Paederomimus Sharp 1885

Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey, 2018, Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae), Zootaxa 4449 (1), pp. 1-95 : 51-57

Paederomimus Sharp 1885

Paederomimus Sharp 1885: 438 –439. Type species: Paederomimus difformiceps Sharp, 1885 , by original designation.

Notes: Paederomimus is the most diverse and widely distributed genus among the Neotropical endemic genera, accounting for 58 species at present. Sharp (1885) erected the genus for 39 species, some of which were organized into three species groups. Never revised after its original description, other species were later described and/or assigned to this genus (see references below) whose monophyly is currently doubtful (Chani- Posse et al. 2018). The only available hypothesis of monophyly known at present is that provided by Sharp (1885: 439) who stated “3. A group for which, if the genus be divided, the name Paederomimus be retained, and of which P. difformiceps may be taken as type ”. Within the context of this study we find adequate to orientate the readers through the study of this genus by doing remarks into those species that fulfill Sharp´s concept of his Paederomimus group 3.

Paederomimus abactus (Sharp 1876)

Philonthus abactus Sharp 1876a: 177 . Type locality: Ega. Type depository: BMNH HT. Distribution : Brazil (Amazonas) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Paederomimus aberrans (Sharp 1876)

Philonthus aberrans Sharp 1876a: 174 . Type locality: Ega . Type depository: BMNH 1 ST. Distribution : Brazil (Amazonas) , Peru. Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; Peru (Newton 2015); distribution within Brazil (Asenjo et al. 2013).

Paederomimus aeneiceps (Sharp 1876)

Philonthus aeneiceps Sharp 1876a: 169 . Type locality: Ega . Type depository: BMNH 2 ST. Distribution : Brazil (Amazonas) , Peru. Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; Peru ( Asenjo 2004), distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus aeneicollis (Sharp 1876)

Philonthus aeneicollis Sharp 1876a: 172 . Type locality: Ega & St. Paulo [Amazonas]. Type depository: BMNH 2 ST: one with abdomen only [ MCP seen]. Distribution: Brazil (Amazonas). Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 (Sharp: 439), to be placed in a further study.

Paederomimus angularius ( Erichson 1840)

Philonthus angularius Erichson 1840: 477 . Type locality: Mexico. Type depository: ZMHB 3 ST [ MCP seen]. Distribution: Mexico. Notes: comb. nov. ex Philonthus ( Sharp 1885: 443) ; original spelling " angularius ", mostly angularis used later ( Bernhauer & Schubert 1914, Herman 2001d: 2723, 2724, Jiménez-Sánchez et al. 2000); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus angusticollis ( Erichson 1840)

Philonthus angularius Erichson 1840: 487 . Type locality: Mexico. Type depository: ZMHB 1 ST [ MCP seen]. Distribution: Mexico, Guatemala. Notes: comb. nov. ex Philonthus ( Sharp 1885: 447) ; distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Paederomimus aterrimus Bernhauer 1917

Paederomimus aterrimus Bernhauer 1917a: 36 . Type locality: West-Columbien: Umgebung von Cali am Rio Cauca (Cañon del Mte Tolima, 1700m). Type depository: FMNH HT. Distribution: Colombia .

Paederomimus basipennis Sharp 1885

Paederomimus basipennis Sharp 1885: 450 . Type locality: Chiriqui, Panama. Type depository: BMNH HT. Distribution: Mexico, Panama. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Paederomimus capitalis (Sharp 1876)

Philonthus capitalis Sharp 1876a: 171 . Type locality: Ega . Type depository: BMNH HT. Distribution : Brazil (Amazonas) , Costa Rica?. Notes: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus capitatus Sharp 1885

Paederomimus capitatus Sharp 1885: 441 . Type locality: Mexico, Jalapa, Playa Vicente. Type depository: BMNH 4 ST: 2 Jalapa, 1 Playa Vicente [ MCP seen]. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus cephalax Sharp 1885

Paederomimus cephalax Sharp 1885: 446 . Type locality: Panama, David . Type depository: BMNH 20 ST. Distribution : Panama .

Paederomimus chontalensis Sharp 1885

Paederomimus chontalensis Sharp 1885: 440 . Type locality: Nicaragua, Chontales . Type depository: BMNH HT ♀. Distribution : Nicaragua ; Panama.

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus cognatus (Sharp 1876)

Philonthus cognatus Sharp 1876a: 169 . Type locality: Ega. Type depository: BMNH 2 ST. Distribution: Brazil (Amazonas). Notes: comb. nov. ex Philonthus ( Sharp 1885: 438) ; preoccupied, junior primary homonym of Philonthus cognatus Stephens 1832: 229 ; conserved, Opinion 2053 ( ICZN 2003); ICZN application to conserve name ( Herman 2001a: 46); distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimu s group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus confertus Sharp 1885

Paederomimus confertus Sharp 1885: 449 . Type locality: Panama, David, Volcan de Chiriqui about 4000 ft. Type depository: BMNH 9 ST: 7 David, 2 Chiriqui [ MCP seen]. Distribution: Costa Rica, Panama.

Paederomimus conformis (Sharp 1876)

Philonthus conformis Sharp 1876a: 175 . Type locality: "Amazons". Type depository: BMNH HT ♂. Distribution : Brazil (Amazonas) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimu s group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus contractus Sharp 1885

Paederomimus contractus Sharp 1885: 448 . Type locality: Guatemala, San Gerónimo, Cubilguitz, Cahabon, Chacoj; Nicaragua. Type depository: BMNH 29 ST: 25 San Gerónimo, 2 Chacoj, 1 Cahabon, 1 Cubilguitz [ MCP seen]; FMNH 2 ST. Distribution: Guatemala, Nicaragua, Costa Rica.

Paederomimus corrosus Sharp 1885

Paederomimus corrosus Sharp 1885: 451 . Type locality: Panama, Bugaba 800–1500 ft. Type depository: BMNH 11 ST [ MCP seen]. Distribution: Costa Rica, Panama.

Paederomimus cribratus Sharp 1885

Paederomimus cribratus Sharp 1885: 450 . Type locality: Panama, Volcan de Chiriqui 2000–3000 ft. Type depository: BMNH 4 ST [ MCP seen]. Distribution : Panama. Note: spelled “ cribricollis ” on Pl. 11, Fig. 24 (teste Sharp 1885: 450).

Paederomimus cribricollis ( Erichson 1840)

Philonthus cribricollis Erichson 1840: 487 . Type locality: Columbia & Mexico. Type depository: ZMHB 4 ST ♀ [ MCP seen]. Distribution: Mexico?, Colombia, Venezuela, Peru, Trinidad, Grenada. Notes: comb. nov. ex Philonthus ( Sharp 1885: 448) ; distribution, also within Mexico ( Navarrete-Heredia et al. 2002); Peru ( Asenjo 2004).

= Paederomimus trinidadensis Bernhauer 1915: 296 . Type locality: Caparo , Trinidad . Type depository: FMNH 2 ST. Distribution : Trinidad. Note: as synonym of P. cribricollis ( Blackwelder 1943: 396) .

Paederomimus cupreonitens Schubert 1909

Paederomimus cupreonitens Schubert 1909: 294 . Type locality: Jalapa region , Veracruz, Mexico. Type depository: ZMHB 1 ST [ MCP seen] ; FMNH 1 ST. Distribution : Mexico . Note: distribution within Mexico (Navarrete- Heredia et al. 2002).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus densiceps Bernhauer 1908

Paederomimus densiceps Bernhauer 1908: 332 . Type locality: Petropolis, Brazil . Type depository: FMNH 1 ST. Distribution : Brazil (Rio de Janeiro) . Note: distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus difformiceps Sharp 1885

Paederomimus difformiceps Sharp 1885: 445 . Type locality: Guatemala, near the city; Capetillo . Type depository: BMNH 28 ST: 25 Guatemala city, 3 Capetillo [ MCP seen] ; FMNH 2 ST. Distribution : Guatemala .

Paederomimus distans Sharp 1885

Paederomimus distans Sharp 1885: 450 . Type locality: Guatemala, Zapote . Type depository: BMNH HT. Distribution : Guatemala ; Panama. Note: Paederomimus group 3 after Sharp (1885: 439) but doubtful assignation (AKH pers. obs.), to be placed in a further study.

Paederomimus flavoguttatus Bernhauer 1908

Paederomimus flavoguttatus Bernhauer 1908: 332 . Type locality: Brasilien . Type depository: FMNH 1 ST. Distribution : Brazil (São Paulo, Santa Catarina) . Note: distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus gentilis Sharp 1885

Paederomimus gentilis Sharp 1885: 439 . Type locality: Mexico: Cerro de Plumas , Jalapa. Type depository: BMNH 3 ST ♀ . Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Paederomimus hilaris ( Fauvel 1874)

Philonthus hilaris Fauvel 1874: 246 (new name for Philonthus laetus Erichson 1840: 477 ). Note: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 368) . Distribution: Colombia, Brazil; Uruguay?, Paraguay?. Note: distribution within Brazil (Asenjo et al. 2013).

= Philonthus laetus Erichson 1840: 477 (preoccupied, junior primary homonym of Philonthus laetus Heer 1839: 259 ). Type locality: Columbia & Brasilia. Type depository: ZMHB 3 ST [ MCP seen]. Distribution: Colombia, Brazil. Notes: as Paederomimus , comb. nov. ex Philonthus ( Sharp 1885: 438) ; as synonym of P. hilaris ( Fauvel 1874: 246) .

Paederomimus imitator Bierig 1937

Paederomimus imitator Bierig 1937: 198 . Type locality: Panamá: Volcán de Chiriqui, 1500m . Type depository: FMNH HT. Distribution : Panama .

Paederomimus insolitus Sharp 1885

Paederomimus insolitus Sharp 1885: 447 . Type locality: Panama, Bugaba 800–1500 ft . Type depository: BMNH 6 ST. Distribution : Panama .

Paederomimus interjectus Bernhauer 1908

Paederomimus interjectus Bernhauer 1908: 333 . Type locality: St. Vincent, W. I . Type depository: FMNH 2 ST. Distribution : Lesser Antilles ( St. Vincent) . Note: distribution within Lesser Antilles (St. Vincent) ( Peck 2010).

= Paederomimus smithi Cameron 1922: 121 , attributed to Fauvel in litteris. Type locality: St. Vincent . Type depository: BMNH #? ST. Distribution : Lesser Antilles ( St. Vincent). Note: as synonym of P. interjectus ( Blackwelder 1943: 397) .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus klimschi Bernhauer 1917

Paederomimus klimschi Bernhauer 1917a: 36 . Type locality: S. Catharina, Brasilien . Type depository: FMNH 2 ST. Distribution : Brazil (Santa Catarina) . Note: distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus longipes (Sharp 1876)

Philonthus longipes Sharp 1876a: 178 . Type locality: Obydos. Type depository: BMNH 1 ST. Distribution: Brazil ( Amazonas , Pará) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus lustralis ( Erichson 1840)

Philonthus lustralis Erichson 1840: 489 . Type locality: ins. Puerto Rico. Type depository: ZMHB 3 ST [ MCP seen]. Distribution : Puerto Rico, St. Thomas ; Trinidad. Note: comb. nov. ex Philonthus ( Sharp 1885: 438) .

= Paederomimus insularis Bernhauer 1908: 333 . Type locality: St. Thomas. Type depository: FMNH 1ST. Distribution: Lesser Antilles (St. Thomas). Note: as synonym of P. lustralis ( Blackwelder, 1943: 398) .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus lustrator (Sharp 1876)

Philonthus lustrator Sharp 1876a: 172 . Type locality: Pará . Type depository: BMNH HT. Distribution: Brazil ( Amazonas , Pará) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus maculipes Sharp 1885

Paederomimus maculipes Sharp 1885: 445 . Type locality: Mexico : Cordova, Jalapa. Type depository: BMNH 3 ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Paederomimus mandibularis Sharp 1885

Paederomimus mandibularis Sharp 1885: 439 . Type locality: Guatemala, Capetillo . Type depository: BMNH HT ♂. Distribution : Guatemala .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus mexicanus Sharp 1885

Paederomimus mexicanus Sharp 1885: 444 . Type locality: Mexico: Xalapa , Veracruz . Type depository: BMNH HT ♂. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Comment. Paederomimus group 3 after Sharp (1885: 439) but doubtful assignation (AKH pers. obs.), to be placed in a further study.

Paederomimus mimeticus ( Sharp 1885)

Belonuchus mimeticus Sharp 1885: 436 . Type locality: Panama, Bugaba 800–1500 ft. Type depository: BMNH 6 ST: 3 ♂, 3 ♀ [ MCP seen]. Distribution: Costa Rica, Panama. Note: comb. nov. ex Belonuchus ( Chani-Posse et al. 2018) .

Comment. Not Paederomimus group 3 after Sharp (1885: 439), to be placed in a further study.

Paederomimus nigrinus Sharp 1885

Paederomimus nigrinus Sharp 1885: 442 . Type locality: Nicaragua, Chontales . Type depository: BMNH HT. Distribution : Nicaragua .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus nigripennis Sharp 1885

Paederomimus nigripennis Sharp 1885: 446 . Type locality: Panama, Bugaba 800–1500 ft, Volcan de Chiriqui 2000–4000 ft . Type depository: BMNH 30 ST: 24 Bugaba , 6 Chiriqui [ MCP seen] ; FMNH 1 ST. Distribution : Panama .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus nigrocyaneus Schubert 1909

Paederomimus nigrocyaneus Schubert 1909: 294 . Type locality: Jalapa region , Veracruz, Mexico. Type depository: ZMHB 1 ST [ MCP seen] ; FMNH 1 ST. Distribution : Mexico . Note: distribution within Mexico (Navarrete- Heredia et al. 2002).

Paederomimus nobilis Sharp 1885

Paederomimus nobilis Sharp 1885: 442 . Type locality: Panama, Volcan de Chiriqui 4000–6000 ft . Type depository: BMNH HT. Distribution : Panama ; Nicaragua.

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus ornatellus Sharp 1885

Paederomimus ornatellus Sharp 1885: 449 . Type locality: Panama, Bugaba 800–1500 ft . Type depository: BMNH 8 ST. Distribution : Panama .

Paederomimus pallidus Bierig 1937

Paederomimus pallidus Bierig 1937: 197 . Type locality: Panamá : France Field (Zona del Canal). Type depository: FMNH 2 ST. Distribution : Panama.

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus palpalis (Sharp 1876)

Philonthus palpalis Sharp 1876a: 173 . Type locality: "Amazons". Type depository: BMNH HT ♂. Distribution : Brazil (Amazonas) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus pectoralis Sharp 1885

Paederomimus pectoralis Sharp 1885: 443 . Type locality: Guatemala: near the city, Capetillo, El Tumbador 2500 ft, Cerro Zunil 4000–5000 ft. Type depository: BMNH 9 ST: 6 Guatemala city, 1 El Tumbador, 1 Capetillo, 1 Cerro Zunil [ MCP seen]. Distribution: Mexico, Guatemala. Note: distribution, also within Mexico (Navarrete- Heredia et al. 2002).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus perforatus Sharp 1885

Paederomimus perforatus Sharp 1885: 451 . Type locality: Panama, Volcan de Chiriqui 20 00– 3000 ft . Type depository: BMNH HT. Distribution : Panama .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus praenobilis Bernhauer 1910

Paederomimus praenobilis Bernhauer 1910: 379 . Type locality: Cordoba, Veracruz . Type depository: FMNH 1 ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Paederomimus propinquus (Sharp 1876)

Philonthus propinquus Sharp 1876a: 176 . Type locality: Ega. Type depository: BMNH HT ♂. Distribution : Brazil (Amazonas) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Paederomimus punctipennis Sharp 1885

Paederomimus punctipennis Sharp 1885: 443 . Type locality: Panama, Bugaba 8 00 – 1500 ft. Type depository: BMNH 2 ST ♀. Distribution : Panama .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus regillus (Sharp 1876)

Philonthus regillus Sharp 1876a: 176 . Type locality: St. Paulo [Amazonas] . Type depository: BMNH 1 ST. Distribution : Brazil (Amazonas) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus salvini Sharp 1885

Paederomimus salvini Sharp 1885: 440 . Type locality: Panama, Bugaba 800–1500 ft, Volcan de Chiriqui 2000– 3000 ft . Type depository: BMNH 12 ST: 10 Bugaba , 2 Chiriqui [ MCP seen]. Distribution: Panama .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus sharpi Bernhauer & Schubert 1914

Paederomimus sharpi Bernhauer & Schubert 1914: 368 (new name for Paederomimus hilaris Sharp 1885: 441 ). Distribution: Panama.

= Paederomimus hilaris Sharp 1885: 441 (preoccupied, junior secondary homonym in Paederomimus of Philonthus hilaris Fauvel 1874: 246 ). Type locality: Panama, Volcan de Chiriqui 3000–4000 ft. Type depository: BMNH HT ♀. Note: as synonym of P. sharpi ( Bernhauer & Schubert 1914: 368) .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus simplex Sharp 1885

Paederomimus simplex Sharp 1885: 444 . Type locality: Guatemala, near the city, San Gerónimo and San Joaquin in Vera Paz. Type depository: BMNH 9 ST: 8 San Gerónimo, 1 San Joaquin [MCP seen]. Distribution: Guatemala;?Panama.

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus sulciceps Bernhauer 1917

Paederomimus sulciceps Bernhauer 1917a: 35 . Type locality: Chanchamayo , Peru . Type depository: FMNH HT. Distribution : Peru . Note: Peru ( Asenjo 2004).

Paederomimus sydowi Bernhauer 1917

Paederomimus sydowi Bernhauer 1917a: 37 . Type locality: Paulo (Umgebung von Ribeirão Preto Fundão). Type depository: FMNH HT, +?; ZMUH 2 +PT [destroyed?]. Distribution : Brazil (Goias, São Paulo) .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus tetricus Blackwelder 1943

Paederomimus tetricus Blackwelder 1943: 395 . Type locality: Trinidad : Port of Spain . Type depository: USNM HT ♀. Distribution : Trinidad .

Paederomimus traili (Sharp 1876)

Philonthus traili Sharp 1876a: 170 . Type locality: Ananá. Type depository: BMNH HT. Distribution : Brazil (Amazonas) . Note: comb. nov. ex Philonthus ( Sharp 1885: 438) ; distribution within Brazil (Asenjo et al. 2013).

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Paederomimus vicinus Sharp 1885

Paederomimus vicinus Sharp 1885: 444 . Type locality: Panama, Volcan de Chiriqui 2000–3000 ft . Type depository: BMNH HT ♀. Distribution : Panama .

Comment. Not Paederomimus group 3 ( Sharp 1885: 439), to be placed in a further study.

Figures

FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm.