus forsteri Scheerpeltz 1960 

Philonthus forsteri Scheerpeltz 1960: 104  . Type locality: Bolivien, Songtal , Cuticucho, etwa 3700m  . Type depository: NMWAbout NMW ( Scheerpeltz ) 3 ST. Distribution: Bolivia  .

Philonthus franzi ( Sáiz 1971) 

Endeius franzi Sáiz 1971: 345  . Type locality: Santiago: El Arrayán. Type depository: Sáiz (in MNNC) HT ♂, 4 PT. Distribution: Chile. Notes: redescription, distribution (Chani-Posse de Maus 2008: 398); comb. nov. ex Endeius (Chani-Posse 2013: 404)  .

Philonthus fulgipennides Newton 2017 

Philonthus fulgipennides Newton 2017: 15  (new name for Philonthus fulgipennis Sharp 1885: 410  ).

Philonthus fulgipennis Sharp 1885: 410  (preoccupied, junior secondary homonym in Philonthus  of Staphylinus fulgipennis Guérin-Méneville 1830  : pl.1). Type locality: Panama, Chiriqui, Volcan de Chiriqui below 4000 ft, Boquete 3500 ft. Type depository: BMNH 29 ST: 23 Chiriqui, 6 Boquete [MCP seen]. Distribution: Panama. Notes: within the Neotropical lineage ( Chani-Posse et al. 2018); not Philonthus  , to be placed in a further study within the Neotropical lineage.

Type material examined. Syntypes ( BMNH): 3 card-mounted on one card, “ Philonthus fulgipennis  / Types D.S./ Volcan de Chiriqui Champion [in Sharp’s hand] // Type (round white label with orange margin) // V. de Chiriqui below 4,000 ft. Champion (white label) // B. C. A. Col. I. 2. Philonthus fulgipennis Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”; 3 card-mounted on one card, “ Philonthus fulgipennis  D.S./ Volcan de Chiriqui Champion [in Sharp’s hand] // V. de Chiriqui below 4,000 ft. Champion (white label) // B. C. A. Col. I. 2. Philonthus fulgipennis Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”; 2 cardmounted on one card and one additional specimen with same locality and collection data as above.

Philonthus furvus Nordmann 1837 

Philonthus furvus Nordmann 1837: 88  . Type locality: Mexico. Type depository: ZMHB 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution: Canada, USA, Mexico, Guatemala, Costa Rica. Notes: redescription ( Smetana 1995: 92); Mexican records ( Sharp 1885, Smetana 1991, 1995, Dugès & Villada 1901); distribution within Mexico ( Navarrete-Heredia et al. 2002).

= Philonthus californicus Mannerheim 1843: 230  . Type locality: California. Type depository: MZHF LT ♂, 1 PLT. Notes: lectotype designation, new synonym of P. furvus ( Smetana 1995: 92)  .

= Philonthus comptus Haldeman 1852: 375  . Type locality: Santa Fé or "journey across the plains" to it [USA: New Mexico]. Type depository: ?? HT. Note: as synonym of P. furvus ( Bernhauer & Schubert 1914: 339)  .

Philonthus gaudichaudii Blanchard 1842 

Philonthus gaudichaudii Blanchard 1842: 81  . Type locality: Rio de Janeiro  . Type depository: MNHNAbout MNHN?. Distribution : Brazil (Rio de Janeiro)  . Note: distribution within Brazil (Asenjo et al. 2013).

Philonthus gentilis Horn 1884 

Philonthus gentilis Horn 1884: 185  . Type locality: Arizona [no specific locality]. Type depository: MCZAbout MCZ (Horn) LT ♂, 5 PLT; (LeConte) 4 PLT. Distribution: USA, Mexico. Note: lectotype designation, redescription ( Smetana 1995: 160); distribution, also within Mexico ( Dugès & Villada 1901, Smetana 1995, Navarrete-Heredia et al. 2002).

Philonthus gracillimus Sharp 1876 

Philonthus gracillimus Sharp 1876a: 168  . Type locality: Ega. Type depository: BMNH HT. Distribution : Brazil (Amazonas)  . Note: distribution within Brazil (Asenjo et al. 2013).

Philonthus grandicollis Horn 1884 

Philonthus grandicollis Horn 1884: 193  . Type locality: [no specific locality; not orig. fixed (California to Vancouver)]. Type depository: MCZAbout MCZ (Horn) LT ♂, 5 PLT; (LeConte) 1 PLT. Distribution: Canada, USA, Mexico. Notes: lectotype designation, redescription, distribution ( Smetana 1995: 300); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus gratiosus Erichson 1840 

Philonthus gratiosus Erichson 1840: 461  . Type locality: Bogotae in Columbia interiore. Type depository: ZMHB 3 ST [ MCPAbout MCP seen]. Distribution: Colombia  .

Comment. Not Philonthus  , to be placed within the Neotropical lineage.

Philonthus griseolus Sharp 1885 

Philonthus griseolus Sharp 1885: 399  . Type locality: Costa Rica: Irazu 6000–7000 ft  . Type depository: BMNH HT ♂. Distribution : Costa Rica  .

Philonthus guatemalenus Sharp 1885 

Philonthus guatemalenus Sharp 1885: 403  . Type locality: Guatemala: San Gerónimo   . Type depository: BMNH HT ♀. Distribution : Guatemala  ; Costa Rica. Note: misspelling “ guatemalensis ” ( Bernhauer & Schubert 1914: 340).

Philonthus guerreroensis Bernhauer 1910 

Philonthus guerreroensis Bernhauer 1910: 380  . Type locality: Omiltemi 8000 ft., Guerrero  . Type depository: FMNH 1View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus hades Chatzimanolis & Engel 2013 

Philonthus hades Chatzimanolis & Engel 2013: 288  . Type locality: Dominican Republic, Santiago [ Oligo-Miocene amber]  . Type depository: AMNHAbout AMNH HT ♀, 1 PT. Distribution: Dominican Rep. [ Miocene fossil]. 

Philonthus havaniensis (Laporte de Castelnau 1835) 

Staphylinus havaniensis Laporte de Castelnau 1835: 116  . Type locality: Ile de Cuba. Type depository: BMNH / MVMA?. Distribution: Cuba, Jamaica, Hispaniola, Puerto Rico, Antigua, Grand Cayman, Mona, Montserrat, St. John, St. Croix. Notes: distribution, also within Cuba ( Peck 2005), Lesser Antilles ( Peck 2011a: Northern Leeward Is.), Cayman Is. (Frank et al. 2011).

= Philonthus obscurus Fauvel 1863: 434  , attributed to Chevrolat (preoccupied, junior secondary homonym in Philonthus  of Staphylinus obscurus Gravenhorst 1802:174  ). Type locality: [no specific locality; Cuba implied]. Type depository: IRSNB 2 ST [MCP seen]. Distribution: Cuba.

Philonthus hepaticus Erichson 1840 

Philonthus hepaticus Erichson 1840: 451  . Type locality: America septentrionali. Type depository: ZMHB LT ♂, 3 PLT. Distribution: Canada, USA, Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Bolivia, Brazil, Chile, Argentina, Cuba, Jamaica, Hispaniola, Puerto Rico, Virgin Is., Bahamas, Lesser Antilles, Trinidad; adventive in Australia, New Zealand. Notes: lectotype designation, redescription ( Smetana 1995: 176); distribution ( Smetana 1995, Chani-Posse 2004, Dugès & Villada 1901, Peck 2010; Chile, Arg.? misidentification, teste Smetana 1995: 177, 179); distribution, also within Mexico ( Navarrete-Heredia et al. 2002), within Cuba ( Peck 2005), within Dominica (Peck 2006), Barbados ( Peck 2009a), Lesser Antilles ( Peck 2010: St. Vincent, Peck 2011a: Northern Leeward Is., Peck et al. 2014: Guadeloupe Archipelago); redescription, distribution ( Chani-Posse 2010: 32); distribution within Brazil (Asenjo et al. 2013).

= Philonthus vilis Erichson 1840: 451  . Type locality: Americae meridionalis  ins. St. Thomae. Type depository: ZMHB LT ♂, 3 PLT. Distribution: Lesser Antilles (St. Thomas I.). Notes: as synonym of P. hepaticus ( Blackwelder 1943: 401)  ; lectotype designation ( Smetana 1995: 176).

= Philonthus nanus Melsheimer 1844: 36  . Type depository: MCZAbout MCZ LT ♂, 5 PLT. Type locality: Pennsylvania. Distribution: USA (Pennsylvania). Notes: as synonym of P. hepaticus ( Gemminger & Harold 1868: 589)  ; lectotype designation ( Smetana 1995: 177).

= Philonthus nanus var. palleolus Melsheimer 1844: 37  . Type depository: MCZ? #?ST. Type locality: [not listed, presumed to be Pennsylvania]. Distribution: USA. Notes: as synonym of P. hepaticus ( Gemminger & Harold 1868: 589)  .

= Philonthus cinctutus Melsheimer 1844: 37  . Type depository: MCZAbout MCZ LT ♀, 2 PLT. Type locality: Pennsylvania. Distribution: USA (Pennsylvania). Notes: probably P. hepaticus ( Horn 1884: 223)  ; as synonym of P. hepaticus ( Fauvel 1891: 112)  ; lectotype designation ( Smetana 1995: 177).

= Philonthus varicolor Boheman 1858: 29  . Type depository: NHRS 1 ST. Type locality: California. Distribution: USA (California). Note: as synonym of P. hepaticus ( Fauvel 1891: 112)  .

= Philonthus parvimanus Sharp1885: 406  . Type locality: Mexico: Cordova; Guatemala near the city, Paso Antonio, Champerico, San Joaquin in Vera Paz; Nicaragua: Chontales; Panama: David. Type depository: BMNH 12+ ST: from all type localities [MCP seen]. Distribution: Mexico etc. Notes: as synonym of P. hepaticus ( Fauvel 1891: 112)  ; synonymy needs checking ( Smetana 1995: 174).

= Philonthus oklahomensis Hatch & Ortenburger 1930: 12  . Type depository: USNM HT ♂. Type locality: Dougherty , Murray Co., Oklahoma. Distribution: USA (Oklahoma). Note  : type data, new synonym of P. hepaticus ( Smetana 1995: 177)  .

Philonthus heteropus Sharp 1887 

Philonthus heteropus Sharp 1887: 788  . Type locality: Mexico (near the city)  , Mexico  . Type depository: BMNH HT ♀. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus hirtiventris Sharp 1885 

Philonthus hirtiventris Sharp 1885: 410  . Type locality: Mexico  : Oaxaca, Mirador  . Type depository: BMNH 2 ST: 1 ♂, 1 ♀. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus hoegei Sharp 1885 

Philonthus hoegei Sharp 1885: 400  . Type locality: Mexico: Las Vigas, Milpas. Type depository: BMNH 26 ST: 23 Las Vigas, 3 Milpas [MCP seen]. Distribution: Mexico. Notes: redescription, distribution ( Márquez & Asiain 2010); key ( Navarrete-Heredia 2003); distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2004, 2006).

Philonthus idoneus Sharp 1885 

Philonthus idoneus Sharp 1885: 411  . Type locality: Guatemala, El Tumbador 2500 ft  . Type depository: BMNH HT ♂. Distribution : Guatemala  .

Philonthus imperialis Bernhauer 1916 

Philonthus imperialis Bernhauer 1916a: 28  . Type locality: West-Columbien: Umgebung von Cali (St. Antonio,

2000m). Type depository: FMNHAbout FMNH HT ♀. Distribution : Colombia  . Note: type material not seen (on loan); after original description probably not Philonthus  .

Philonthus incertus Solsky 1868 

Philonthus incertus Solsky 1868: 135  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZINAbout ZIN? (Solsky) #?ST. Distribution: USA, Mexico. Notes: redescription, distribution ( Smetana 1995: 221); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

= Philonthus gracilicornis Horn 1884: 190  (preoccupied, junior primary homonym of Philonthus gracilicornis Heer 1839: 274  ). Type locality: Arizona. Type depository: MCZAbout MCZ (Horn) LT ♂, 5 PLT, (LeConte) 8 PLT?. As synonym of P. incertus ( Sharp 1887: 788)  ; lectotype designation ( Smetana 1995: 222).

Philonthus indigaceus Fauvel 1891 

Philonthus indigaceus Fauvel 1891: 110  . Type locality: Caracas & Nouvelle-Grenade [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 2 ST [one without head, MCPAbout MCP seen]. Distribution: Colombia, Venezuela.

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus iopterus Erichson 1840 

Philonthus iopterus Erichson 1840: 928  . Type locality: Bogotae in Columbia interiore. Type depository: ZMHB 3 ST: ♂ [ MCPAbout MCP seen]. Distribution: Colombia, Venezuela, Peru. Note: Peru ( Plavilstschikov 1929).

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus iris Sharp 1885 

Philonthus iris Sharp 1885: 401  , attributed to Dugès in litteris. Type locality: Mexico: Cuernavaca, Capulalpam. Type depository: BMNH 2 ST: 1 Cuernavaca, 1 Capulalpam [MCP seen]. Distribution: Mexico. Notes: Durango in error (Navarrete-Heredia 1995, reference and locality refer to P. iteratus  , not P. iris  ); distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).

Philonthus jalapensis Bernhauer 1910 

Philonthus jalapensis Bernhauer 1910: 382  . Type locality: Jalapa, Veracruz. Type depository: ?? #?ST (not at FMNH). Distribution: Mexico. Note: to Belonuchus  ? (after Smetana 1995); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus jenseni Bernhauer 1912 

Philonthus jenseni Bernhauer 1912a: 42  . Type locality: Argentinien: Chaunar-Region [orig. pub.], "Channar Reg." [LT]. Type depository: FMNH LT ♀, 4 PLT; MACN 1 PLT. Distribution: Argentina. Notes: lectotype designation, redescription, distribution ( Chani-Posse 2010: 35).

Philonthus klimschi Bernhauer 1916 

Philonthus klimschi Bernhauer 1916a: 38  . Type locality: Brasilien: S. Catharina. Type depository: FMNHAbout FMNH HT ♂  . Distribution: Brazil (Santa Catarina). Note: distribution within Brazil (Asenjo et al. 2013); 

Philonthus labilis Erichson 1840 

Philonthus labilis Erichson 1840: 500  . Type locality: Guyana Venezuelensi  . Type depository: ZMHB #? ST. Distribution : Venezuela  .

Philonthus laetipennis Sharp 1885 

Philonthus laetipennis Sharp 1885: 409  . Type locality: Mexico: Misantla   , Veracruz  . Type depository: BMNH 3 ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus laeviventris Schubert 1909 

Philonthus laeviventris Schubert 1909: 291  . Type locality: Jalapa region  , Veracruz, Mexico. Type depository: 

ZMHB (Schubert) 1 ST ♂ [ MCPAbout MCP seen]; FMNHAbout FMNH 1ST. Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus lecontei Horn 1884 

Philonthus lecontei Horn 1884: 222  . Type locality: " California ". Type depository: MCZAbout MCZ (Horn) LT ♂, 3 PLT; (LeConte) 1 PLT. Distribution: Canada, USA, Mexico. Notes: lectotype designation, redescription, distribution ( Smetana 1995: 467); Mexico "northern Mexico " ( Casey 1915: 442); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus limaensis Solsky 1872 

Philonthus limaensis Solsky 1872: 300  . Type locality: Lima  . Type depository: ZINAbout ZIN? (Solsky) #? ST. Distribution : Peru. Note: Peru ( Asenjo 2004). 

Philonthus lineatoventris Bruch 1915  , nomen nudum

Philonthus lineatoventris Bruch 1915: 502  , attributed to Bernhauer in litteris; nomen nudum. Locality: Rio Negro. Distribution: Argentina. Note: not in Fernández et al. (2007) or Bachmann et al. (2017).

Philonthus loensis ( Coiffait & Sáiz 1968) 

Endeius loensis Coiffait & Sáiz 1968: 357  . Type locality: Guatín, Antofagasta, Chile. Type depository: MNNCAbout MNNC HT ♂ (teste Chani-Posse de Maus 2008: 390) + 2 PT; Sáiz (in MNNCAbout MNNC) 12 PT. Distribution: Chile. Notes: redescription (Chani-Posse de Maus 2008: 388); comb. nov. ex Endeius (Chani-Posse 2013: 404)  ; distribution within Chile ( Ferrú & Elgueta 2011).

Philonthus lomatus Erichson 1840 

Philonthus lomatus Erichson 1840: 482  . Type locality: America septentrionali. Type depository: ZMHB LT ♂, 5 PLT. Distribution: Canada, USA, Haiti. Note: Haiti ( Frank 1983); lectotype designation, redescription, distribution ( Smetana 1995: 390).

= Philonthus georgianus Sachse 1852: 143  . Type locality: Georgia. Type depository: ?? (Sachse) #?ST. Distribution: USA ( Georgia). Note: as synonym of P. lomatus ( Fauvel 1869: 488)  .

Philonthus longicornis Stephens 1832  [adventive? in CASA]

Philonthus longicornis Stephens 1832: 237  , attributed to Kirby MSS. Type locality: near London. Type depository: BMNH? (Stephens) #?ST. Distribution: Palaearctic, Oriental, Korea (S); adventive? in Canada, USA, Mexico, Costa Rica, Colombia, Peru, Bolivia, Brazil, Uruguay, Argentina, Chile, Juan Fernández Is, Cuba, Haiti, Puerto Rico, Lesser Antilles, Australia, New Zealand, New Caledonia, Norfolk I., Tahiti, Ethiopia, South Africa, Madagascar, Réunion, Mauritius. Notes: “cosmopolitan” ( Herman 2001d); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution including Brazil (Chani-Posse 2004); redescription ( Smetana 1995: 244, Chani-Posse 2010: 35, Hromádka 2012a); Lesser Antilles ( Peck 2011a: Northern Leeward Is.), Peru ( Asenjo 2004), also within Brazil (Asenjo et al. 2013); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Philonthus perplexus Fairmaire & Germain 1862: 431  . Type locality: Santiago, Chile. Type depository: MNHN LT ♂, 2 PLT. Distribution: Chile, Easter Island. Notes: lectotype designation, resynonymized (Chani-Posse de Maus 2009: 37); ex synonym of P. longicornis  , distribution, validity ( Coiffait & Sáiz 1968: 360).

= Philonthus fumosus Solsky 1868: 134  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]  . Type depository: ZINAbout ZIN? ( Solsky ) #? ST. Distribution: Mexico. Note: as synonym of P. longicornis ( Sharp 1885: 416)  . 

Philonthus lucidanus Blackwelder 1944 

Philonthus lucidanus Blackwelder 1944: 134  (new name for Philonthus lucidus Fauvel 1891: 116  ). Distribution: Colombia, Venezuela.

= Philonthus lucidus Fauvel 1891: 116  (preoccupied, junior secondary homonym in Philonthus  of Staphylinus lucidus Gravenhorst 1802: 21  ). Type locality: Colonie Tovar, Lagonita & Nouvelle-Grenade [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 3 ST [ MCPAbout MCP seen].

Philonthus lugubris ( Sáiz 1971) 

Endeius lugubris Sáiz 1971: 347  . Type locality: Coquimbo: Río Los Molles   . Type depository: Sáiz (in MNNCAbout MNNC) HT ♂, 6 PT. Distribution: Chile. Notes: redescription, distribution ( Chani-Posse de Maus 2008: 399); comb. nov. ex Endeius (Chani-Posse 2013: 404)  . 

Philonthus lynchi Bernhauer & Schubert 1914 

Philonthus lynchi Bernhauer & Schubert 1914: 345  (new name for Philonthus affinis Lynch-Arribálzaga 1884: 160  ). Distribution: Peru, Paraguay, Uruguay, Argentina. Notes: redescription, distribution including Uruguay, Paraguay, Peru ( Chani-Posse 2010: 38).

= Philonthus affinis Lynch-Arribálzaga 1884: 160  (preoccupied, junior primary homonym of Philonthus affinis Roth 1851: 118  ). Type locality: Baradero. Type depository: MACN HT ♀. Distribution: Argentina.

Philonthus macilentus Sharp 1885 

Philonthus macilentus Sharp 1885: 423  . Type locality: Mexico  : Cordova  . Type depository: BMNH 2 ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus melampus Smetana 1991 

Philonthus melampus Smetana 1991: 236  . Type locality: Mexico  : Chiapas, Mt  . Tzontehuitz, 9500 ft. Type depository: CNCAbout CNC HT ♂, 25 PT. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus melancholicus Sharp 1885 

Philonthus melancholicus Sharp 1885: 425  . Type locality: Mexico: Capulalpam (Oaxaca?). Type depository: BMNH HT ♀  . Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus melanopus Sharp 1885 

Philonthus melanopus Sharp 1885: 405  . Type locality: Mexico: Jalapa; Costa Rica: Volcan de Irazu 6000–7000 ft; Panama: Volcan de chiriqui, 2500–4000 ft. Type depository: BMNH 6 ST: all type localities [ MCPAbout MCP seen]. Distribution: Mexico, Costa Rica, Panama. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus metallifer Bernhauer 1916 

Philonthus metallifer Bernhauer 1916a: 31  , attributed to Fauvel in litteris. Type locality: Bolivien & Peru. Type depository: FMNHAbout FMNH 2 ST. Distribution: Bolivia, Peru. Note: Peru ( Asenjo 2004).

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus mnemon Smetana 1995 

Philonthus mnemon Smetana 1995: 306  . Type locality: Arizona, Chiric Mts., Ash Spg., 7mi SW Portal, 6000 ft. Type depository: CNCAbout CNC HT ♂, etc. Distribution: USA, Mexico. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus mollis Sharp 1885 

Philonthus mollis Sharp 1885: 408  . Type locality: Panama: San Feliz. Type depository: BMNH 4 ST. Distribution   : Panama.

Philonthus monachus Bernhauer 1908 

Philonthus monachus Bernhauer 1908: 335  . Type locality: Paraguay. Type depository: FMNHAbout FMNH 1 ST. Distribution: Paraguay, Brazil (based on conspecific material in FMNHAbout FMNH). Note: Brazil (São Paulo, Rio de Janeiro); not in Asenjo et al. (2013).

Philonthus morosus Casey 1915 

Philonthus morosus Casey 1915: 441  . Type locality: Lake & San Francisco Cos., California  . Type depository:

USNMAbout USNM LT ♀, 2 PLT. Distribution: Canada, USA, Mexico. Notes: lectotype designation, redescription, distribution in Mexico ( Smetana 1995: 452); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus navarretei Márquez & Asiain 2014 

Philonthus navarretei Márquez & Asiain 2014: 297  , availability of Philonthus navarretei Márquez & Asiain 2010: 280  by designation of type depository. Type locality: México: Oaxaca: Santiago Yosondua, camino a Santa María Yolotepec, 2398m, N16°51'13", W97°33'43.6". Type depository: UAEH HT ♂, 31 PT; CZUG 2 PT; ZMUC 1 PT. Distribution: Mexico.

= Philonthus navarretei Márquez & Asiain 2010: 280  , unavailable, no holotype depository indicated (Art. 16.4.2). Note: noted as nomen nudum in Navarrete-Heredia & Newton (2013: 132).

Philonthus neptunus Smetana 1995 

Philonthus neptunus Smetana 1995: 436  . Type locality: Florida, Monroe Co., Middle Torch Key, Sec. 17 Lazelle Place. Type depository: CNCAbout CNC HT ♂, etc. Distribution: USA, Bahamas (South Bimini); Jamaica.

Philonthus nigerrimus Erichson 1840 

Philonthus nigerrimus Erichson 1840: 434  . Type locality: Mexico. Type depository: ZMHB LT, 1 PLT. Distribution: Mexico. Note: lectotype designation, redescription and distribution ( Smetana 1991: 235); distribution within Mexico ( Sharp 1885, Dugès & Villada 1901, Smetana 1991, Navarrete-Heredia et al. 2002).

Philonthus occultus Sharp 1885 

Philonthus occultus Sharp 1885: 407  . Type locality: Guatemala  . Type depository: BMNH HT ♂. Distribution : Guatemala  .

Philonthus ochromerus Sharp 1885 

Philonthus ochromerus Sharp 1885: 406  . Type locality: Mexico: Cordova; Guatemala near the city, San Gerónimo, Zapote, Capetillo, Calderas, Cerro Zunil, Lanquin, Senahu, San Joaquin in Vera Paz; Panama: Bugaba, Volcan de Chiriqui 2000–4000 ft. Type depository: BMNH 12+ST: several, from all type localities [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala, Costa Rica, Panama. Notes: Costa Rica ( Champion 1913); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus oenotrus Smetana 1991 

Philonthus oenotrus Smetana 1991: 228  . Type locality: Mexico: Chiapas, Mt. Tzontehuitz, 9500 ft Type depository: CNC HT ♂, 3 PT. Distribution: Mexico. Note: Guanajuato in error (Navarrete-Heredia 1995, reference and locality refer to “ optatus  ”, not “ oenotrus  ”); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus opacigaster ( Bierig 1940) 

Philonthus (Gabrius) opacigaster Bierig 1940: 143  . Type locality: Volcán de Chiriqui, como 1500m, Panamá  . Type depository: FMNHAbout FMNH HT. Distribution : Panama  .

Philonthus ophthalmicus Bernhauer 1941 

Philonthus ophthalmicus Bernhauer 1941: 288  , also 1951: 278. Type locality: Ica, s Peru  . Type depository: FMNH 1View Materials ST. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Philonthus orphanus Erichson 1840 

Philonthus orphanus Erichson 1840: 452  (senior primary homonym of Philonthus orphanus Erichson 1840: 499  ). Type locality: Caracas Columbiae. Type depository: ZMHB LT ♂, 3 PLT. Distribution: Venezuela. Notes: as valid species, lectotype designation, redescription and distribution ( Smetana 1992: 135).

Philonthus palcoensis Bernhauer 1941 

Philonthus palcoensis Bernhauer 1941: 288  , also 1951: 277. Type locality: Palco, 2600m, etc., s Peru. Type

depository: FMNH 1View Materials ST. Distribution : Peru  . Note : Peru ( Asenjo 2004). Note: type material not seen (on loan); after original description probably not Philonthus  . 

Philonthus pallipes Blanchard 1842 

Philonthus pallipes Blanchard 1842: 82  . Type locality: "îles du Parana, en face de Corrientes " [orig. pub., Herman 2001d]; " Montevideo " [LT, but judged in error, Chani-Posse 2010: 42]. Type depository: MNHNAbout MNHN LT ♀. Distribution: Colombia, Venezuela, Ecuador, Bolivia, Brazil, Paraguay, Uruguay, Argentina, Cuba. Notes: lectotype designation, redescription, distribution including Colombia, Cuba etc. ( Chani-Posse 2010: 40); distribution within Brazil (Asenjo et al. 2013).

= Philonthus depressus Steinheil 1869: 253  . Type locality: Porfido , Sierra de Mendoza [ Argentina]  . Type depository: MNHNAbout MNHN? HT ♀. Note : as synonym of P. pallipes ( Bernhauer & Schubert 1914: 349)  . 

= Philonthus pallidipes  ; Lynch-Arribálzaga 1884: 152, attributed to Blanchard; emendation or misspelling of pallipes  ?. Type locality: Republica Argentina prope Corrientes. Note: as synonym of P. pallipes ( Bernhauer & Schubert 1914: 349)  .

Philonthus palmatus Sharp 1885 

Philonthus palmatus Sharp 1885: 397  . Type locality: Mexico  : Jalapa, Oaxaca, La Parada. Type depository: BMNH 4 ST: 2 Jalapa, 1 Oaxaca, 1 Parada [ MCPAbout MCP seen]. Distribution : Mexico  ; Guatemala  . Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002). Note: Generic affiliation unclear.

Philonthus pauper Bernhauer 1916 

Philonthus pauper Bernhauer 1916a: 32  . Type locality: West-Columbien: Umgebung von Cali im Caucatale (Alto de las cruces, 2200m)  . Type depository: FMNH 2View Materials ST. Distribution : Colombia  . Note: type material not seen (on loan); generic affiliation unclear after original description, probably within the Neotropical lineage.

Philonthus pauxillus Solsky 1868 

Philonthus pauxillus Solsky 1868: 133  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZINAbout ZIN? (Solsky) #?ST. Distribution: USA, Mexico, Panama, Venezuela, Trinidad, Brazil, Paraguay, Argentina, Galapagos Is. Notes: redescription ( Smetana 1995: 179, Chani-Posse 2010: 43); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution including Argentina (Chani-Posse 2004), also Venezuela, Brazil, Paraguay ( Chani-Posse 2010); as synonym of P. hepaticus ( Blackwelder 1944)  but valid ( Smetana 1995); distribution within Galápagos Is. ( Newton & Peck 2006); distribution within Brazil (Asenjo et al. 2013).

Philonthus pavidus Erichson 1847 

Philonthus pavidus Erichson 1847: 89  . Type locality: [ Peru, east side]  . Type depository: ZMHB 4 ST [ MCPAbout MCP seen]. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Philonthus peratriceps Scheerpeltz 1933 

Philonthus peratriceps Scheerpeltz 1933: 1355  (new name for Philonthus atriceps Sharp 1885: 404  ). Distribution: Mexico. Notes: distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006); distribution ( Navarrete-Heredia 2012); ex extenal refuse piles Atta mexicana  ( Navarrete-Heredia 2001, 2012).

= Philonthus atriceps Sharp 1885: 404  (preoccupied, junior primary homonym of Philonthus alumnus var. atriceps Horn 1884:199  ). Type depository: BMNH HT ♂. Type locality: Mexico: Jalapa, Las Vigas.

Philonthus piceatus Nordmann 1837 

Philonthus piceatus Nordmann 1837: 85  . Type locality: Mexico. Type depository: ZMHB 1 ST ♂ [ MCPAbout MCP seen]. Distribution: USA, Mexico, Guatemala, Costa Rica, Panama. Notes: Mexican records ( Smetana 1995, Márquez-Luna 1998); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

= Philonthus iridiventris Solsky 1868: 128  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]  . Type depository: ZINAbout ZIN? ( Solsky ) #? ST. Distribution: Mexico. Note: as synonym of P. piceatus ( Sharp 1885: 400)  . 

= Philonthus mexicanus Sharp 1876b: 430  . Type locality: city of Mexico  . Type depository: BMNH 8 ST. Distribution : Mexico. Note: as synonym of P. piceatus ( Sharp 1885: 400)  . 

Philonthus politus ( Linnaeus 1758)  [adventive in CASA]

Staphylinus politus Linnaeus 1758: 422  . Type locality: Europa. Type depository: LSUK 6 ST? (http:// www.linnean-online.org/), not seen by Smetana (1995). Distribution: Europe, north Africa, Canary Is., Madeira, Azores, Iceland, Russia, Armenia, Azerbaijan, Georgia, Turkey, Iran, Turkmenistan, Kazakhstan, Mongolia, China, India (KA); adventive in Canada, USA, Mexico, Venezuela, Chile, Argentina, Australia, New Zealand, Chatham Is. Notes: Argentina ( Scheerpeltz 1972); redescription, distribution, no Mexican records ( Smetana 1995: 73); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); redescription, distribution ( Chani-Posse 2010: 44).

Philonthus pollens Márquez & Asiain 2010 

Philonthus pollens Márquez & Asiain 2010: 272  . Type locality: México  : Morelos: Cempoala [= Zempoala ]. Type depository: CNINAbout CNIN HT ♂  . Distribution: distribution within Mexico (Márquez et al. 2015).

Philonthus prescottianus Sharp 1887 

Philonthus prescottianus Sharp 1887: 787  . Type locality: [no specific locality; not fixed]. Type depository: BMNH 2 ST: 1 Salazar, 1 Amecameca [ MCPAbout MCP seen]. Distribution : Mexico  . Notes: distribution comment (Navarrete- Heredia 1995)  ; distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus prismalis Fauvel 1891 

Philonthus prismalis Fauvel 1891: 114  . Type locality: Colonie Tovar, Lagonita & Nouvelle-Grenade [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 1 ST ♀ [ MCPAbout MCP seen]. Distribution: Colombia, Venezuela.

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus prolixicornis Sharp 1885 

Philonthus prolixicornis Sharp 1885: 424  . Type locality: Guatemala, Torola   . Type depository: BMNH HT ♂. Distribution : Guatemala  .

Philonthus proportionalis Schubert 1909 

Philonthus proportionalis Schubert 1909: 293  . Type locality: Jalapa region  , Veracruz, Mexico  . Type depository: ZMHB ( Schubert ) 1 ST [ MCPAbout MCP seen]. Distribution: Mexico  . Note: distribution within Mexico (Navarrete- Heredia et al. 2002).

Philonthus pulcher Bernhauer 1908 

Philonthus pulcher Bernhauer 1908: 335  . Type locality: Peru  . Type depository: FMNH 1View Materials ST ♂, 1ST ♀. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus pulchricolor Bernhauer 1923 

Philonthus pulchricolor Bernhauer 1923: 63  (new name for Philonthus quadricolor Bernhauer 1916: 30  ). Distribution: Colombia. Note: type material not seen (on loan); generic affiliation unclear after original description.

= Philonthus quadricolor Bernhauer 1916a: 30  (preoccupied, junior primary homonym of Philonthus quadricolor Fauvel 1879: 99  ). Type locality: West-Columbien: Umgebung von Cali im Caucatal (Rio Aquatal, 1800m). Type depository: FMNHAbout FMNH 1ST.

Philonthus punctifer Nordmann 1837 

Philonthus punctifer Nordmann 1837: 107  . Type locality: Mexico. Type depository: ZMHB 1 ST ♀ [ MCPAbout MCP seen]. Distribution: Mexico, Costa Rica. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus punctipennis ( Solier 1849)  Staphylinus punctipennis Solier 1849: 319  . Type locality: Santa Rosa, Chile. Type depository: MNHN?; IRSNB LT

♂, 1 PLT, see Coiffait & Sáiz (1968: 358, from the Fauvel collection, not the original series!). Distribution: Chile, Argentina, Peru. Notes: preoccupied, junior primary homonym of Staphylinus punctipennis Lacordaire 1835: 409  , but not congeneric after 1899, conserved, Opinion 2053 ( ICZN 2003); lectotype designation, redescription, distribution ( Coiffait & Sáiz 1968: 358), distribution ( Scheerpeltz 1972: 20), redescription, distribution (Chani-Posse de Maus 2008: 392); comb. nov. ex Endeius (Chani-Posse 2013: 404)  .

Philonthus pyropterus Kraatz 1859 

Philonthus pyropterus Kraatz 1859b: 12  (new name for Staphylinus rufipennis Solier 1849: 317  ). Distribution: Chile; adventive in New Zealand. Notes: as valid, adventive in New Zealand ( Kuschel 1990); as synonym of P. hepaticus ( Fauvel 1878: 546)  and considered as such by subsequent authors (e.g., Coiffait & Sáiz 1968, Herman 2001d). Smetana (1995: 179) noted " hepaticus  " by Coiffait & Sáiz (1968) is a misidentification. It is unclear whether Coiffait & Sáiz (1968: 355) examined Solier´s type material, but their redescription of P. hepaticus  actually belongs to that of P. bonariensis ( Chani-Posse 2010: 15)  .

Staphylinus rufipennis Solier 1849: 317  (preoccupied, junior primary homonym of Staphylinus rufipennis Fabricius 1801: 597  and junior primary homonym of Staphylinus rufipennis Gravenhorst 1802: 40  ). Type locality: Santa Rosa & Illapel [Chile]. Type depository: MNHN? #? ST.

Philonthus quadraticeps Boheman 1858 

Philonthus quadraticeps Boheman 1858: 28  . Type locality: Buenos Ayres. Type depository: NHRSAbout NHRS LT ♂. Distribution: Bolivia, Brazil, Uruguay, Paraguay, Argentina. Notes: lectotype designation, redescription (Chani-Posse de Maus 2009: 35), distribution including Uruguay, Paraguay, Brazil (Chani-Posse 2004), also Bolivia ( Chani-Posse 2010); distribution within Brazil (Asenjo et al. 2013); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Philonthus tenebrosus Boheman 1858: 28  . Type locality: Buenos Ayres. Type depository: NHRSAbout NHRS LT ♂, 1 PLT. Distribution: Argentina. Note: lectotype designation, as synonym of P. quadraticeps (Chani-Posse de Maus 2009: 35)  .

Philonthus quadrulus Horn 1884 

Philonthus quadrulus Horn 1884: 221  . Type locality: Arizona; California [no specific locality]. Type depository: MCZAbout MCZ LT ♂, 6 PLT (Horn col.), 1 PLT (LeConte col.). Distribution: USA, Mexico. Note: distribution in Mexico ( Horn 1894); lectotype designation, redescription, no Mexican records given ( Smetana 1995: 314); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

= Philonthus sagax Casey 1915: 445  . Type locality: Nogales , Arizona. Type depository: USNMAbout USNM LT ♂  . Distribution: USA (Arizona). Notes : lectotype designation, new synonym of P. quadrulus ( Smetana 1995: 314)  . 

Philonthus rectangulus Sharp 1874  [adventive in CASA]

Philonthus rectangulus Sharp 1874: 42  . Type locality: Nagasaki [ Japan]. Type depository: BMNH 22 ST: 22 from Japan, but nothing about " Nagasaki " [ MCPAbout MCP seen]. Distribution: Europe, Russia, Azores, Madeira, Canary Is., Turkey, Afghanistan, Kazakhstan, Japan, Korea (S), China, Taiwan, Thailand, Nepal, Bhutan, Indonesia, New Guinea; adventive in Tahiti, Hawaii, New Zealand, Canada, USA, Mexico, Brazil, Chile, Argentina, Uruguay, Cuba, Costa Rica etc. Notes: Chile ( Coiffait & Sáiz 1968); New Zealand ( Kuschel 1990); Argentina, Cuba, Costa Rica, Uruguay (Chani-Posse 2004); redescription, distribution ( Smetana 1995: 216, Chani-Posse 2010: 47); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution within Brazil (Asenjo et al. 2013).

Philonthus rectilaterus Sharp 1885 

Philonthus rectilaterus Sharp 1885: 425  . Type locality: Mexico: Las Vigas; Guatemala, Dueñas. Type depository: BMNH 12 ST: 10 Las Vigas, 2 Dueñas [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica. Notes: distribution ( Scheerpeltz 1971b: 205); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus rhadamanthus Chatzimanolis & Engel 2013 

Philonthus rhadamanthus Chatzimanolis & Engel 2013: 289  . Type locality: Dominican Republic, Santiago [Oligo- 

Miocene amber]. Type depository: AMNH HT ♀; FMNH 2 PT. Distribution: Dominican Rep. [Miocene fossil].

Philonthus richterianus Bruch 1915  , nomen nudum

Philonthus richterianus Bruch 1915: 503  , attributed to Bernhauer in litteris; nomen nudum. Locality: Misiones. Distribution: Argentina. Note: not in Fernández et al. (2007) or Bachmann et al. (2017).

Philonthus ridiculus Blackwelder 1944 

Philonthus ridiculus Blackwelder 1944: 134  (new name for Philonthus grandiceps Bernhauer 1916a: 33  ). Distribution: Colombia.

= Philonthus grandiceps Bernhauer 1916a: 33  (preoccupied, junior primary homonym of Philonthus grandiceps Kraatz 1859a: 95  and junior primary homonym of Philonthus grandiceps Poppius 1909: 24  ). Type locality: West-Columbien: Umgebung von Cali im Caucatale. Type depository: FMNHAbout FMNH 2 ST.

Comment. Not Philonthus  , to be placed in a further study.

Philonthus riedeli Bernhauer 1916 

Philonthus riedeli Bernhauer 1916a: 39  . Type locality: Brasilien: S. Paulo (Umgebung von Ribeirao Preto Fundão). Type depository: FMNHAbout FMNH HT ♂  . Distribution: Brazil (São Paulo). Note: distribution within Brazil (Asenjo et al. 2013); 

Philonthus rubromaculatus Blanchard 1842 

Philonthus rubromaculatus Blanchard 1842: 82  . Type locality: Montevideo province [orig. pub.]. Type depository: MNHNAbout MNHN LT ♂, 1 PLT. Distribution: Uruguay, Argentina, Chile. Notes: lectotype designation, redescription, distribution including Chile ( Chani-Posse 2010: 50).

Philonthus ruficauda Bernhauer 1934 

Philonthus ruficauda Bernhauer 1934a: 119  . Type locality: Tucuman prov. Argentina. Type depository: FMNHAbout FMNH HT ♀  . Distribution: Argentina.Note: redescription, distribution ( Chani-Posse 2010).

Philonthus rufoplagiatus Solsky 1868 

Philonthus rufoplagiatus Solsky 1868: 130  . Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #?ST. Distribution: Mexico. Notes: as synonym of P. piceatus ( Sharp 1885: 400)  , but as valid? (see Navarrete-Heredia 1995); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus rufotibialis Márquez & Asiain 2010 

Philonthus rufotibialis Márquez & Asiain 2010: 270  . Type locality: Guatemala: San Marcos, 2 km E Tajumulco y 500m al volcán, 2062m, N15°04.99', W91°55.563'. Type depository: MZFCAbout MZFC HT ♂, 1 PT; UAEH 1 PT. Distribution: Mexico, Guatemala.

Philonthus rufulus Horn 1884 

Philonthus alumnus var. rufulus Horn 1884: 199  , attributed to Fauvel. Type locality: Pennsylvania (lectotype). Type depository: MCZAbout MCZ LT ♂, 7 PLT (LeConte col.). Distribution: USA, Mexico, Canada, Mexico, Cuba, Bahamas (South Bimini), Cayman Is. Notes: as P. alumnus  , Mexico distribution ( Horn 1894); valid sp. not synonym of P. alumnus  , lectotype designation, redescription, distribution ( Smetana 1995: 430); distribution, also within Mexico ( Navarrete-Heredia et al. 2002), within Cuba ( Peck 2005); distribution within (and including) Cayman Is. (Frank et al. 2011).

= Philonthus alumnus  ; Horn 1884: 198 (misidentification, not Erichson 1840)," Porto Rico "; Florida to Arizona and as far north as New York. Distribution: USA, Puerto Rico. Note: lectotype designation and redescription for P. alumnus Erichson ( Smetana 1994: 341)  ; correction of misidentification ( Smetana 1995: 434).

= Philonthus alumnus var. atriceps Horn 1884: 199  , attributed to Fauvel. Type locality: New Jersey (lectotype). Type depository: MCZAbout MCZ LT ♂, 7 PLT (LeConte col.). Distribution  : USA. Notes : lectotype designation, as synonym of P. rufulus ( Smetana 1995: 430)  . 

Philonthus sallaei Sharp 1885 

Philonthus sallaei Sharp 1885: 398  . Type locality: Mexico: La Parada  , Oaxaca  . Type depository: BMNH 3 ST: 1 ♂, 2 ♀, from Parada [ MCPAbout MCP seen]. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus scintillatus Blackwelder 1944 

Philonthus scintillatus Blackwelder 1944: 134  (new name for Philonthus scintillans Solsky 1868: 131  ). Distribution: Mexico.

Philonthus scintillans Solsky 1868: 131  (preoccupied, junior secondary homonym in Philonthus  of Staphylinus scintillans Gravenhorst 1806: 70  , not congeneric when replaced before 1961 or since, but replacement name in use (Art. 59.3)). Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #?ST. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus semicupreus Fauvel 1891 

Philonthus semicupreus Fauvel 1891: 114  . Type locality: Colonie Tovar, Lagonita & Nouvelle-Grenade [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 1 ST [ MCPAbout MCP seen]. Distribution: Colombia, Venezuela.

Comment. Not Philonthus  , to be placed in a further study.

Philonthus semicyaneus Schubert 1909 

Philonthus semicyaneus Schubert 1909: 290  . Type locality: Jalapa region  , Veracruz, Mexico  . Type depository: ZMHB ( Schubert ) ST? #?. Distribution: Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus semiruber Horn 1884 

Philonthus semiruber Horn 1884: 187  . Type locality: Arizona; California [no specific locality]. Type depository: IRSNBAbout IRSNB LT ♀, 1 PLT, USNMAbout USNM 1 PLT. Distribution: Canada, USA, Mexico. Note: distribution in Mexico ( Sharp 1887); lectotype designation, redescription and distribution ( Smetana 1995: 109); distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus semunciarius Tottenham 1940 

Philonthus semunciarius Tottenham 1940: 20  . Type locality: Costa Rica?: El Congo, 750m. Type depository: BMNH? (Tottenham) HT, etc; FMNH 1View Materials PT. Distribution: Costa Rica  .

Philonthus sericans ( Gravenhorst 1802) 

Staphylinus sericans Gravenhorst 1802: 171  . Type locality: America septentr. Type depository: ZMHB LT ♂, #? PLT (under combined heading “ P. brunneus  ” and “ P. sericans  ”, see Smetana 1995: 320). Distribution: Canada, USA, Mexico, Brazil, Argentina. Notes: as synonym of P. brunneus ( Erichson 1840: 486)  ; as valid species, lectotype designation and redescription ( Smetana 1995: 319), distribution including Brazil (Chani-Posse 2004); Argentina in Scheerpeltz (1933: 1335, as P. brunneus  ); distribution, also within Mexico (Navarrete- Heredia et al. 2002); redescription, ( Chani-Posse 2010); distribution within Brazil (Asenjo et al. 2013).

= Staphylinus brunneus Gravenhorst 1802: 172  (preoccupied, junior primary homonym of Staphylinus brunneus Paykull 1789: 63  ). Type locality: America septentr. Type depository: ZMHB LT ♂, #? PLT (under combined heading “ P. brunneus  ” and “ P. sericans  ”, see Smetana 1995: 320). Note: as synonym of P. sericans ( Smetana 1995: 319)  .

= Staphylinus dimidiatus Say 1830: 37  , also 1834: 450. Type locality: Pennsylvania & Missouri. Type depository: ?? #?ST. Note: as synonym of P. ventralis ( Erichson 1840: 929)  ; as synonym of P. brunneus ( Bernhauer & Schubert 1914: 331)  ; as synonym of P. sericans ( Smetana 1995: 319)  .

= Philonthus picatus Kirby 1837: 93  (preoccupied, junior secondary homonym in Philonthus  of Quedius picatus Stephens 1832: 220  ). Type locality: Canada: "Lat. 54°". Type depository: BMNH? (Kirby) 2ST. Note: as synonym of P. brunneus ( Horn 1876: 151)  ; as synonym of P. sericans ( Smetana 1995: 319)  .

= Philonthus brevis Melsheimer 1844: 37  . Type locality: Pennsylvania. Type depository: MCZAbout MCZ LT, 2 PLT? (des. Smetana 1995: 321). Note: as synonym of P. brunneus ( Bernhauer & Schubert 1914: 331)  ; as synonym of P. sericans ( Smetana 1995: 319)  .

= Philonthus solitarius Sharp 1885: 424  . Type locality: Mexico: Jalapa. Distribution: Mexico. Type depository: BMNH HT ♀. Note: as synonym of P. sericans  ( Smetana 1995: 2, abstract, not in text!).

= Philonthus iaynesi Bernhauer 1934a: 117  . Type locality: Tucuman: Parque Aconquija. Distribution: Argentina, Brazil. Type depository: FMNHAbout FMNH 1 ST. Note: as synonym of P. sericans (Chani-Posse 2004: 221)  .

Philonthus seticornis Sharp 1885 

Philonthus seticornis Sharp 1885: 410  . Type locality: Guatemala, Cerro Zunil 4000–5000 ft  . Type depository: BMNH 11 ST: Cerro Zunil [ MCPAbout MCP seen]. Distribution: Guatemala  .

Philonthus silvaticus Cameron 1922 

Philonthus silvaticus Cameron 1922: 119  . Type locality: Jamaica  . Type depository: BMNH ( Cameron ) 9 ST. Distribution: Jamaica  .

Philonthus silvestris Bernhauer 1929 

Philonthus silvestris Bernhauer 1929: 194  . Type locality: Desierto de los Leones , Distrito Federal  . Type depository: FMNH 1View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus speculicollis Schubert 1909 

Philonthus speculicollis Schubert 1909: 292  . Type locality: Jalapa region  , Veracruz, Mexico  . Type depository: ZMHB ( Schubert ) 1 ST ♂ [ MCPAbout MCP seen]. Distribution: Mexico  . Note: distribution within Mexico (Navarrete- Heredia et al. 2002).

Philonthus speculipennis Bernhauer 1921 

Philonthus speculipennis Bernhauer 1921b: 75  . Type locality: West-Kolumbien: Umgebung von Cali am Cauca ( Canon del Mte Tolima, 1700m)  . Type depository: FMNHAbout FMNH HT. Distribution : Colombia. Note  : type material not seen (on loan); after original description probably not Philonthus  .

Philonthus stenocephalus Scheerpeltz 1972 

Philonthus stenocephalus Scheerpeltz 1972: 18  (new name for Philonthus angusticeps Bernhauer 1934a: 118  ). Distribution: Argentina, Bolivia; Chile?. Notes: redescription, distribution ( Bolivia in text, not map or specimen data) ( Chani-Posse 2010: 57).

= Philonthus angusticeps Bernhauer 1934a: 118  (preoccupied, junior primary homonym of Philonthus angusticeps Bernhauer 1933: 57  ). Type locality: Tucuman prov. Type depository: FMNHAbout FMNH LT ♂, 1 PLT. Distribution: Argentina. Notes: lectotype designation (as stenocephalus Scheerpeltz  : Chani-Posse 2010: 57; as angusticeps Bernhauer  : Chani-Posse 2010: 68).

Philonthus stolzmanni Bernhauer 1916 

Philonthus stolzmanni Bernhauer 1916b: 187  , attributed to Fauvel in litteris. Type locality: Bolivien  . Type depository: FMNHAbout FMNH HT. Distribution : Bolivia  .

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus stygialis Sharp 1885 

Philonthus stygialis Sharp 1885: 405  . Type locality: Guatemala, Balheu in Vera Paz; Panama, Peña Blanca 3000– 4000 ft. Type depository: BMNH 4 ST: 2 Peña Blanca, 2 Balheu [ MCPAbout MCP seen]. Distribution: Guatemala, Panama.

Philonthus subpunctipennis ( Coiffait & Sáiz 1968) 

Endeius subpunctipennis Coiffait & Sáiz 1968: 358  . Type locality: Chili. Type depository: Sáiz (in MNNCAbout MNNC) HT ♂  ; MNNC 2View Materials PT. Distribution : Chile. Notes: redescription (Chani-Posse de Maus 2008: 395); comb. nov. ex Endeius (Chani-Posse 2013: 404)  . 

Philonthus succinctus Guérin-Méneville 1844 

Philonthus succinctus Guérin-Méneville 1844: 13  . Type locality: Nouvelle-Grenade: régions tempérées et froides. Type depository: ?? #?ST. Distribution: Colombia, Peru. Notes: Peru ( Asenjo 2004); it probably belongs to Leptopeltus  .

Philonthus suspectus Erichson 1840 

Philonthus suspectus Erichson 1840: 465  . Type locality: prope Caracas Columbiae [= Venezuela]. Type depository: ZMHB 4 ST: 2 ♂, 2 ♀ [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala, Colombia, Venezuela, Brazil, Uruguay. Notes: Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution including Colombia, also within Brazil (Asenjo et al. 2013).

= Philonthus quadraticollis Sharp 1885: 409  . Type locality: Mexico: Cordova, Jalapa  ; Guatemala: San Gerónimo ; South America, Rio de Janeiro. Type depository: BMNH 5 + ST. Note: as synonym of P. suspectus ( Bernhauer & Schubert 1914: 357)  . 

Comment. Not Philonthus  , to be placed in a further study.

Philonthus tachiniformis ( Say 1830) 

Philonthus tachiniformis Say 1830: 37  , also 1834: 450. Type locality: Mexico  . Type depository: ?? #? ST. Distribution : Mexico  . Notes: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus tenuimarginatus Scheerpeltz 1960 

Philonthus tenuimarginatus Scheerpeltz 1960: 112  . Type locality: Cochabamba ( Bolivien), in der nächsten Umgebung seines Wohnsitzes, 2600m  . Type depository: NMWAbout NMW ( Scheerpeltz ) HT ♂. Distribution: Bolivia  .

Philonthus testaceipennis Erichson 1840 

Philonthus testaceipennis Erichson 1840: 435  . Type locality: Mexico. Type depository: ZMHB #?ST. Distribution: Mexico. Notes: distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2004, 2006); key ( Navarrete-Heredia 2003); redescription, distribution ( Márquez & Asiain 2010).

Philonthus titschacki Bernhauer 1941 

Philonthus titschacki Bernhauer 1941: 286  , also 1951: 276. Type locality: Hac. Huayuri, s Peru  . Type depository: FMNH 2View Materials ST. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Philonthus trapeziceps Scheerpeltz 1960 

Philonthus trapeziceps Scheerpeltz 1960: 107  . Type locality: Yungas de Palmar , 1250m. Type depository: NMWAbout NMW (Scheerpeltz) HT ♂. Distribution: Bolivia  .

Philonthus trepidus Erichson 1840 

Philonthus trepidus Erichson 1840: 489  . Type locality: ins. St. Johannis & Puerto Rico. Type depository: ZMHB LT ♂, 5 PLT. Distribution: Puerto Rico, Virgin Is., Antigua. Notes: lectotype designation, redescription, distribution ( Smetana 1994: 346); as valid species, ex synonym of P. havaniensis ( Smetana 1994)  ; distribution, also within Lesser Antilles ( Peck 2011a: Northern Leeward Is.),

Philonthus triangulum Horn 1884 

Philonthus triangulum Horn 1884: 202  . Type locality: [no specific locality; not orig. fixed (Washington Terr., Nevada, California)]. Type depository: MCZAbout MCZ (Horn) LT ♂, 8 PLT; (LeConte) 8 PLT; USNMAbout USNM 3 PLT. Distribution: Canada, USA, Mexico. Notes: lectotype designation, redescription, distribution in Mexico ( Smetana 1995: 338); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus trinitatis Blackwelder 1943 

Philonthus trinitatis Blackwelder 1943: 418  . Type locality: Trinidad: St. Augustine   . Type depository: MCZAbout MCZ HT. Distribution : Trinidad  .

Philonthus tristipennis Bernhauer 1941 

Philonthus tristipennis Bernhauer 1941: 289  , also 1951: 278. Type locality: Querobamba , 3520m, etc., s Peru  . Type depository: FMNH 2View Materials ST. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Philonthus trochilus Solsky 1872 

Philonthus trochilus Solsky 1872: 299  . Type locality: Venezuela [Amérique du Sud]  . Type depository: ZINAbout ZIN? (Solsky) #? ST. Distribution : Venezuela  . Note: type material not seen; generic affiliation unclear after original description, probably not Philonthus  .

Philonthus umbratus Sharp 1885 

Philonthus umbratus Sharp 1885: 424  . Type locality: Mexico: Las Vigas; Guatemala, Paso Antonio; Panama, Bugaba, Volcan de Chiriqui 4000–6000 ft, Peña Blanca 3000–4000 ft. Type depository: BMNH 4+ ST: from all type localities [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala, Panama. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus vagepunctatus Schubert 1909 

Philonthus vagepunctatus Schubert 1909: 292  . Type locality: Jalapa region  , Veracruz, Mexico  . Type depository: ZMHB ( Schubert ) 1 ST ♂ [bad condition, MCPAbout MCP seen]. Distribution: Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Philonthus varians ( Paykull 1789)  [adventive in CASA]

Staphylinus varians Paykull 1789: 45  . Type locality: [no specific locality]. Type depository: NHRSAbout NHRS 3 ST (1 labeled as LT by Drugmand in 1995 but evidently not published). Distribution: Europe, Russia (east to west), Armenia, Georgia, Turkey to Syria, Iran, Kazakhstan, Kyrgyzstan, Uzbekistan, Mongolia, north Africa, Afrotropical, South Africa, Madagascar, India, Myanmar, Tahiti; adventive in Canada, USA, Cuba, St. John, St. Vincent, Venezuela, Argentina, Chile. Notes: Argentina in Scheerpeltz (1972: 19); redescription, distribution ( Smetana 1995: 235; Chani-Posse 2010: 62, including Chile); distribution, also within Cuba ( Peck 2005), Lesser Antilles ( St. Vincent) ( Peck 2010).

Philonthus ventralis ( Gravenhorst 1802)  [adventive in CASA]

Staphylinus ventralis Gravenhorst 1802: 174  . Type locality: Lusitania. Type depository: ZMHB LT. Distribution: Palearctic, Iran; adventive in Canada, USA, Mexico, Costa Rica, French Guiana, Cuba, Jamaica, Hispaniola, Puerto Rico, Virgin Is., Lesser Antilles, Galapagos Is., Australia, New Zealand, New Caledonia, Afrotropical, etc. Notes: lectotype designation and redescription, " Mexico " only ( Smetana 1995: 191); New Zealand (Hammond ms); distribution ( Frank 1983); distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution, also within Galápagos Is. ( Newton & Peck 2006), Cuba ( Peck 2005), Dominica (Peck 2006), Lesser Antilles ( Peck 2010: St. Vincent, Peck 2011a: Northern Leeward Is.; Peck 1916); distribution within Cayman Is. (Frank et al. 2011); redescription, distribution ( Hromádka 2012b).

Philonthus viridiniger Bernhauer 1916 

Philonthus viridiniger Bernhauer 1916a: 30  . Type locality: West-Columbien: Umgebung von Cali im Caucatal  . Type depository: FMNHAbout FMNH HT ♀?. Distribution : Colombia  . Note: type material not seen (on loan); generic affiliation unclear after original description, probably within the Neotropical lineage.

Philonthus waterhousei Cameron 1922 

Philonthus waterhousei Cameron 1922: 120  , attributed to Fauvel in litteris. Type locality: St. Vincent & Grenada. Type depository: BMNH 10 ST: 8 St. Vincent, 2 Grenada [ MCPAbout MCP seen]. Distribution: Grenada, St. Vincent.

Comment. Not Philonthus  , to be placed in a further study.

Philonthus whymperi Sharp 1891 

Philonthus whymperi Sharp 1891: 40  . Type locality: Hacienda of Guachala (9217 ft), Antisanilla to Piñantura (11000 ft), Machachi (9–10000 ft). Type depository: BMNH 5 ST: 1 Guachala , 2 Antisanilla , 2 Machachi [ MCPAbout MCP seen]. Distribution: Ecuador  .

Comment. Not Philonthus  , to be placed in a further study within the Neotropical lineage.

Philonthus yaqui Navarrete-Heredia 2003 

Philonthus yaqui Navarrete-Heredia 2003: 2  . Type locality: Yécora , 1400m, Sonora, Mexico  . Type depository: CZUGAbout CZUG HT ♂, 20 PT, FMNHAbout FMNH etc. 1+PT. Distribution : Mexico. Notes: redescription, distribution ( Márquez & Asiain 2010). 

MCP

Pontificia Universidade Catolica do Rio Grande do Sul

NMW

Naturhistorisches Museum, Wien

ZIN

Russian Academy of Sciences, Zoological Institute, Zoological Museum

MCZ

Museum of Comparative Zoology

USNM

Smithsonian Institution, National Museum of Natural History

FMNH

Field Museum of Natural History

IMLA

Fundacion e Instituto Miguel Lillo

MACN

Museo Argentino de Ciencias Naturales Bernardino Rivadavia

MCSN

Museo Civico di Storia Naturale "Giacomo Doria"

MRSN

Museo Regionale di Scienze Naturali

IRSNB

Institut Royal des Sciences Naturelles de Belgique

MNNC

Museo Nacional de Historia Natural, Santiago

AMNH

American Museum of Natural History

MNHN

Museum National d'Histoire Naturelle

CNC

Canadian National Collection of Insects, Arachnids, and Nematodes

CNIN

Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico

NHRS

Swedish Museum of Natural History, Entomology Collections

MZFC

Museo de Zoologia "Alfonso L. Herrera"

CZUG

Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Loc

Philonthus Stephens 1829

Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey 2018

2018
Loc

Philonthus

Stephens 1829a : 23 Stephens (1829b : 279 Curtis (1829: 25) Dillwyn (1829: 68)

Loc

Spatulonthus

Tottenham 1955 : 154 Coiffait & Sáiz (1968: 360) Smetana (1995: 51)

Loc

Endeius Coiffait & Sáiz 1968 : 355

Coiffait & Sáiz 1968 : 355

Loc

Philonthus abdominalis

Erichson 1847 : 89

Loc

Philonthus accedens

Sharp 1885 : 403

Loc

Philonthus acromyrmecis

Scheerpeltz 1976 : 135

Loc

Philonthus actinus

Bernhauer 1917c : 103

Loc

Philonthus actocharis

Sharp 1887 : 788

Loc

Philonthus aenescens

Solsky 1870 : 262

Loc

Philonthus aeruginosus

Nordmann 1837 : 95 Chani-Posse 2010: 12

Loc

Philonthus aliiseriatus

Bruch 1915 : 501

Loc

Philonthus alius

Bernhauer & Schubert 1914 : 328 Sharp 1885 : 396

Loc

Philonthus crassulus

Sharp 1885 : 396 Horn 1884 : 213

Loc

Philonthus alticola

Sharp 1885 : 400

Loc

Philonthus alumnus

Erichson 1840 : 490 Smetana 1994: 342

Loc

Philonthus alutaceus

Horn 1884 : 183 Smetana 1995: 106

Loc

Philonthus ambiguus

Erichson 1840 : 441

Loc

Philonthus apheles

Solsky 1868 : 132

Loc

Philonthus apicetestaceus

Scheerpeltz 1960 : 106

Loc

Philonthus argus

Herman 2001a : 46 Bernhauer 1934a : 118 Chani-Posse 2010: 13

Loc

Philonthus tucumanensis

Bernhauer 1934a : 118 Bernhauer 1927 : 245 Chani-Posse 2010: 14

Loc

Philonthus arizonensis

Horn 1884 : 203 Smetana 1995: 374

Loc

Philonthus atramentarius

Erichson 1840 : 440

Loc

Philonthus audanti

Blackwelder 1943 : 406

Loc

Philonthus auripennis

Bernhauer 1916a : 37

Loc

Philonthus avicola

Bernhauer 1942 : 25

Loc

Philonthus belonuchiformis

Bruch 1915 : 501 Bernhauer (1915: 145)

Loc

Philonthus bolivianus

Bernhauer 1916a : 35

Loc

Philonthus bonariensis

Bernhauer 1909 : 245 Maus 2009 : 38 Chani-Posse 2010: 15

Loc

Philonthus catamarcanus

Bernhauer 1916a : 36 Maus 2009 : 38 Chani-Posse 2010: 19

Loc

Philonthus weiseri

Bernhauer 1921c : 169 Maus 2009 : 38

Loc

Philonthus brachypterus

Solsky 1872 : 297

Loc

Philonthus brasilianus

Bernhauer 1916a : 40

Loc

Philonthus bruchianus

Chani-Posse 2010 : 20

Loc

Philonthus caliensis

Bernhauer 1916a : 33

Loc

Philonthus callosipennis

Scheerpeltz 1960 : 100

Loc

Philonthus caribaeus

Bierig 1940 : 141

Loc

Philonthus carinulatus

Sharp 1885 : 402

Loc

Philonthus catamarcensis

Bruch 1915 : 501 Bachmann et al. (2017 : 44

Loc

Philonthus cavifrons

Sharp 1885 : 408

Loc

Philonthus centralis

Sharp 1885 : 422

Loc

Philonthus chiriquensis

Sharp 1885 : 404

Loc

Philonthus chlorocephalus

Solsky 1872 : 301

Loc

Philonthus (Philonthus) columbianus Baudi di Selve 1848 : 126

Selve 1848 : 126 Herman 2001c : 1594

Loc

Philonthus corallipennis

Sharp 1876a : 166

Loc

Philonthus cordilleranus

Bernhauer 1916a : 34

Loc

Philonthus cribrellus

Fauvel 1891 : 112

Loc

Philonthus cribriceps

Bernhauer 1921c : 105

Loc

Philonthus cribripennis

Sharp 1885 : 403

Loc

Philonthus cribriventris

Bernhauer 1912a : 43 Maus 2009 : 39 Chani-Posse 2010: 25

Loc

Philonthus catamarcanus var. densior

Bernhauer 1916a : 36 Maus 2009 : 39

Loc

Philonthus crinitus

Schubert 1909 : 293

Loc

Staphylinus cruentatus

Gmelin 1790 : 2035 Smetana (1995: 233) Smetana 1995: 232

Loc

Philonthus cyaneoaureus

Bernhauer 1935 : 90

Loc

Philonthus deplanatus

Sharp 1885 : 423

Loc

Philonthus diamantinus

Bernhauer 1917c : 102

Loc

Philonthus discedens

Schubert 1909 : 291

Loc

Staphylinus discoideus

Gravenhorst 1802 : 38 Smetana 1995: 186

Loc

Philonthus emelinae Coiffait & Sáiz 1968 : 354

Coiffait & Sáiz 1968 : 354 Maus 2009 : 39

Loc

Philonthus discretus

Sharp 1876b : 428

Loc

Philonthus dispersus

Fauvel 1891 : 111

Loc

Philonthus duploseriatus

Solsky 1868 : 136

Loc

Philonthus eryx

Smetana 1995 : 306

Loc

Philonthus excellens

Bernhauer 1916a : 37

Loc

Philonthus exemplaris

Wendeler 1956 : 231

Loc

Philonthus fassli

Bernhauer 1916a : 29

Loc

Philonthus fenyesi

Bernhauer 1910 : 381

Loc

Philonthus feralis

Erichson 1840 : 469 Smetana 1994: 346 Posse 2010 : 65

Loc

Philonthus ferreipennis

Horn 1884 : 202 Smetana 1995: 348

Loc

Philonthus ferreipennis stigmatias

Smetana 1995 : 350 Smetana 1995: 349

Loc

Philonthus fibularius

Erichson 1840 : 442

Loc

Philonthus boucardi

Solsky 1868 : 127 Sharp 1885: 404 Bernhauer & Schubert 1914: 338

Loc

Philonthus figulus

Erichson 1840 : 464 Chani-Posse 2010: 28

Loc

Philonthus amazonicus

Sharp 1876a : 165 Sharp 1885: 415

Loc

Philonthus fissilis

Sharp 1885 : 399

Loc

Philonthus flavolimbatus

Erichson 1840 : 471 Smetana 1995 : 201 Smetana 1995 : 200 Chani-Posse 2010 : 29

Loc

Philonthus apicipennis Lynch-Arribálzaga 1884: 155

Chani-Posse 2010: 30 Fauvel 1887: 232

Loc

Philonthus wacoensis

Casey 1915 : 430 Smetana 1995: 200

Loc

Philonthus flohri

Sharp 1876b : 429 Scheerpeltz 1971b: 205

Loc

Philonthus floresi

Chani-Posse 2010 : 31

Loc

Philonthus formicarius

Bernhauer 1908 : 334

Loc

Philonthus forsteri

Scheerpeltz 1960 : 104

Loc

Endeius franzi Sáiz 1971 : 345

Sáiz 1971 : 345 Maus 2008 : 398

Loc

Philonthus fulgipennides

Sharp 1885 : 410

Loc

Philonthus fulgipennis

Sharp 1885 : 410

Loc

Philonthus furvus

Nordmann 1837 : 88 Smetana 1995: 92

Loc

Philonthus californicus

Mannerheim 1843 : 230 Smetana 1995: 92

Loc

Philonthus comptus

Haldeman 1852 : 375 Bernhauer & Schubert 1914: 339

Loc

Philonthus gaudichaudii

Blanchard 1842 : 81

Loc

Philonthus gentilis

Horn 1884 : 185 Smetana 1995: 160

Loc

Philonthus gracillimus

Sharp 1876a : 168

Loc

Philonthus grandicollis

Horn 1884 : 193 Smetana 1995: 300

Loc

Philonthus gratiosus

Erichson 1840 : 461

Loc

Philonthus griseolus

Sharp 1885 : 399

Loc

Philonthus guatemalenus

Sharp 1885 : 403 Bernhauer & Schubert 1914: 340

Loc

Philonthus guerreroensis

Bernhauer 1910 : 380

Loc

Philonthus hades

Chatzimanolis & Engel 2013 : 288

Loc

Staphylinus havaniensis Laporte de Castelnau 1835 : 116

Castelnau 1835 : 116

Loc

Philonthus obscurus

Fauvel 1863 : 434 Gravenhorst 1802 :174

Loc

Philonthus hepaticus

Erichson 1840 : 451 Smetana 1995: 176 Smetana 1995 : 177 Chani-Posse 2010: 32

Loc

Philonthus vilis

Erichson 1840 : 451 Blackwelder 1943: 401 Smetana 1995: 176

Loc

Philonthus nanus

Melsheimer 1844 : 36 Gemminger & Harold 1868: 589 Smetana 1995: 177

Loc

Philonthus nanus var. palleolus

Melsheimer 1844 : 37 Gemminger & Harold 1868: 589

Loc

Philonthus cinctutus

Melsheimer 1844 : 37 Horn 1884: 223 Fauvel 1891: 112 Smetana 1995: 177

Loc

Philonthus varicolor

Boheman 1858 : 29 Fauvel 1891: 112

Loc

Philonthus parvimanus

Sharp1885 : 406 Fauvel 1891: 112 Smetana 1995: 174

Loc

Philonthus oklahomensis

Hatch & Ortenburger 1930 : 12 Smetana 1995: 177

Loc

Philonthus heteropus

Sharp 1887 : 788

Loc

Philonthus hirtiventris

Sharp 1885 : 410

Loc

Philonthus hoegei

Sharp 1885 : 400

Loc

Philonthus idoneus

Sharp 1885 : 411

Loc

Philonthus imperialis

Bernhauer 1916a : 28

Loc

Philonthus incertus

Solsky 1868 : 135 Smetana 1995: 221

Loc

Philonthus gracilicornis

Horn 1884 : 190 Sharp 1887: 788 Smetana 1995: 222

Loc

Philonthus indigaceus

Fauvel 1891 : 110

Loc

Philonthus iopterus

Erichson 1840 : 928

Loc

Philonthus iris

Sharp 1885 : 401

Loc

Philonthus jalapensis

Bernhauer 1910 : 382

Loc

Philonthus jenseni

Bernhauer 1912a : 42 Chani-Posse 2010: 35

Loc

Philonthus klimschi

Bernhauer 1916a : 38

Loc

Philonthus labilis

Erichson 1840 : 500

Loc

Philonthus laetipennis

Sharp 1885 : 409

Loc

Philonthus laeviventris

Schubert 1909 : 291

Loc

Philonthus lecontei

Horn 1884 : 222 Smetana 1995: 467 Casey 1915: 442

Loc

Philonthus limaensis

Solsky 1872 : 300

Loc

Philonthus lineatoventris

Bruch 1915 : 502

Loc

Endeius loensis Coiffait & Sáiz 1968 : 357

Coiffait & Sáiz 1968 : 357 Maus 2008 : 390 Maus 2008 : 388

Loc

Philonthus lomatus

Erichson 1840 : 482 Smetana 1995: 390

Loc

Philonthus georgianus

Sachse 1852 : 143 Fauvel 1869: 488

Loc

Philonthus longicornis

Stephens 1832 : 237 Smetana 1995 : 244 Chani-Posse 2010 : 35

Loc

Philonthus perplexus

Fairmaire & Germain 1862 : 431 Maus 2009 : 37 Coiffait & Sáiz 1968: 360

Loc

Philonthus fumosus

Solsky 1868 : 134 Sharp 1885: 416

Loc

Philonthus lucidanus

Blackwelder 1944 : 134 Fauvel 1891 : 116

Loc

Philonthus lucidus

Fauvel 1891 : 116 Gravenhorst 1802 : 21

Loc

Endeius lugubris Sáiz 1971 : 347

Sáiz 1971 : 347 Maus 2008 : 399

Loc

Philonthus lynchi

Bernhauer & Schubert 1914 : 345 Chani-Posse 2010: 38

Loc

Philonthus affinis Lynch-Arribálzaga 1884: 160

Roth 1851 : 118

Loc

Philonthus macilentus

Sharp 1885 : 423

Loc

Philonthus melampus

Smetana 1991 : 236

Loc

Philonthus melancholicus

Sharp 1885 : 425

Loc

Philonthus melanopus

Sharp 1885 : 405

Loc

Philonthus metallifer

Bernhauer 1916a : 31

Loc

Philonthus mnemon

Smetana 1995 : 306

Loc

Philonthus mollis

Sharp 1885 : 408

Loc

Philonthus monachus

Bernhauer 1908 : 335

Loc

Philonthus morosus

Casey 1915 : 441

Loc

Philonthus navarretei Márquez & Asiain 2014 : 297

Márquez & Asiain 2014 : 297 Márquez & Asiain 2010 : 280

Loc

Philonthus navarretei Márquez & Asiain 2010 : 280

Márquez & Asiain 2010 : 280 Navarrete-Heredia & Newton (2013: 132)

Loc

Philonthus neptunus

Smetana 1995 : 436

Loc

Philonthus nigerrimus

Erichson 1840 : 434 Smetana 1991: 235

Loc

Philonthus occultus

Sharp 1885 : 407

Loc

Philonthus ochromerus

Sharp 1885 : 406

Loc

Philonthus oenotrus

Smetana 1991 : 228

Loc

Philonthus (Gabrius) opacigaster

Bierig 1940 : 143

Loc

Philonthus ophthalmicus

Bernhauer 1941 : 288

Loc

Philonthus orphanus

Erichson 1840 : 452 Erichson 1840 : 499 Smetana 1992: 135

Loc

Philonthus palcoensis

Bernhauer 1941 : 288

Loc

Philonthus pallipes

Blanchard 1842 : 82 Chani-Posse 2010 : 42 Chani-Posse 2010: 40

Loc

Philonthus depressus

Steinheil 1869 : 253 Bernhauer & Schubert 1914: 349

Loc

Philonthus pallidipes

Bernhauer & Schubert 1914: 349

Loc

Philonthus palmatus

Sharp 1885 : 397

Loc

Philonthus pauper

Bernhauer 1916a : 32

Loc

Philonthus pauxillus

Solsky 1868 : 133 Smetana 1995 : 179 Chani-Posse 2010 : 43

Loc

Philonthus pavidus

Erichson 1847 : 89

Loc

Philonthus peratriceps

Scheerpeltz 1933 : 1355 Sharp 1885 : 404

Loc

Philonthus atriceps

Sharp 1885 : 404 Horn 1884 :199

Loc

Philonthus piceatus

Nordmann 1837 : 85

Loc

Philonthus iridiventris

Solsky 1868 : 128 Sharp 1885: 400

Loc

Philonthus mexicanus

Sharp 1876b : 430 Sharp 1885: 400

Loc

Staphylinus politus

Linnaeus 1758 : 422 Smetana 1995: 73 Chani-Posse 2010: 44

Loc

Philonthus pollens Márquez & Asiain 2010 : 272

Márquez & Asiain 2010 : 272

Loc

Philonthus prescottianus

Sharp 1887 : 787

Loc

Philonthus prismalis

Fauvel 1891 : 114

Loc

Philonthus prolixicornis

Sharp 1885 : 424

Loc

Philonthus proportionalis

Schubert 1909 : 293

Loc

Philonthus pulcher

Bernhauer 1908 : 335

Loc

Philonthus pulchricolor

Bernhauer 1923 : 63

Loc

Philonthus quadricolor

Bernhauer 1916a : 30 Fauvel 1879 : 99

Loc

Philonthus punctifer

Nordmann 1837 : 107

Loc

Philonthus punctipennis (

Solier 1849 : 319

Loc

Philonthus pyropterus

Kraatz 1859b : 12 Solier 1849 : 317 Fauvel 1878: 546 Smetana (1995: 179) Coiffait & Sáiz (1968: 355) Chani-Posse 2010: 15

Loc

Staphylinus rufipennis

Solier 1849 : 317 Fabricius 1801 : 597 Gravenhorst 1802 : 40

Loc

Philonthus quadraticeps

Boheman 1858 : 28 Maus 2009 : 35

Loc

Philonthus tenebrosus

Boheman 1858 : 28 Maus 2009 : 35

Loc

Philonthus quadrulus

Horn 1884 : 221 Smetana 1995: 314

Loc

Philonthus sagax

Casey 1915 : 445 Smetana 1995: 314

Loc

Philonthus rectangulus

Sharp 1874 : 42 Smetana 1995 : 216 Chani-Posse 2010 : 47

Loc

Philonthus rectilaterus

Sharp 1885 : 425 Scheerpeltz 1971b: 205

Loc

Philonthus rhadamanthus

Chatzimanolis & Engel 2013 : 289

Loc

Philonthus richterianus

Bruch 1915 : 503

Loc

Philonthus ridiculus

Blackwelder 1944 : 134 Bernhauer 1916a : 33

Loc

Philonthus grandiceps

Bernhauer 1916a : 33 Kraatz 1859a : 95 Poppius 1909 : 24

Loc

Philonthus riedeli

Bernhauer 1916a : 39

Loc

Philonthus rubromaculatus

Blanchard 1842 : 82 Chani-Posse 2010: 50

Loc

Philonthus ruficauda

Bernhauer 1934a : 119

Loc

Philonthus rufoplagiatus

Solsky 1868 : 130 Sharp 1885: 400

Loc

Philonthus rufotibialis Márquez & Asiain 2010 : 270

Márquez & Asiain 2010 : 270

Loc

Philonthus alumnus var. rufulus

Horn 1884 : 199 Smetana 1995: 430

Loc

Philonthus alumnus

Horn 1884 : 198 Smetana 1994: 341 Smetana 1995: 434

Loc

Philonthus alumnus var. atriceps

Horn 1884 : 199 Smetana 1995: 430

Loc

Philonthus sallaei

Sharp 1885 : 398

Loc

Philonthus scintillatus

Blackwelder 1944 : 134 Solsky 1868 : 131

Loc

Philonthus scintillans

Solsky 1868 : 131 Gravenhorst 1806 : 70

Loc

Philonthus semicupreus

Fauvel 1891 : 114

Loc

Philonthus semicyaneus

Schubert 1909 : 290

Loc

Philonthus semiruber

Horn 1884 : 187 Smetana 1995: 109

Loc

Philonthus semunciarius

Tottenham 1940 : 20

Loc

Staphylinus sericans

Gravenhorst 1802 : 171 Smetana 1995 : 320 Erichson 1840: 486 Smetana 1995: 319 Scheerpeltz (1933 : 1335

Loc

Staphylinus brunneus

Gravenhorst 1802 : 172 Paykull 1789 : 63 Smetana 1995 : 320 Smetana 1995: 319

Loc

Staphylinus dimidiatus

Say 1830 : 37 Erichson 1840: 929 Bernhauer & Schubert 1914: 331 Smetana 1995: 319

Loc

Philonthus picatus

Kirby 1837 : 93 Stephens 1832 : 220 Horn 1876: 151 Smetana 1995: 319

Loc

Philonthus brevis

Melsheimer 1844 : 37 Smetana 1995 : 321 Bernhauer & Schubert 1914: 331 Smetana 1995: 319

Loc

Philonthus solitarius

Sharp 1885 : 424

Loc

Philonthus iaynesi

Bernhauer 1934a : 117

Loc

Philonthus seticornis

Sharp 1885 : 410

Loc

Philonthus silvaticus

Cameron 1922 : 119

Loc

Philonthus silvestris

Bernhauer 1929 : 194

Loc

Philonthus speculicollis

Schubert 1909 : 292

Loc

Philonthus speculipennis

Bernhauer 1921b : 75

Loc

Philonthus stenocephalus

Scheerpeltz 1972 : 18 Bernhauer 1934a : 118 Chani-Posse 2010: 57

Loc

Philonthus angusticeps

Bernhauer 1934a : 118 Chani-Posse 2010 : 57 Chani-Posse 2010 : 68

Loc

Philonthus stolzmanni

Bernhauer 1916b : 187

Loc

Philonthus stygialis

Sharp 1885 : 405

Loc

Endeius subpunctipennis Coiffait & Sáiz 1968 : 358

Coiffait & Sáiz 1968 : 358 Maus 2008 : 395

Loc

Philonthus succinctus Guérin-Méneville 1844 : 13

Guérin-Méneville 1844 : 13

Loc

Philonthus suspectus

Erichson 1840 : 465

Loc

Philonthus quadraticollis

Sharp 1885 : 409 Bernhauer & Schubert 1914: 357

Loc

Philonthus tachiniformis

Say 1830 : 37

Loc

Philonthus tenuimarginatus

Scheerpeltz 1960 : 112

Loc

Philonthus testaceipennis

Erichson 1840 : 435

Loc

Philonthus titschacki

Bernhauer 1941 : 286

Loc

Philonthus trapeziceps

Scheerpeltz 1960 : 107

Loc

Philonthus trepidus

Erichson 1840 : 489 Smetana 1994: 346

Loc

Philonthus triangulum

Horn 1884 : 202 Smetana 1995: 338

Loc

Philonthus trinitatis

Blackwelder 1943 : 418

Loc

Philonthus tristipennis

Bernhauer 1941 : 289

Loc

Philonthus trochilus

Solsky 1872 : 299

Loc

Philonthus umbratus

Sharp 1885 : 424

Loc

Philonthus vagepunctatus

Schubert 1909 : 292

Loc

Staphylinus varians

Paykull 1789 : 45 Scheerpeltz (1972: 19) Smetana 1995 : 235 Chani-Posse 2010 : 62

Loc

Staphylinus ventralis

Gravenhorst 1802 : 174 Smetana 1995: 191

Loc

Philonthus viridiniger

Bernhauer 1916a : 30

Loc

Philonthus waterhousei

Cameron 1922 : 120

Loc

Philonthus whymperi

Sharp 1891 : 40

Loc

Philonthus yaqui

Navarrete-Heredia 2003 : 2