Belonuchus Nordmann 1837

Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey, 2018, Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae), Zootaxa 4449 (1), pp. 1-95: 9-30

publication ID

https://doi.org/10.11646/zootaxa.4449.1.1

publication LSID

lsid:zoobank.org:pub:0734DC4E-2C9B-4CD2-B8FE-8F94589200E6

persistent identifier

http://treatment.plazi.org/id/215187C8-4C20-B355-FF6B-FC00F4ACD3FC

treatment provided by

Plazi

scientific name

Belonuchus Nordmann 1837
status

 

Belonuchus Nordmann 1837 

Belonuchus Nordmann 1837: 129  . Type species: Staphylinus haemorrhoidalis Fabricius, 1801  , by monotypy.

= Musicoderus Sharp 1885: 455  . Type species: Musicoderus cephalotes Sharp 1885  , des. by Lucas 1920: 425. Notes: as subgenus of Belonuchus ( Bernhauer 1916c: 266)  ; as synonym of Belonuchus ( Blackwelder 1943: 421)  .

Notes: There are 165 valid species of Belonuchus  distributed in CASA. A recent study ( Chani-Posse et al. 2018) shows Belonuchus  as a genus of Neotropical origin and generic limits so far undefined. The current concept of the genus is that of Smetana (1995), Chani-Posse (2014a) and Chani-Posse et al. (2018) but most of its Neotropical endemic species were never revised after being described and they do not fit into this concept. Species that may belong to the so called ‘ Belonuchus haemorrhoidalis  clade’ ( Chani-Posse et al. 2018) are here denoted with an asterisk *.

* Belonuchus abnormalis ( Sharp 1885)  , comb. nov. ( Figs. 1–2View FIGURES 1–12)

Philonthus abnormalis Sharp 1885: 397  . Type locality: Mexico: Oaxaca. Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

Remarks. Sharp explicitly mentioned two specimens (male and female) as part of the type series. The two specimens listed below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 1–2View FIGURES 1–12): card-mounted, “ ♂ (in ink) Philonthus abnormalis  Type D.S. Oaxaca. Mexico. Höge [in Sharp’s hand] // Type (round white label with red margin) // Oaxaca, Mexico. Hoege. (white label) // B. C. A. Col. I. 2. Philonthus abnormalis, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotype ♀ ( BMNH): with same locality and collection data as lectotype, and additional printed white label “Sp. Figured”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  ; determined as “ Belonuchus  ” by J. L. Navarrete-Heredia in 2002.

Belonuchus aequalis Sharp 1876 

Belonuchus aequalis Sharp 1876a: 161  . Type locality: Pará, Ega, St. Paulo [Amazonas] & Conceicão, Rio Mauhes.

Type depository: BMNH 6 ST: 4 Ega, 1 Pará, 1 St. Paulo [ MCPAbout MCP seen]  . FMNH 1View Materials ST: 1 Ega. Distribution: Brazil  . Notes: distribution in Brazil (Asenjo et al. 2013).

* Belonuchus agilis Erichson 1840 

Belonuchus agilis Erichson 1840: 423  . Type locality: Cuba. Type depository: ZMHB 2 ST: 1♂, 1♀ [ MCPAbout MCP seen].

Distribution: USA, Dominican Rep., Cuba, Jamaica, Haiti, Colombia. Notes: distribution including USA and Dominican Rep. ( Frank 2004); distribution, also within Cuba ( Peck 2005).

= Belonuchus chevrolati Fauvel 1863: 432  . Type locality: La Havane, Cuba. Type depository: IRSNBAbout IRSNB HT ♂ [ MCPAbout MCP seen]. Note: as synonym of B. agilis ( Fauvel 1869: 489)  .

Belonuchus albovariegatus Bernhauer 1916 

Belonuchus albovariegatus Bernhauer 1916c: 276  . Type locality: West-Columbien: Umgebung Cali am Rio Cauca, St. Antonio, 2000 m. Type depository: FMNH LT ♂, 2 PLT (des. Chani-Posse & Solodovnikov 2017). Distribution: Colombia.

= Linoderus navarretei López-García & Méndez-Rojas 2014: 86  . Type locality: Colombia, Quindío Dept., Filandia, Cruces, Parque Regional Natural Barbas-Bremen , 4°42’6.4” N 75°38’16.3” W, 2000 mGoogleMaps  . Type depositories: UNCB HT ♂, 13 PT; CIUQ 3 PT. Note : as synonym of B. albovariegatus ( Chani-Posse & Solodovnikov 2017: 90)  . 

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus alternans ( Sharp 1885) 

Philonthus alternans Sharp 1885: 418  . Type locality: Mexico: Jalapa, Oaxaca. Type depository: BMNH 3 ST; FMNH 1ST. Distribution: Mexico. Notes: comb. nov. ex Philonthus ( Navarrete-Heredia et al. 2002: 334)  ; distribution within Mexico ( Márquez 2006).

* Belonuchus alternatus Sharp 1885 

Belonuchus alternatus Sharp 1885: 430  . Type locality: Guatemala, near the City, San Geronimo; Nicaragua, Chontales; Panama, Bugaba. Type depository: BMNH #? ST: several, from all type localities [ MCPAbout MCP seen]. FMNHAbout FMNH 4 ST: Panama, Bugaba. Distribution: Panama, Nicaragua, Guatemala. Note: misspelled as alternans  in error by Bernhauer & Schubert (1914: 369) (not Philonthus alternans Sharp 1885: 418  ).

* Belonuchus altissimus Bernhauer 1916 

Belonuchus altissimus Bernhauer 1916c: 275  . Type locality: Monte Socorro, West-Cordilleren, West-Columbien [3800m on type label]. Type depository: FMNHAbout FMNH HT. Distribution : Colombia  .

Belonuchus altivagans Bernhauer 1916 

Belonuchus altivagans Bernhauer 1916c: 272  . Type locality: West-Columbien: Pacho, 2000m, Ost-Cordilleren, West-Columbien. Type depository: FMNH 2View Materials ST. Distribution : Colombia  .

Comment. Not Belonuchus  , to be placed in a further study.

Belonuchus aluticollis (Bernhauer 1921) 

Philonthus aluticollis Bernhauer 1921c: 107  . Type locality: Cueva Iturbe, 3700m, Jujuy, Argentina [LT without "Cueva"]. Type depository: FMNH LT ♀, MACN 2 PLT. Distribution: Argentina. Note: comb. nov. ex Philonthus  and lectotype designation ( Chani-Posse 2010: 8); redescription ( Chani-Posse 2011b: 215).

Belonuchus amplipennis Fauvel 1891 

Belonuchus amplipennis Fauvel 1891: 117  . Type locality: Colonie Tovar, San Esteban [ Venezuela]. Type depository: IRSNBAbout IRSNB 2 ST [ MCPAbout MCP seen]. Distribution: Venezuela.

Belonuchus amplus Blackwelder 1943 

Belonuchus amplus Blackwelder 1943: 425  . Type locality: St. Lucia: 0.5 mile northwest of St. Remy on road to Soufrière (2.5 miles southeast of Soufrière)  . Type depository: USNMAbout USNM HT ♀, 1 PT ♀. Distribution: Lesser Antilles ( St. Lucia)  .

Belonuchus andinus Bernhauer 1916 

Belonuchus andinus Bernhauer 1916c: 278  (attributed to Fauvel in litt.). Type locality: Yuracarès, Bolivien. Type depository: FMNH 1View Materials ST; ZMUH 1View Materials +ST? [destroyed?]  ; IRSNB 2View Materials +3? ST: only two of them in Bernhauer´s hand [ MCPAbout MCP seen]. Distribution : Bolivia. 

Belonuchus angusticeps Scheerpeltz 1960 

Belonuchus (Musicoderus) angusticeps Scheerpeltz 1960: 117  . Type locality: Cochabamba ( Bolivien), in den Yungas de Palmar , 2000m  . Type depository: NMW 2View Materials ST; FMNH 2View Materials ST. Distribution : Bolivia  .

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus angusticollis Bernhauer 1906 

Belonuchus angusticollis Bernhauer 1906: 330  . Type locality: Chanchamayo , Peru  . Type depository: FMNHAbout FMNH HT. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Belonuchus antiguae Blackwelder 1943 

Belonuchus antiguae Blackwelder 1943: 434  . Type locality: Antigua: near summit of Boggy Peak , parish of St. Mary, elevation 1300 feet  . Type depository: USNMAbout USNM HT ♂, 5 PT; BMNH 1 PT; FMNH 1View Materials PT. Distribution: Antigua. Notes: distribution within Lesser Antilles (Northern Leeward Is.) ( Peck 2011a); generic affiliation unclear. 

Belonuchus apicicornis Bernhauer 1916 

Belonuchus (Musicoderus) apicicornis Bernhauer 1916c: 270  . Type locality: West-Columbien: Cali am Rio Cauca ( Rio Aquatal , 1800m; S. Antonio, 2000m; Rio Vitaco, 2200m). Type depository: FMNH 2View Materials ST;? 1 ST. Distribution: Colombia  .

* Belonuchus apiciventris ( Sharp 1885) 

Philonthus apiciventris Sharp 1885: 419  . Type locality: Mexico: Juquila. Type depository: BMNH HT ♂. Distribution: Mexico. Notes: in cave collections ( Navarrete-Heredia & Márquez-Luna 1993); cited as apaciventris, characters. habits, natural history notes, host mushrooms (Navarrete-Heredia 1995); comb. nov. ex Philonthus ( Jiménez-Sánchez et al. 2000)  ; distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).

Belonuchus argentinus Bernhauer 1916 

Belonuchus argentinus Bernhauer 1916c: 278  . Type locality: Misiones, Argentina. Type depositary: FMNHAbout FMNH HT, 2 PT; MLPA 8 PT (as ST in error, Fernandez et al. 2007). Distribution: Argentina, Brazil, Bolivia. Notes: MLPA "ST", Fernandez et al. 2007, but original reference to "Das typische Stück …"; name cited by Bruch (1915: 503) as nomen nudum, attributed to Bernhauer in litteris; distribution in Brazil (Asenjo et al. 2013); generic affiliation unclear.

* Belonuchus armatus Sharp 1876 

Belonuchus armatus Sharp 1876a: 163  . Type locality: Abacaxis. Type depository: BMNH HT ♂. Distribution : Brazil (Amazonas)  . Note: distribution in Brazil (Asenjo et al. 2013).

Belonuchus arrogans Bernhauer 1916 

Belonuchus arrogans Bernhauer 1916c: 274  . Type locality: West-Columbien : Ost-Cordilleren, Pacho, 2000 m  . Type depository: FMNHAbout FMNH HT ♂. Distribution : Colombia. Note: generic affiliation unclear. 

* Belonuchus asperatus Bernhauer 1906 

Belonuchus asperatus Bernhauer 1906: 336  . Type locality: Pachitea, Peru. Type depository: FMNHAbout FMNH HT ♂. Distribution: Colombia, Ecuador, Peru, Bolivia. Notes: distribution ( Scheerpeltz 1960: 120; Asenjo 2004).

Belonuchus atticida ( Wasmann 1925) 

Plociopterus atticida Wasmann 1925: 71  . Type locality: Colonia alpina, Rio d. Janeiro [LT]. Type depository: NHMEAbout NHME LT ♂, #? PLT. Distribution: Brazil, Paraguay. Notes: comb. nov. ex Plociopterus  and lectotype designation ( Chatzimanolis & Chani-Posse 2016: 214); with Acromyrmex niger  , Atta sexdens  [orig.; Navarrete-Heredia 2001].

= Hesperus eidmanni Scheerpeltz 1936: 484  . Type locality: Villa Rica in Paraguay [LT]; Umgebung von Mendes, E. d. Rio [4 PLT; Brazil]. Type depository: NMWAbout NMW (Scheerpeltz) LT ♂, 4 PLT (des. Chatzimanolis & Chani- Posse 2016: 214). Distribution: Brazil, Paraguay. Notes: distribution within Brazil (Asenjo et al. 2013); as synonym of B. atticida ( Chatzimanolis & Chani-Posse 2016: 214)  ; in Atta sexdens  fungus chamber [orig.]; also Acromyrmex lobicornis  fungus chamber ( Navarrete-Heredia 2001).

Belonuchus avidus Erichson 1840 

Belonuchus avidus Erichson 1840: 423  . Type locality: Novae Valenciae in Columbia [ Venezuela, teste Herman 2001d: 1594]. Type depository: ZMHB 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution: Colombia, Venezuela.

Belonuchus barbicornis Bernhauer 1916 

Belonuchus (Musicoderus) barbicornis Bernhauer 1916c: 271  . Type locality: West-Columbien: Alto de las cruces, 2200m & S. Antonio, 2200m. Type depository: FMNH 5View Materials ST. Distribution : Colombia  .

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus basiventris ( Sharp 1885) 

Philonthus apiciventris Sharp 1885: 418  . Type locality: Mexico: Cordova, Puebla, Yolos, Oaxaca, Jalapa. Type depository: BMNH 2 ST. Distribution: Mexico. Notes: comb. nov. ex Philonthus ( Jiménez-Sánchez et al. 2000)  ; in Atta mexicana  external debris [ Navarrete-Heredia 2001; distribution ( Márquez & Asiain 2012).

* Belonuchus batesi Sharp 1876 

Belonuchus batesi Sharp 1876a: 157  . Type locality: Ega. Type depository: BMNH HT. Distribution : Brazil (Amazonas)  . Note: distribution in Brazil (Asenjo et al. 2013).

* Belonuchus bidens Sharp 1885 

Belonuchus bidens Sharp 1885: 430  . Type locality: Mexico, Jalapa, Oaxaca, Mirador, Tuxtla; Nicaragua, Chontales. Type depository: BMNH 5 ST: from all type localities [ MCPAbout MCP seen]. Distribution: Nicaragua, Mexico. Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2004, 2006, Márquez & Asiain 2012, Asiain et al. 2015).

Belonuchus bipunctatus Bernhauer 1935 

Belonuchus bipunctatus Bernhauer 1935: 90  . Type locality: Kolumbien; Cali (Villa Elvira, 1800m). Type depository: FMNHAbout FMNH 2 ST; BMNH #?. Distribution: Colombia, Costa Rica.

* Belonuchus boops Sharp 1885 

Belonuchus boops Sharp 1885: 435  . Type locality: Panama, Bugaba   . Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution : Panama  .

Belonuchus boxi Bernhauer 1934 

Belonuchus boxi Bernhauer 1934b: 212  . Type locality: Tucumán  . Type depository: FMNH 1View Materials ST ♂. Distribution : Argentina. Note: generic affiliation unclear. 

Belonuchus brasilianus Bernhauer 1917 

Belonuchus brasilianus Bernhauer 1917b: 19  . Type locality: S. Catharina, Brasilien  . Type depository: FMNHAbout FMNH HT. Distribution : Brazil (Santa Catarina)  . Note: distribution in Brazil (Asenjo et al. 2013); generic affiliation unclear.

Belonuchus breviceps Fauvel 1891 

Belonuchus breviceps Fauvel 1891: 117  . Type locality: Colonie Tovar [ Venezuela]. Type depository: IRSNBAbout IRSNB 2 ST [ MCPAbout MCP seen]. Distribution: Venezuela. Note: generic affiliation unclear.

Belonuchus bruchi Bernhauer 1912 

Belonuchus bruchi Bernhauer 1912b: 175  . Type locality: Buenos Aires prov., Argentina. Type depository: FMNH HT. Distribution: Argentina. Note: not bruchi Bernhauer 1934b  (see duplicatus  ).

* Belonuchus bruchianus Bernhauer 1911 

Belonuchus bruchianus Bernhauer 1911: 413  . Type locality: Misiones prov., Argentina  . Type depository: FMNHAbout FMNH HT ♀. Distribution : Argentina  .

Belonuchus bugnioni Fauvel 1901 

Belonuchus bugnioni Fauvel 1901: 86  . Type locality: Martinique. Type depository: IRSNBAbout IRSNB HT ♀ [ MCPAbout MCP seen]. Distribution: Martinique, Guadeloupe. Note: distribution within (and endemic to) Lesser Antilles ( Peck 2011b: Martinique, Peck et al. 2014: Guadeloupe Archipelago). Note: generic affiliation unclear.

= Belonuchus eximius Bernhauer 1917c: 105  . Type locality: Guadeloupe  . Type depository: FMNH 2View Materials ST. Distribution : Guadeloupe. Note: as synonym of B. bugnioni ( Blackwelder 1943: 428)  . 

Belonuchus cacao Blackwelder 1943 

Belonuchus cacao Blackwelder 1943: 434  . Type locality: Trinidad: one-half mile south-southwest of Manzanilla junction on the Plum Road. Type depository: USNMAbout USNM HT ♂, 1 PT ♀. Distribution: Venezuela, Trinidad. Note: Venezuela, biology, in Heliconia  bracts ( Frank & Barrera 2010).

Belonuchus candens ( Erichson 1840) 

Philonthus candens Erichson 1840: 460  . Type locality: Bogotae in Columbia interiore. Type depository: ZMHB 1+3 ST: 1 ♂ in Erichson´s hand, 3 ♀ [ MCPAbout MCP seen]. Distribution: Colombia, Brazil, Peru. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 369)  ; distribution including Peru, Brazil questioned (Asenjo et al. 2013).

Comment. Not Belonuchus  , to be placed in a further study.

Belonuchus cariniventris Bernhauer 1918 

Belonuchus cariniventris Bernhauer 1918: 225  . Type locality: Mexico: Jalappa [Xalapa, Veracruz]  . Type depository: FMNH 3View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002); generic affiliation unclear.

Belonuchus castaneus Chani-Posse 2006 

Belonuchus castaneus Chani-Posse 2006: 366  (new name for Belonuchus richteri Bernhauer 1922: 15  ).

= Belonuchus richteri Bernhauer 1922: 15  (preoccupied, junior secondary homonym in Belonuchus  of Philonthus richteri Bernhauer 1912a: 44  ). Type locality: Misiones, Argentina. Type depository: FMNHAbout FMNH LT ♂, 2 PLT (des. Chani-Posse 2011b: 219). Distribution: Argentina.

Belonuchus cavipennis Bernhauer 1916 

Belonuchus (Musicoderus) cavipennis Bernhauer 1916c: 267  , attributed to Fauvel in litteris. Type locality: Yuracarès , Bolivien  . Type depository: FMNH 1View Materials ST; ZMUH 1View Materials +ST?. Distribution : Bolivia. Note: generic affiliation unclear. 

* Belonuchus celatus ( Sharp 1885)  , comb. nov. ( Figs. 3–4View FIGURES 1–12)

Philonthus celatus Sharp 1885: 420  . Type locality: Mexico: Mirador   , Oaxaca, Tehuacan, Yolos, Cuernavaca. Type depository: BMNH 10 ST: from all type localities [ MCPAbout MCP seen]. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Remarks. Sharp did not specify the number of specimens he examined for naming this species, but it seems likely that he based his description on at least two specimens of different sex. Ten specimens were located in the BMNH under this name. All the ten specimens listed below agree with the original description by Sharp (1885) and are considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated (BMNH, Figs. 3–4View FIGURES 1–12): card-mounted, “ Philonthus celatus  Type D.S. Oaxaca. Höge” [in Sharp’s hand] // Type (round white label with red margin) // Oaxaca, Mexico. Hoege (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 2 ♀, one of them “Cuernavaca” (white label) [in Sharp’s hand], the other one “Yolos” (white label) [in Sharp’s hand], both of them with the following additional labels, “Mexico. Salle. Coll (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp.  (white label)”; 1 ♂, 4 ♀, card-mounted, ♂ with “ Philonthus celatus Mirador. Höge  ” [in Sharp’s hand]), “Mirador, Mexico. Hoege (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp.  (white label)”; 2 ♀, card-mounted (one of them also “ Philonthus celatus  Type D.S. Tehuacan. Höge” [in Sharp’s hand]), “Tehuacan, Mexico. Hoege (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp.  (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

Belonuchus cephalotes ( Sharp 1885) 

Musicoderus cephalotes Sharp 1885: 455  . Type locality: Panama, Bugaba 800 to 1500 feet. Type depository: BMNH 12 ST: all of them with labels as "Bugaba" but only two of them with the elevation data [ MCPAbout MCP seen]; FMNHAbout FMNH 3 ST. Distribution: Panama, Venezuela. Notes: comb. nov. ex Musicoderus ( Scheerpeltz 1933: 1377)  ; Venezuela, biology, in Heliconia  bracts ( Frank & Barrera 2010); generic affiliation unclear.

* Belonuchus cifuentesi Rodríguez & Navarrete-Heredia 2016 

Belonuchus cifuentesi Rodríguez & Navarrete-Heredia 2016: 445  . Type locality: Mexico  : Jalisco, Teocuitatlán de Corona, Cerro de García , 20°09´N 103°19´W, 2314 m. Type depository: CZUGAbout CZUG HT ♂, 117 PT ( CZUGAbout CZUG, MUD, CNINAbout CNIN)GoogleMaps  . Distribution: Mexico. Note: in decomposing agave (Agave inaequidens) [orig.].

* Belonuchus clypeatus Sharp 1876 

Belonuchus clypeatus Sharp 1876a: 160  . Type locality: Ega   . Type depository: BMNH HT. Distribution : Brazil (Amazonas)  , Peru. Note: Peru (Newton 2015); distribution within Brazil (Asenjo et al. 2013).

Belonuchus coelestinus Bernhauer 1908 

Belonuchus coelestinus Bernhauer 1908: 330  . Type locality: St. Vincent. Type depository: NMWAbout NMW 1+ST, FMNHAbout FMNH 1ST. Distribution: Lesser Antilles ( St. Vincent, Grenada). Note: distribution within Lesser Antilles ( St. Vincent) ( Peck 2010). Note: generic affiliation unclear.

= Belonuchus coeruleus Cameron 1922: 122  . Type locality: St. Vincent. Type depository: BMNH 2ST. Distribution: Lesser Antilles ( St. Vincent, Grenada). Notes: as synonym of B. coelestinus ( Blackwelder 1943: 426)  .

= Belonuchus smithi Cameron 1922: 122  (as synonym of B. coeruleus  ), attributed to Fauvel in litteris. Type locality: St. Vincent. Type depository: BMNH 2 ST. Distribution: Lesser Antilles ( St. Vincent, Grenada). Note: as synonym of B. coelestinus ( Blackwelder, 1943: 426)  ; unavailable ( ICZN 1999: Art. 11.6).

Belonuchus coelestis Bernhauer 1916 

Belonuchus (Musicoderus) coelestis Bernhauer 1916c: 266  (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien  . Type depository: FMNH 1View Materials ST; ZMUH 1View Materials +ST? Distribution : Bolivia  .

Comment. Not Belonuchus  , to be placed in a further study.

Belonuchus coeruleispennis Motschulsky 1858 

Belonuchus coeruleispennis Motschulsky 1858: 668  . Type locality: Brésil intérieur. Type depository: ZMUM? Distribution: Brazil. Note: subsequently spelled coeruleipennis (e.g., Motschulsky 1870: 49; Bernhauer & Schubert 1914: 369), but coeruleispennis  maintained as correct spelling by Herman (2001d: 2522); distribution within Brazil unknown (Asenjo et al. 2013).

* Belonuchus cognatus Sharp 1885 

Belonuchus cognatus Sharp 1885: 429  . Type locality: British Honduras, Belize; Guatemala, El Reposo, Las Mercedes, San Isidro, Mirandilla; Nicaragua, Chontales; Panama, Bugaba. Type depository: BMNH #? ST: several, from all type localities [ MCPAbout MCP seen]; IRSNBAbout IRSNB 5 ST: 4 Bugaba, two of them labelled as “Ext-ypis” [ MCPAbout MCP seen]; 1 Chontales [ MCPAbout MCP seen]; FMNHAbout FMNH 3 ST: 3 Bugaba. Distribution: Belize, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad, Brazil, Argentina, St. Vincent. Notes: distribution within Lesser Antilles ( St. Vincent) ( Peck 2010); within (and including) Brazil (Asenjo et al. 2013).

* Belonuchus colon ( Sharp 1885) 

Philonthus colon Sharp 1885: 419  . Type locality: Mexico: Cordova. Type depository: BMNH 2 ST. Distribution: Mexico. Notes: comb. nov. ex Philonthus  , distribution ( Navarrete-Heredia et al. 2002: 334); distribution within Mexico ( Márquez 2006).

Belonuchus columbicus Bernhauer 1916 

Belonuchus (Musicoderus) columbicus Bernhauer 1916c: 271  . Type locality: West-Columbien: Umgebung von Cali ( St. Antonio , 2000m). Type depository: FMNH 2View Materials ST;? 1 ST. Distribution: Colombia  .

Comment. Not Belonuchus  , to be placed in a further study.

Belonuchus connexus ( Say 1830) 

Staphylinus connexus Say 1830: 35  , also 1834: 448. Type locality: Mexico. Type depository:? #?ST. Distribution: Mexico. Note: comb. nov. ex Staphylinus ( Erichson 1840: 927)  ; distribution within Mexico (Navarrete- Heredia et al. 2002). Note: generic affiliation unclear.

Belonuchus convexus ( Bernhauer 1908) 

Musicoderus convexus Bernhauer 1908: 331  . Type locality: Pozuzo, Peru. Type depository: FMNHAbout FMNH HT ♀?: Distribution: Peru, Nicaragua. Notes: comb. nov. ex Musicoderus  , distribution Nicaragua ( Scheerpeltz 1933: 1377); Peru ( Asenjo 2004).

Belonuchus cordilleranus Bernhauer 1916 

Belonuchus (Musicoderus) cordilleranus Bernhauer 1916c: 270  . Type locality: Columbien: Ost-Cordilleren, 450– 900m. Type depository: FMNH 3View Materials ST. Distribution : Colombia. Note: generic affiliation unclear. 

* Belonuchus corticalis ( Sharp 1885)  , comb. nov. ( Figs. 5–6View FIGURES 1–12)

Philonthus corticalis Sharp 1885: 413  . Type locality: Panama, Volcan de Chiriqui 2000–3000 ft  . Type depository: BMNH HT ♂. Distribution : Panama  .

Remarks. The “unique” specimen in BMNH is confirmed as the holotype.

Type material examined. Holotype ♂ ( BMNH, Figs. 5–6View FIGURES 1–12): card-mounted, “ ♂ (in ink) Philonthus corticalis  Type D.S. V. de Chiriqui 2–3000 ft. [in Sharp’s hand] // Type (round white label with red margin) // V. de Chiriqui 2– 3000 ft., 2–3000 ft. Champion (white label) // B. C. A. Col. I. 2. Philonthus corticalis, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

Belonuchus cribriceps Bernhauer 1921 

Belonuchus cribriceps Bernhauer 1921c: 172  . Type locality: San Roque, Corrientes [ Argentina]. Type depository: FMNHAbout FMNH 1 ST. Distribution: Argentina, Brazil. Note: distribution within (and including) Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.

Belonuchus danforthi Blackwelder 1943 

Belonuchus danforthi Blackwelder 1943: 429  . Type locality: Puerto Rico: about 13 miles east of Maricao and north of Yauco, elevation about 3000 feet  . Type depository: USNMAbout USNM HT ♂. Distribution : Puerto Rico  .

* Belonuchus decipiens Sharp 1876 

Belonuchus decipiens Sharp 1876a: 159  . Type locality: Ega. Type depository: BMNH 6 ST, FMNHAbout FMNH 1 ST. Distribution: Brazil, Argentina, Paraguay; Madagascar?; Indonesia (Lombok)? Notes: redescription ( Silvestri 1945); distribution, including Argentina and Paraguay ( Chani-Posse 2011b); records from Brazil only (Asenjo et al. 2013).

* Belonuchus dichrous Erichson 1840 

Belonuchus dichrous Erichson 1840: 420  . Type locality: Monte Orizaba in Mexico. Type depository: ZMHB 4 ST: 2 in Bernhauer´s hand " dichrous  ", 2 with "Orizaba" [ MCPAbout MCP seen]. Distribution: Mexico, Guatemala, Venezuela. Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).

* Belonuchus diversicornis Bernhauer 1917 

Belonuchus diversicornis Bernhauer 1917c: 104  . Type locality: Espirito Santo, Brasilien  . Type depository: FMNHAbout FMNH HT. Distribution : Brazil (Espírito Santo)  . Note: distribution within Brazil (Asenjo et al. 2013).

* Belonuchus docilis Sharp 1885 

Belonuchus docilis Sharp 1885: 435  . Type locality: Guatemala, Pantaleon; Panama, Bugaba, Tolé. Type depository: BMNH 8 ST: 2 Pantaleón, 3 Bugaba, 3 Tolé [ MCPAbout MCP seen]. FMNHAbout FMNH 2 ST: 2 Bugaba. Distribution: Guatemala,? Costa Rica, Panama, Brazil. Note: distribution within (and including) Brazil (Asenjo et al. 2013).

Belonuchus dominicus Blackwelder 1943 

Belonuchus dominicus Blackwelder 1943: 431  . Type locality: Dominica: Hillsboro Estate, at mouth of the Layou River, Leeward side. Type depository: USNMAbout USNM HT ♂, 4 PT; Blackwelder 1 PT; FMNHAbout FMNH 1 PT. Distribution: Dominica Note: distribution within Dominica (Peck 2006); generic affiliation unclear.

Belonuchus duplicatus Blackwelder 1944 

Belonuchus duplicatus Blackwelder 1944: 137  (new name for Belonuchus bruchi Bernhauer 1934: 313  ).

= Belonuchus bruchi Bernhauer 1934b: 213  (preoccupied, junior primary homonym of Belonuchus bruchi Bernhauer 1912: 175  ). Type locality: Tucuman, Argentina. Type depository: FMNH HT. Distribution: Argentina. Note: generic affiliation unclear.

Belonuchus ecuadorensis Bernhauer 1935 

Belonuchus (Musicoderus) ecuadorensis Bernhauer 1935: 91  . Type locality: Ecuador  . Type depository: BMNH #?ST; FMNH 1View Materials ST. Distribution : Ecuador  .

* Belonuchus ephippiatus ( Say 1830) 

Staphylinus ephippiatus Say 1830: 35  , also 1834: 448. Type locality: Mexico. Type depository: MCZAbout MCZ NT ♂. Distribution: USA, Mexico, Colombia, Venezuela; adventive in Hawaii. Notes: comb. nov. ex Staphylinus ( Erichson 1840: 927)  ; neotype designation, redescription ( Smetana 1995: 746); distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).

= Belonuchus simplex Sharp 1885: 432  . Type locality: Mexico: Oaxaca. Type depository: BMNH LT ♂, 2 PLT; FMNH 1 PLT?. Notes: as synonym of B. ephippiatus ( Fauvel 1901: 86)  ; lectotype designation ( Smetana 1995: 748).

= Belonuchus punctiventris Casey 1915: 446  . Type locality: Benson, Arizona. Type depository: USNMAbout USNM LT ♂, 13 PLT. Notes: synonym of B. ephippiatus ( Smetana 1995: 746)  ; lectotype designation ( Smetana 1995: 748).

= Belonuchus laticeps Casey 1915: 446  . Type locality: near Indio, California. Type depository: USNM LT ♂. Notes: synonym of B. ephippiatus ( Smetana 1995: 746)  ; lectotype designation ( Smetana 1995: 749).

= Belonuchus jacobianus Casey 1915: 447  . Type locality: San Diego, California. Type depository: USNM LT ♂. Notes: synonym of B. ephippiatus ( Smetana 1995: 746)  ; lectotype designation ( Smetana 1995: 749).

= Belonuchus schaefferi Cooper 1933: 545  . Type locality: Brownsville , Texas  . Type depository: CUICAbout CUIC HT ♀; AMNH 1View Materials PT. Note : synonym of B. ephippiatus ( Smetana 1995: 746)  . 

* Belonuchus erichsoni Bernhauer 1917 

Belonuchus erichsoni Bernhauer 1917d: 223  . Type locality: [Mexiko]. Type depository: FMNH HT ♂. Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).

* Belonuchus erythropterus Solsky 1868 

Belonuchus erythropterus Solsky 1868: 137  . Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #? ST, not seen by Smetana (1995). Distribution: USA (TX), Mexico. Note: redescription, distribution ( Smetana 1995: 752); distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).

Belonuchus excelsus Bernhauer 1917 

Belonuchus excelsus Bernhauer 1917b: 19  . Type locality: St. Antonio , 2000m, Umgebung von Cali, West- Columbien. Type depository: FMNHAbout FMNH HT. Distribution: Colombia  .

* Belonuchus extremus ( Sharp 1885)  , comb. nov. ( Figs. 7–8View FIGURES 1–12)

Philonthus extremus Sharp 1885: 415  . Type locality: Guatemala, San Gerónimo. Type depository: BMNH HT. Distribution : Guatemala  .

Remarks. The “unique” specimen in BMNH is confirmed as the holotype.

Type material examined. Holotype ♂ (BMNH, Figs. 7–8View FIGURES 1–12): card-mounted, “ Philonthus extremus  Type D.S. San Geronimo, Guatemala. Champion. [in Sharp’s hand] // Type (round white label with red margin) // San Geronimo, Verapaz. Champion. (white label) // B. C. A. Col. I. 2. Philonthus extremus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus falsus Bernhauer 1917 

Belonuchus falsus Bernhauer 1917c: 105  . Type locality: Venezuela  . Type depository: FMNH 1View Materials ST; ZMUH 1View Materials +ST?. Distribution : Venezuela  .

* Belonuchus fassli Bernhauer 1917 

Belonuchus fassli Bernhauer 1917a: 37  . Type locality: West-Columbien: Umgebung von Cali am Rio Cauca (Cañon del Mte Tolima, 1700m). Type depository: FMNHAbout FMNH HT. Distribution: Colombia  .

Belonuchus flavicoxis (Bernhauer 1912) 

Philonthus flavicoxis Bernhauer 1912a: 43  . Type locality: Tucuman prov., Argentina. Type depository: FMNH LT ♀. Distribution: Argentina. Notes: comb. nov. ex Philonthus  and lectotype designation ( Chani-Posse 2010: 8); distribution (Chani-Posse 2004); redescription ( Chani-Posse 2011b: 221).

* Belonuchus flavipennis Solsky 1870 

Belonuchus flavipennis Solsky 1870: 263  . Type locality: " Mexique "  . Type depository: ZINAbout ZIN? HT ♀. Distribution : Mexico  ; Costa Rica?  , Panama?  . Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).

* Belonuchus flavocaudatus Bernhauer 1921 

Belonuchus flavocaudatus Bernhauer 1921c: 170  . Type locality: Hornadita , 3400m, Jujuy, Argentina  . Type depository: FMNH 1View Materials ST; MACN 1View Materials ST. Distribution : Argentina  .

Belonuchus forsteri Scheerpeltz 1960 

Belonuchus (Musicoderus) forsteri Scheerpeltz 1960: 115  . Type locality: Bolivien, Yungas de Arepucho , Sihuencas, 2200–2500m  . Type depository: NMWAbout NMW ( Scheerpeltz ) HT ♀. Distribution: Bolivia  .

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus fossulatus Wasmann 1887 

Belonuchus fossulatus Wasmann 1887: 414  . Type locality: Blumenau (Prov. St. Catharina, Brasil)  . Type depository: NHMEAbout NHME (Wasmann) 1ST ♀. Distribution : Brazil (Santa Catarina)  . Note: distribution within Brazil (Asenjo et al. 2013).

Belonuchus frater Bernhauer 1906 

Belonuchus frater Bernhauer 1906: 333  . Type locality: Peru  . Type depository: FMNH 1View Materials ST. Distribution : Peru  . Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.

* Belonuchus gagates Erichson 1840  Belonuchus gagates Erichson 1840: 424  . Type locality: Puerto Rico & Cuba. Type depository: ZMHB LT ♂, 7

PLT. Distribution: USA (FL), Cuba, Jamaica, Hispaniola, Puerto Rico, Cayman Is., St. Thomas, St. John, Montserrat, Bahamas. Note: lectotype designation, redescription and distribution ( Smetana 1995: 737); distribution, also within Cuba ( Peck 2005); distribution within (and including) Cayman Is. (Frank et al. 2011).

= Belonuchus gowdeyi Bernhauer 1935: 91  . Type locality: Hope Gardens, Jamaica. Type depository: FMNH LT, BMNH PLT?. Distribution: Jamaica. Note: lectotype designation ( Smetana 1995: 739); as synonym of B. gagates ( Blackwelder 1943: 424)  .

= Belonuchus minax Erichson 1840: 424  . Type locality: ins. St. Thomae & St. Johannis. Type depository: ZMHB 4 ST [ MCPAbout MCP seen]. Distribution: Lesser Antilles (St. Thomas). Note: synonym of B. gagates ( Blackwelder 1943: 424)  ; not in Smetana (1995: 737).

= Musicoderus nigrocaeruleus Cameron 1913: 175  . Type locality: Jamaica, Newcastle district, 3000 ft above sea level. Type depository: BMNH (Cameron) HT ♂. Distribution: Jamaica. Notes: as synonym of B. gagates ( Blackwelder 1943: 424)  ; although both male and female characters were used in the original description, Cameron (1913) only refers to a “type” deposited in his collection. Smetana (1995: 738) examined eight conspecific specimens under this name in the Cameron collection at the BMNH, one of them being the specimen chosen as “Type” by the author, thus the holotype.

Belonuchus ganglbaueri Bernhauer 1906 

Belonuchus ganglbaueri Bernhauer 1906: 331  . Type locality: Bolivien  . Type depository: FMNH 1View Materials ST, NMWAbout NMW #? ST. Distribution : Bolivia. Note: generic affiliation unclear. 

Belonuchus godmani ( Sharp 1885) 

Philonthus godmani Sharp 1885: 413  . Type locality: Mexico: Guatemala: Calderas [LT]. Type depository: BMNH LT ♀, 7+ PLT. Distribution: Mexico, Guatemala, Costa Rica, Panama. Notes: comb. nov. ex Philonthus ( Navarrete-Heredia et al. 2002)  ; distribution, also within Mexico ( Navarrete-Heredia 2002, Márquez 2004, 2006); lectotype designation, redescription, distribution (Márquez 2011: 206).

Comment. Not Belonuchus  , to be placed in a further study.

Belonuchus goeldii ( Wasmann 1925) 

Plociopterus goeldii Wasmann 1925: 72  . Type locality: Colonia alpina, Rio d. Janeiro [LT]. Type depository: NHMEAbout NHME LT ♀, #? PLT. Distribution: Brazil, Paraguay, Bolivia. Notes: comb. nov. ex Plociopterus  and lectotype designation ( Chatzimanolis & Chani-Posse 2016: 215); with Acromyrmex niger  [orig.; Navarrete-Heredia 2001].

Belonuchus gracilis ( Sharp 1885) 

Musicoderus gracilis Sharp 1885: 456  . Type locality: Nicaragua, Chontales  . Type depository: BMNH HT ♀. Distribution : Nicaragua  .

* Belonuchus haemohrroidalis ( Fabricius 1801) 

Staphylinus haemorrhoidalis Fabricius 1801: 596  . Type locality: America meridionali. Type depository: ZMUK 4ST (until recently at ZMUCAbout ZMUC, Zimsen 1964): 3ST (former Sehested-Tønder Lund collection), 1ST (former Fabricius´s personal collection, seen AFN 2010). Distribution: Colombia, Venezuela, French Guiana, Peru, Bolivia, Brazil, Paraguay, Argentina. Notes: comb. nov. ex Staphylinus  , cited as haemoroidalis ( Nordmann 1837: 130); preoccupied, junior primary homonym of Staphylinus haemorrhoidalis Gmelin 1790: 2036  and junior primary homonym of Staphylinus haemorrhoidalis Olivier 1795  : (42) 11; ICZN application to conserve name ( Herman 2001a: 42), conserved in Opinion 2053 ( ICZN 2003); distribution including Peru, also within Brazil (Asenjo et al. 2013).

Belonuchus hispaniolus Blackwelder 1943 

Belonuchus hispaniolus Blackwelder 1943: 422  . Type locality: Dominican Republic: San Francisco Mountains   . Type depository: USNMAbout USNM HT ♀, 1PT ♀. Distribution : Dominican Rep., Haiti. Note: Haiti ( Frank & Barrera 2010). 

Belonuchus holisoides Sharp 1876 

Belonuchus holisoides Sharp1876a: 161  . Type locality: Ega , Brazil  . Type depository: BMNH 2 ST: 1 Ega together with other 4 specimens from probably a different species, all of them under the name " Metoponcus holisoides Sharp  "[ MCPAbout MCP seen]. Distribution : Brazil (Amazonas)  , Argentina. Note: distribution only in Brazil (Asenjo et al. 2013).

Comment. It probably belongs to Holisus Erichson. 

Belonuchus iconnicoffi Plavilstschikov 1929  , nomen nudum

Belonuchus iconnicoffi Plavilstschikov 1929: 30  , as "n. sp. (Bernhauer in litteris)"; nomen nudum, no description. Distribution: Peru. Note: listed in error as valid species in Blackwelder (1944) and Asenjo (2004); not in Herman (2001d)

Belonuchus ielskii ( Solsky 1872) 

Philonthus ielskii Solsky 1872: 296  . Type locality: Puna, Lima [Amérique du Sud]. Type depository: ZIN? (Solsky) #?ST. Distribution: Peru. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 370)  ; as jelskii ( Herman 2001d: 2527, Asenjo 2004: 61). Note: generic affiliation unclear.

* Belonuchus impressifrons Sharp 1876 

Belonuchus impressifrons Sharp 1876a: 162  . Type locality: Ega. Type depository: BMNH HT. Distribution: Brazil ( Amazonas , São Paulo)  . Note: distribution within Brazil (Asenjo et al. 2013).

* Belonuchus infimus ( Sharp 1885)  , comb. nov.

Philonthus infimus Sharp 1885: 414  . Type locality: Guatemala, Las Mercedes 3000 ft, Chiacam; Panama, Volcan de Chiriqui. Type depository: BMNH 8 ST: 5 Las Mercedes, 2 Chiriqui, 1 Chiacam [ MCPAbout MCP seen]. Distribution: Guatemala, Panama.

Remarks. Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on more than two specimens, including male/s and female/s. Eight specimens were located in the BMNH under this name. The specimens below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated (BMNH): card-mounted (with additional label “Sp. figured”), “ Philonthus infimus Las Mercedes  3000 ft. Guatemala Champion. [in Sharp’s hand] // Las Mercedes 3000 ft. Champion. (white label) // B. C. A. Col. I. 2. Philonthus infimus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 2 ♂, 2 ♀, card-mounted (one ♀ with additional “Type” round white label with red margin and “Type D.S” on the mounting card [in Sharp’s hand]), “ Philonthus infimus Las Mercedes  3000 ft. Guat. Champion. [in Sharp’s hand] // Las Mercedes 3000 ft. Champion. (white label) // B. C. A. Col. I. 2. Philonthus infimus, Sharp.  (white label)”; 2 ♀, card-mounted, “ Philonthus infimus  D.S. [in Sharp’s hand] // V. de Chiriqui 4–3000 ft Champion (white label) // B. C. A. Col. I. 2. Philonthus infimus, Sharp.  (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus iridescens Solsky 1868 

Belonuchus iridescens Solsky 1868: 140  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]  . Type depository: ZINAbout ZIN? ( Solsky ) #? ST. Distribution: Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Belonuchus irregularis Bernhauer 1916 

Belonuchus irregularis Bernhauer 1916c: 275  . Type locality: West-Columbien: Pacho, 2000m, Ost-Cordilleren, West-Columbien. Type depository: FMNHAbout FMNH HT. Distribution : Colombia. Note: generic affiliation unclear. 

* Belonuchus iteratus ( Sharp 1887)  , comb. nov.

Philonthus iteratus Sharp 1887: 788  . Type locality: Durango (Villa Lerdo). Type depository: BMNH 23 ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Remarks. Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on several specimens, including male/s and female/s. Twenty-three specimens were located in the BMNH under this name.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus jalappensis Bernhauer 1917 

Belonuchus jalappensis Bernhauer 1917d: 224  . Type locality: Mexico: Jalappa [Xalapa, Veracruz]  . Type depository: FMNH 2View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Belonuchus laevipennis Bernhauer 1942 

Belonuchus laevipennis Bernhauer 1942: 25  . Type locality: Cartago ( Pino ), 1000m [ Costa Rica]  . Type depository: FMNH 3View Materials ST. Distribution : Costa Rica. Note: generic affiliation unclear. 

* Belonuchus latecinctus ( Sharp 1885)  , comb. nov. ( Figs. 9–10View FIGURES 1–12)

Philonthus latecinctus Sharp 1885: 415  . Type locality: Guatemala, Cerro Zunil 4000–5000 ft, San Gerónimo. Type depository: BMNH 3 ST: 2 Cerro Zunil , 1 San Gerónimo [ MCPAbout MCP seen]. Distribution: Guatemala  .

Remarks. Sharp based his description on three specimens which were located in the BMNH under this name. The specimens below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated (BMNH, Figs. 9–10View FIGURES 1–12): card-mounted, “ Philonthus latecinctus  Type D.S. Cerro Zunil 4–6000 ft. Guatem. Champion. [in Sharp’s hand] // Type (round white label with red margin) // Cerro Zunil, Guatemala. Champion. (white label) // B. C. A. Col. I. 2. Philonthus latecinctus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 1 ♂ with same locality and collection data as lectotype and additional printed white label “Sp. Figured”; 1 ♀, cardmounted, “ Philonthus latecinctus  D.S. San Gerónimo Champion. [in Sharp’s hand] //San Gerónimo, Guatemala. Champion. (white label) // B. C. A. Col. I. 2. Philonthus latecinctus, Sharp.  (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus latro Erichson 1840 

Belonuchus latro Erichson 1840: 422  . Type locality: [no specific locality]. Type depository: ZMHB 1 ST ♀ [ MCPAbout MCP seen]. Distribution : Brazil (São Paulo)  ; South America? Note: distribution within (and including) Brazil (Asenjo et al. 2013). 

Belonuchus longiceps Bernhauer 1906 

Belonuchus longiceps Bernhauer 1906: 332  . Type locality: Peru. Type depository: FMNHAbout FMNH HT. Distribution: Peru, Bolivia. Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.

* Belonuchus lucilius ( Sharp 1885)  , comb. nov.

Philonthus lucilius Sharp 1885: 415  . Type locality: Panama, Volcan de Chiriqui 2000–3000 ft  . Type depository: BMNH HT. Distribution : Panama  .

Remarks. The “unique” specimen in BMNH is confirmed as the holotype.

Type material examined. Holotype ♂ ( BMNH): card-mounted, “ Philonthus lucilius  Type D.S. V. de Chiriqui 2– 3000 ft  . Champion. [in Sharp’s hand] // Type (round white label with red margin) // V. de Chiriqui 2–3000 ft  . Champion. (white label) // B. C. A. Col. I. 2. Philonthus lucilius, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

Belonuchus magnificus Bernhauer 1916 

Belonuchus magnificus Bernhauer 1916c: 277  . Type locality: West-Kolumbien: Umgebung von Cali (S. Antonio, 2000m). Type depository: FMNH 3View Materials ST. Distribution : Colombia. Note: generic affiliation unclear. 

Belonuchus meridionalis Bernhauer 1916 

Belonuchus meridionalis Bernhauer 1916c: 269  . Type locality: Argentinien [illeg. locality] & Bolivia. Type depository: FMNHAbout FMNH 2 ST, ZMUHAbout ZMUH 1+ST. Distribution: Argentina, Bolivia. Note: generic affiliation unclear.

Belonuchus mexicanus Solsky 1864 

Belonuchus mexicanus Solsky 1864: 440  . Type locality: Mexique. Type depository: ZINAbout ZIN? (Solsky) HT. Distribution: Mexico, Brazil, Argentina. Notes: redescription ( Silvestri 1945); distribution within Mexico ( Navarrete-Heredia et al. 2002); distribution within Brazil (Asenjo et al. 2013).

= Belonuchus mexicanus var. femoralis Solsky 1864: 440  . Type locality: Mexique. Type depository: ZIN? (Solsky) 2 ST. Distribution: Mexico. Notes: not in Bernhauer & Schubert (1914), Blackwelder (1944), Herman (2001d), but as synonym of B. mexicanus in Navarrete-Heredia et al. (2002: 335)  .

Belonuchus militaris Bernhauer 1921 

Belonuchus militaris Bernhauer 1921c: 170  . Type locality: Sao Paulo, Brazil  . Type depository: FMNH 1View Materials ST. Distribution : Brazil (São Paulo)  . Note: distribution within Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.

Belonuchus minor Bruch 1915  , nomen nudum

Belonuchus minor Bruch 1915: 504  , attributed to Bernhauer in litteris; nomen nudum. Locality: Misiones. Depository: MLPA (MLP 180). Distribution: Argentina. Note: see Fernández et al. (2007: 38).

Belonuchus miripennis Bernhauer 1916 

Belonuchus (Musicoderus) miripennis Bernhauer 1916c: 267  . Type locality: Cañon del Mte. Tolima, 1700m, Umgebung von Cali am Cauca, Columbien. Type depository: FMNH 3View Materials ST. Distribution : Colombia  .

Belonuchus modestus Fauvel 1891 

Belonuchus modestus Fauvel 1891: 118  . Type locality: Colonie Tovar [ Venezuela]. Type depository: IRSNBAbout IRSNB (Fauvel) 3 ST [ MCPAbout MCP seen]. Distribution: Venezuela, Colombia. Note: generic affiliation unclear.

* Belonuchus montanellus Bernhauer 1916 

Belonuchus montanellus Bernhauer 1916c: 272  . Type localities: Cañon del Mte. Tolima, 1700m; Alto de las cruces, 2200m (Umgebung von Cali am Cauca). Type depository: FMNH 4View Materials ST. Distribution : Colombia  .

Belonuchus montanicus Blackwelder 1944 

Belonuchus montanicus Blackwelder 1944: 137  (new name for Belonuchus montanus Bernhauer 1916: 273  ).

= Belonuchus montanus Bernhauer 1916c: 273  (preoccupied, junior primary homonym of Belonuchus montanus Fauvel 1895: 267  ). Type localities: West-Columbien: Cañon del Mte. Tolima, 1700m; Rio Vitaco, 2200m; Villa Elvira, 1800m. Type depository: FMNH 3 ST. Distribution: Colombia.

Belonuchus monticola Bernhauer 1917 

Belonuchus monticola Bernhauer 1917b: 18  . Type locality: Rio Vitaco , 2200m, Umgebung von Cali am Rio Cauca, West-Columbien  . Type depository: FMNHAbout FMNH HT. Distribution : Colombia 

* Belonuchus mordens Erichson 1840 

Belonuchus mordens Erichson 1840: 422  . Type locality: Brasilia. Type depository: ZMHB 1 ST ♂ [ MCPAbout MCP seen]. Distribution: Peru, Brazil, Argentina, Bolivia. Notes: frequent guest of Melipona  nests in Brazil ( Wasmann 1904: 482, Kistner 1982); distribution, including Peru and Bolivia ( Chani-Posse 2011b), within Brazil (Asenjo et al. 2013).

Belonuchus moritzi Bernhauer 1906 

Belonuchus moritzi Bernhauer 1906: 331  . Type locality: Venezuela  . Type depository: FMNH 2View Materials ST. Distribution : Venezuel. Note: generic affiliation unclear. 

Belonuchus multipunctatus ( Coiffait 1981) 

Endeius multipunctatus Coiffait 1981: 297  . Type locality: Galapagos: I. Santa Cruz versant S.E., basse altitude. Type depository: IRSNB HT ♂; IRSNB, MNHN 258 PT. Distribution: Galapagos Is. (Santa Cruz). Notes: comb. nov. ex Endeius (Chani Posse de Maus 2008: 386)  ; redescription ( Chani-Posse 2011b: 222); distribution within Galápagos Is. ( Newton & Peck 2006).

Belonuchus mundus Erichson 1840 

Belonuchus mundus Erichson 1840: 425  . Type locality: ins. St. Vincentii. Type depository: ZMHB? #?ST. Distribution: Lesser Antilles ( St. Lucia, St. Vincent, Grenada), Brazil. Notes: distribution within Lesser Antilles ( St. Vincent) ( Peck 2010); within (and including) Brazil (Asenjo et al. 2013).

* Belonuchus muticus (Sharp 1876)  , comb. nov. ( Figs. 11–12View FIGURES 1–12)

Philonthus muticus Sharp 1876a: 167  . Type locality: Ega. Type depository: BMNH 4 ST: 3 Ega , 1 Sao Paulo [ MCPAbout MCP seen]. Distribution: Brazil (Amazonas)  . Note: distribution within Brazil (Asenjo et al. 2013).

Remarks. Sharp explicitly mentioned four specimens of both sexes as part of the type series. The specimens listed below agree with the original description by Sharp (1876a) and is considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 11–12View FIGURES 1–12): card-mounted, “ ♂ // Type (round white label with red margin) // Ega (round green label) [in Sharp’s hand]// S America Brazil // Philonthus muticus  Type D.S. [in Sharp’s hand] // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 2 ♀ (one without head), card mounted, same locality and collection data as lectotype but with “ SYNTYPE ” (rounded white label with blue margin).

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus nigerrimus Bernhauer 1918 

Belonuchus nigerrimus Bernhauer 1918: 225  . Type locality: Mexico: Jalappa [Xalapa, Veracruz]  . Type depository: FMNH 2View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Belonuchus oakleyi Blackwelder, 1943 

Belonuchus oakleyi Blackwelder 1943: 422  . Type locality: Puerto Rico: Mayagüez  . Type depository: USNMAbout USNM HT. Distribution : Puerto Rico  .

* Belonuchus optatus ( Sharp, 1885)  , comb. nov. ( Figs. 13–14View FIGURES 13–24)

Philonthus optatus Sharp 1885: 427  . Type locality: Mexico  : Guanajuato  . Type depository: BMNH HT ♀. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Remarks. The “unique” specimen in BMNH is confirmed as the holotype.

Type material examined. Holotype ♀ (BMNH, Figs. 13–14View FIGURES 13–24): card-mounted, “ Philonthus optatus  Type D.S. Guanajuato. Mexico. Sallé. [in Sharp’s hand] // Type (round white label with red margin) //Guanajuato (white label) // Mexico. Salle Coll. (white label) // B. C. A. Col. I. 2. Philonthus optatus, Sharp.  (white label) // Belonuchus formosus  apud Sallé [in Sallé’s hand]”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus oxyporinus ( Sharp 1885) 

Philonthus oxyporinus Sharp 1885: 419  . Type locality: Mexico: Oaxaca. Type depository: BMNH HT ♀. Distribution: Mexico. Notes: to Belonuchus  (after Smetana 1995); distribution, biology (Navarrete-Heredia 1995, Jiménez-Sánchez et al. 2000); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Belonuchus palustris Bernhauer 1916 

Belonuchus (Musicoderus) palustris Bernhauer 1916c: 268  . Type locality: Alto de las Cruces, 2200m. Type depository: FMNHAbout FMNH HT. Distribution : Colombia  .

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus panamensis Sharp 1885 

Belonuchus panamensis Sharp 1885: 434  . Type localities: Nicaragua, Chontales; Panama, Volcan de Chiriqui, Bugaba, Tolé. Type depository: BMNH 11 ST: from all type localities [ MCPAbout MCP seen]. Distribution: Nicaragua, Costa Rica, Panama.

Belonuchus paradoxipunctatus Bernhauer 1921 

Belonuchus paradoxipunctatus Bernhauer 1921c: 171  . Type locality: San Roque, Corrientes [ Argentina]. Type depository: FMNHAbout FMNH 1 ST. Distribution: Argentina, Brazil. Note: distribution within (and including) Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.

Belonuchus paradoxus Bernhauer 1906 

Belonuchus paradoxus Bernhauer 1906: 333  . Type locality: Pachitea , Peru  . Type depository: FMNHAbout FMNH HT. Distribution : Peru  . Note: Peru ( Asenjo 2004).

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus pectinipes Bernhauer 1910 

Belonuchus pectinipes Bernhauer 1910: 379  . Type locality: Cordoba, Veracruz  . Type depository: FMNH 4View Materials ST. Distribution : Mexico  . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Belonuchus peruvianus Plavilstschikov 1929  , nomen nudum

Belonuchus peruvianus Plavilstschikov 1929: 30  , as "n. sp. (Bernhauer in litteris)"; nomen nudum, no description. Distribution: Peru. Note: as valid sp. in Blackwelder (1944) and Asenjo (2004); not in Herman (2001d).

* Belonuchus piceolus Bernhauer 1935 

Belonuchus piceolus Bernhauer 1935: 90  . Type locality: Brasilien  . Type depository: BMNH #?ST; FMNH 1View Materials ST. Distribution : Brazil  . Note: distribution within Brazil unknown (Asenjo et al. 2013).

Belonuchus pictipennis Sharp 1885 

Belonuchus pictipennis Sharp 1885: 436  . Type locality: Bugaba , Panama  . Type depository: BMNH 2 ST ♂. Distribution : Panama  .

Comment. It may belong to Paederomimus Sharp  (Sharp: 436, “very closely allied to B. mimeticus  ”) but see Paederomimus mimeticus (Sharp)  .

Belonuchus planiceps Erichson 1840 

Belonuchus planiceps Erichson 1840: 425  . Type locality: Columbia. Type depository: ZMHB 4 ST [ MCPAbout MCP seen]. Distribution: Costa Rica, Colombia, Venezuela.

Comment. It probably belongs to Holisus Erichson  ( Hyptiomina  ).

* Belonuchus platypterus ( Sharp 1885)  , comb. nov. ( Figs. 15–16View FIGURES 13–24)

Philonthus platypterus Sharp 1885: 412  . Type locality: Guatemala, Cerro Zunil 4 000  5000 feet. Type depository: BMNH 3 ST. Distribution : Guatemala  .

Remarks. Sharp explicitly mentioned three specimens (two males and one female) as part of the type series. The specimens listed below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 15–16View FIGURES 13–24): card-mounted, “ ♂ (in ink) Philonthus platypterus  Type D.S. Cerro Zunil. 4–5000 ft. [in Sharp’s hand] // Type (round white label with red margin) // Cerro Zunil. Guatemala. Champion. // B. C. A. Col. I. 2. Philonthus platypterus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 1 ♂, 1 ♀, same locality and collection data as lectotype but male with additional printed white label “Sp. figured”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus pollens Sharp 1885 

Belonuchus pollens Sharp 1885: 431  . Type localities: North America; Mexico: Oaxaca; Guatemala: near the city; Honduras. Type depository: BMNH 8 ST: from all type localities except North America [ MCPAbout MCP seen]; FMNHAbout FMNH 2 ST. Distribution: Mexico, Guatemala, Honduras ["North America" in error]. Notes: not in USA or Canada ( Smetana 1995: 733); distribution within Mexico ( Navarrete-Heredia et al. 2002).

Belonuchus posticalis Laporte de Castelnau 1840 

Staphylinus posticalis Laporte de Castelnau 1840: 176  . Type locality: Brésil  . Type depository: ?? #? ST. Distribution : Brazil  . Note: comb. nov. ex Staphylinus ( Bernhauer & Schubert 1914: 371)  ; distribution within Brazil unknown (Asenjo et al. 2013).

Belonuchus prasinipennis Bernhauer 1917 

Belonuchus prasinipennis Bernhauer 1917b: 18  (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien  . Type depository: FMNH 1View Materials ST; ZMUH 1View Materials ST. Distribution : Bolivia  .

Belonuchus pulchripennis Bernhauer 1908 

Belonuchus pulchripennis Bernhauer 1908: 330  . Type locality: Montana de Nancho, 7000', central Peru  . Type depository: FMNH 2View Materials ST; ?? 1 ST. Distribution : Peru  . Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.

Belonuchus purpuripennis Bernhauer 1916 

Belonuchus purpuripennis Bernhauer 1916c: 278  . Type locality: West-Columbien: Umgebung Cali am Rio Cauca, St. Antonio, 2000 m. Type depository: FMNHAbout FMNH 2 ST. Distribution: Colombia, Bolivia. Note: generic affiliation unclear.

Belonuchus richteri (Bernhauer, 1912) 

Philonthus (Chroaptomus) richteri Bernhauer 1912a: 44  . Type locality: Tucuman prov., Argentina. Type depository: FMNH HT ♀. Distribution: Argentina. Notes: comb. nov. ex Chroaptomus ( Chani-Posse 2006: 365)  ; redescription ( Chani-Posse 2011b: 223).

Comment. Not Belonuchus  , to be placed in a further study.

* Belonuchus rufipennis ( Fabricius 1801) 

Staphylinus rufipennis Fabricius 1801: 596  . Type locality: America meridionali. Type depository: ZMUK 3 ST (until recently at ZMUCAbout ZMUC, Zimsen 1964), not seen by Smetana (1995). Distribution: Canada, USA, Mexico, Guatemala, Belize, El Salvador, Nicaragua, Panama, Colombia, Venezuela, French Guiana, Trinidad, Tobago, Jamaica, Peru, Brazil, Argentina, Galapagos Is. (Isabela, S. Crist., S. Cruz), Hawaii; introduced for biocontrol in Italy but not established. Notes: as synonym of B. formosus ( Erichson 1840: 426)  ; introduction for biocontrol, biology, immature stages ( Silvestri 1945); redescription and distribution ( Smetana 1995: 726); in Atta mexicana  external debris (occasional) ( Navarrete-Heredia 2001); distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006); distribution within Galapagos Is. ( Newton & Peck 2006); distribution within Brazil (Asenjo et al. 2013); in Heliconia  bracts ( Frank & Barrera 2010); distribution in carcasses in Argentina ( Aballay et al. 2014).

= Staphylinus formosus Gravenhorst 1806: 72  . Type depository: ZMHB LT ♀, 1 PLT. Type locality: Carolina, USA (" Carolina "). Note: as synonym of B. rufipennis  ( Fauvel 1877: xxvii). Note: lectotype designation ( Smetana 1995: 728).

= Belonuchus pallipes Melsheimer 1844: 35  . Type depository: MCZ? #?ST. Type locality: Pennsylvania. Distribution: USA. Note: as synonym of B. rufipennis  ( Fauvel 1877: xxvii); no specimens under this name in the Melsheimer collection at the MCZ ( Smetana 1995: 728).

= Cafius apicalis Dejean 1833: 63  , unavailable, published in synonymy of unavailable name ( ICZN 1999: Art. 11.6). Distribution: Jamaica. Note: as synonym of B. formosus ( Erichson 1840: 426)  .

* Belonuchus rufiventris ( Sharp 1887)  , comb. nov. ( Figs. 17–18View FIGURES 13–24)

Philonthus rufiventris Sharp 1887: 787  . Type locality: Mexico: "Amecameca in Morelos " (=Est. Mexico?). Type depository: BMNH HT ♀  . Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).

Remarks. The unique female specimen in BMNH is confirmed as the holotype.

Type material examined. Holotype ♀ (BMNH, Figs. 17–18View FIGURES 13–24): card-mounted, “ Philonthus rufiventris  Type D.S. Amecameca. Hoge [in Sharp’s hand] // Type (round white label with red margin) // Amecameca, Morelos. Höge. // B. C. A. Col. I. 2. Philonthus rufiventris, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus rufocaudus ( Sharp 1885)  , comb. nov. ( Figs. 19–20View FIGURES 13–24)

Philonthus rufocaudus Sharp 1885: 412  . Type locality: Panama, Bugaba , Volcan in Chiriqui  . Type depository: BMNH 8 ST: 5 Bugaba , 3 Chiriqui [ MCPAbout MCP seen]. Distribution: Panama  .

Remarks. Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on at least one male and one female. Eight specimens were located in the BMNH under this name. The specimens below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 19–20View FIGURES 13–24): card-mounted, “ ♂ (in ink) Philonthus rufocaudus  Type D.S. Panama. Bugaba. Champion [in Sharp’s hand] // Type (round white label with red margin) // Bugaba  , Panama. Champion. // B. C. A. Col. I. 2. Philonthus rufocaudus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 1 ♂, 3 ♀, card-mounted, with same locality and collection data as lectotype  ; 3 ♀, card-mounted, “V. de Chriqui 25–4,000 ft. Champion (white label) // B. C. A. Col. I. 2. Philonthus rufocaudus, Sharp.  (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

* Belonuchus rufopygus ( Sharp 1885)  , comb. nov. ( Figs. 21–22View FIGURES 13–24)

Philonthus rufopygus Sharp 1885: 411  . Type locality: Panama: Bugaba. Type depository: BMNH 8 ST. Distribution: Panama [ Mexico in error]. Note: reported from Mexico ( Zaragoza Caballero 1976, but based on misidentified specimens [J.L. Navarrete-Heredia, unpublished]).

Remarks. Sharp explicitly mentioned several specimens as part of the type series. The specimens listed below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.

Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 21–22View FIGURES 13–24): card-mounted, “ ♂ (in ink) Philonthus  rufo-pygus Type D.S. Panama. Bugaba. Champion [in Sharp’s hand] // Type (round white label with red margin) // Bugaba  , Panama. Champion. // B. C. A. Col. I. 2. Philonthus rufopygus, Sharp.  (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 7 ♀ (2 of them mounted together), cardmounted, with same locality and collection data as lectotype  .

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

Belonuchus rugicollis Bernhauer 1916 

Belonuchus rugicollis Bernhauer 1916c: 276  . Type locality: Peru  . Type depository: FMNHAbout FMNH HT Distribution: Peru  . Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.

* Belonuchus satyrus Erichson 1840 

Belonuchus satyrus Erichson 1840: 421  . Type locality: Columbia. Type depository: ZMHB? #?ST. Distribution: Panama, Colombia, Venezuela, Trinidad; Costa Rica. Notes: biology, in Heliconia  bracts ( Frank & Barrera 2010).

* Belonuchus serraticornis (Sharp 1876)  , comb. nov. ( Figs. 23–24View FIGURES 13–24)

Philonthus serraticornis Sharp 1876a: 179  . Type locality: Ega. Type depository: BMNH HT. Distribution : Brazil (Amazonas)  . Note: distribution within Brazil (Asenjo et al. 2013).

Remarks. The “unique” specimen in BMNH is confirmed as the holotype.

Type material examined. Holotype ♂ (incomplete genitalia) ( BMNH, Figs. 23–24View FIGURES 13–24): card-mounted, “ ♂ // Type (round white label with red margin) // Ega (round green label) [in Sharp’s hand]// S America Brazil // Philonthus serraticornis  Type D.S. [in Sharp’s hand] // Sharp Coll. 1905.-313. (white label)”.

Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018)  .

Belonuchus setiger Sharp 1876 

Belonuchus setiger Sharp 1876a: 164  . Type locality: Ega. Type depository: BMNH 2 ST: 2 ♂ [ MCPAbout MCP seen]. Distribution : Brazil (Amazonas)  . Note: distribution within Brazil (Asenjo et al. 2013).

* Belonuchus sharpi Bernhauer 1907 

Belonuchus sharpi Bernhauer 1907: 285  (replacement name for Belonuchus grandiceps Sharp 1876a: 158  ). Distribution: Brazil (Amazonas, Pará), Peru. Note: distribution within Brazil (Asenjo et al. 2013).

= Belonuchus grandiceps Sharp 1876a: 158  (preoccupied, junior secondary homonym in Belonuchus  of Philonthus grandiceps Kraatz 1859a: 95  ). Type locality: Ega, Tapajos & St. Paulo [Amazonas]. Type depository: BMNH 5 ST. Note: Peru ( Asenjo 2004).

Belonuchus silvestris Bernhauer 1917 

Belonuchus silvestris Bernhauer 1917b: 17  . Type locality: Alto de las Cruces, 2200m, Cali am Rio Cauca, West- Columbien. Type depository: FMNHAbout FMNH HT. Distribution : Colombia  .

Belonuchus similis Sharp 1885 

Belonuchus similis Sharp 1885: 436  . Type locality: Panama, Volcan de Chiriqui 2500–6000 ft  . Type depository: BMNH 3 ST; FMNH 1View Materials ST. Distribution : Panama. Note: generic affiliation unclear. 

Belonuchus simillimus Bernhauer 1917 

Belonuchus simillimus Bernhauer 1917c: 106  . Type locality: Villa Elvira 1800m, Westkolumbien. Type depository: FMNHAbout FMNH HT. Distribution : Colombia  .

Belonuchus skalitskyi Bernhauer 1906 

Belonuchus skalitskyi Bernhauer 1906: 334  . Type locality: Pumananea, Peru. Type depository: FMNHAbout FMNH HT. Distribution: Peru, Bolivia. Note: Peru ( Asenjo 2004).

Comment. Not Belonuchus  , to be placed in a further study.

Belonuchus spinicornis ( Champion 1913) 

Musicoderus spinicornis Champion 1913: 4  . Type locality: Costa Rica, Pitahaya , 1400m  . Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [ MCPAbout MCP seen]. Distribution : Costa Rica  .

* Belonuchus stenoderus Sharp 1885 

Belonuchus stenoderus Sharp 1885: 434  . Type locality: Panama, Bugaba. Type depository: BMNH 4 ST; FMNHAbout FMNH 1 ST. Distribution: Costa Rica, Panama. Note: the syntype in FMNHAbout FMNH does not belong to the ‘ Belonuchus haemorrhoidalis  clade’ ( Chani-Posse et al. 2018).

Belonuchus subaeneipennis Bernhauer 1916 

Belonuchus (Musicoderus) subaeneipennis Bernhauer 1916c: 269  . Type locality: West-Columbien: Umgebung von Cali am Rio Cauca (Cañon del Mte Tolima, 1700m). Type depository: FMNH 1View Materials ST;? 1 ST. Distribution : Colombia  .

Belonuchus subaeneus Bernhauer 1916 

Belonuchus subaeneus Bernhauer 1916c: 268  (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien  . Type depository: FMNH 1View Materials ST; ZMUH 1View Materials +ST?. Distribution : Bolivia  .

Belonuchus subsellatus Bernhauer 1916 

Belonuchus subsellatus Bernhauer 1916c: 276  (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien  . Type depository: FMNH 1View Materials ST, ZMUH 1View Materials + ST. Distribution : Bolivia. Note: generic affiliation unclear. 

Belonuchus superbus Bernhauer 1916 

Belonuchus superbus Bernhauer 1916c: 273  . Type localities: Alto de las cruces, 2200m  ; Rio Vitaco , 2000m  ; Rio Aquatal , 1800m  ; S. Antonio, 2000m; Villa Elvira, 1800m. Type depository: FMNH 3View Materials ST;? 2 ST. Distribution : Colombia. Note: generic affiliation unclear. 

* Belonuchus supernus (Herman 2001)  , comb. nov. ( Figs. 47–48View FIGURES 45–52)

Philonthus supernus Herman 2001a: 48  (new name for Philonthus excelsus Bernhauer 1941: 287  ). Distribution: Peru.

= Philonthus excelsus Bernhauer 1941: 287  , also 1951: 277 (preoccupied, junior secondary homonym in Philonthus  of Hesperus excelsus Cameron 1931: 361  , and junior primary homonym of Philonthus excelsus Cameron 1932: 147  ). Type locality: Ocaña, 3100m, etc., s Peru, Oberh. Paleo 3240m, S Peru. Type depository: FMNHAbout FMNH 21 ST: 1 Ocaña. Note: Peru ( Asenjo 2004).

Remarks. Although references about three different type localities on different dates and a male character are given in the original description, there is no indication about how many specimens were examined. Two specimens were located in the FMNHAbout FMNH under this name. The specimen below agrees with the original description and considered to have been a syntype.

Type material examined. Syntype ♂ (FMNH, Figs. 47–48View FIGURES 45–52): card mounted with cutouts at head-thorax and abdomen // Süd – Peru Oberh. Paleo, 3240m 10.4.1936 [in Bernhauer’s hand] (green label) // Hbg. – Süd – Peru Sammelreise 1936. Eing. Nr. 1, 1937 (green label) // 219 (white label) // Philonthus excelsus Brh. Cotypus  (white label) // excelsus Benh.  Typus Philonthus  [in Bernhauer’s hand](red label) // Chicago NHMus M.Bernhauer Collection (white label)”.

* Belonuchus taczanovskii ( Solsky 1872) 

Philonthus taczanovskii Solsky 1872: 303  . Type locality: Monte Rico, Perou. Type depository: ZINAbout ZIN? (Solsky) #?ST. Distribution: Peru, Ecuador. Note: Peru ( Asenjo 2004).

Belonuchus terminalis (Laporte de Castelnau 1840) 

Staphylinus terminalis Laporte de Castelnau 1840: 176  . Type locality: Brésil. Type depository:? #? ST. Distribution: Brazil. Notes: comb. nov. ex Staphylinus ( Bernhauer & Schubert 1914: 371)  ; preoccupied, junior primary homonym of Staphylinus terminalis Erichson 1839: 396  ; conserved by Opinion 2053 ( ICZN 2003); distribution within Brazil unknown (Asenjo et al. 2013).

* Belonuchus tremolerasi Bernhauer 1912 

Belonuchus tremolerasi Bernhauer 1912a: 40  . Type locality: Uruguay. Type depository: FMNHAbout FMNH 1 ST. Distribution: Uruguay, Argentina, Paraguay.

Belonuchus trinitatis Blackwelder 1943 

Belonuchus trinitatis Blackwelder 1943: 423  . Type locality: Trinidad: Department of Agriculture Stock Farm at St  . Joseph, 6 miles east of Port of Spain  . Type depository: USNMAbout USNM HT ♂, 4 PT; FMNH 1View Materials PT. Distribution : Trinidad  .

* Belonuchus trochanterinus ( Sharp 1885) 

Philonthus trochanterinus Sharp 1885: 417  . Type locality: Guatemala, Capetillo. Type depository: BMNH #? ST. Distribution: Mexico, Guatemala. Note: comb. nov. ex Philonthus  , distribution within Mexico (Navarrete- Heredia et al. 2002: 335).

Belonuchus tucumanensis Bernhauer 1934 

Belonuchus tucumanensis Bernhauer 1934b: 213  . Type locality: Tucumán. Type depository: FMNH 1View Materials HT. Distribution : Argentina  .

* Belonuchus variolosus Solsky 1870 

Belonuchus variolosus Solsky 1870: 263  . Type locality: " Cayenne " [ French Guiana]  . Type depository: ZINAbout ZIN? ( Solsky ) #? ST. Distribution : French Guiana (Cayenne). Note: generic affiliation unclear. 

= Belonuchus variolaris  ; Solsky 1872: 305, misspelling.

Belonuchus venezolanus Bernhauer 1917 

Belonuchus venezolanus Bernhauer 1917c: 104  . Type localities: Venezuela & Columbien. Type depository: FMNHAbout FMNH 3ST; NMWAbout NMW 1+ST?; ZMUHAbout ZMUH 1+ST?. Distribution: Venezuela, Colombia. Note: generic affiliation unclear.

Belonuchus viridipennis Baudi di Selve 1848 

Belonuchus viridipennis Baudi di Selve 1848: 125  . Type locality: Novae Valenciae in Columbia [ Venezuela, teste Herman 2001c: 1594]. Type depository: MRSNAbout MRSN / MCSNAbout MCSN? #?ST. Distribution: Mexico, Guatemala, Panama, Colombia, Venezuela. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002). Note: generic affiliation unclear (based on non-type material in FMNHAbout FMNH).

= Belonuchus dejectus Sharp 1885: 433  . Type localities: Mexico: Cordova, Jalapa, Las Vigas, Mirador; Guatemala, Capetillo, Cerro Zunil, Balheu, Tamahu, San Joaquin, Purula, El Tumbador, San Gerónimo; Panama: Volcan de Chiriqui 2500–4000 ft. Type depository: BMNH 14+ST. Distribution: Mexico, Guatemala, Panama. Note: as synonym of B. viridipennis ( Sharp 1887: 789)  .

Belonuchus vulneratus Bernhauer 1906 

Belonuchus vulneratus Bernhauer 1906: 335  . Type locality: Pachitea , Peru  . Type depository: FMNHAbout FMNH HT. Distribution : Peru  . Note: Peru ( Asenjo 2004). Note: not Belonuchus  , to be placed in a further study.

Belonuchus weiserianus (Bernhauer 1921) 

Philonthus weiserianus Bernhauer 1921c: 106  . Type locality: Hornadita, 3400m, Jujuy, Argentina. Type depository: FMNH LT ♀. Distribution: Argentina. Note: comb. nov. ex Philonthus  and lectotype designation ( Chani-Posse 2010: 8); redescription ( Chani-Posse 2011b: 224).

Belonuchus xantholinoides Bernhauer 1917 

Belonuchus xantholinoides Bernhauer 1917c: 106  . Type localities: Villa Elvira 1800m & S. Antonio 2200m, Westkolumbien. Type depository: FMNH 2View Materials ST. Distribution : Colombia  .

* Belonuchus xanthomelas Solsky 1868 

Belonuchus xanthomelas Solsky 1868: 139  . Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #?ST. Distribution: USA (TX), Mexico. Note: generic placement (Navarrete- Heredia 1995); distribution, also within Mexico ( Navarrete-Heredia et al. 2002; Márquez-Luna 2001, Márquez 2006).

* Belonuchus xanthopterus ( Nordmann 1837) 

Philonthus xanthopterus Nordmann 1837: 99  . Type locality: Parae in America meridionali. Type depository: ZMHB 1 ST ♂ [ MCPAbout MCP seen]. Distribution: Brazil, Peru, Panama?, Ecuador?. Note: comb. nov. ex Philonthus ( Erichson 1840: 421)  ; Peru ( Soukup 1942: 177); distribution within Brazil (Asenjo et al. 2013).

* Belonuchus xanthopus Solsky 1868 

Belonuchus xanthopus Solsky 1868: 139  . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca]  . Type depository: ZINAbout ZIN? ( Solsky ) #? ST. Distribution: Mexico  . Notes: generic placement (Navarrete-Heredia 1995); distribution within Mexico ( Navarrete-Heredia et al. 2002).

* Belonuchus zunilensis ( Sharp 1885) 

Philonthus zunilensis Sharp 1885: 418  . Type locality: Guatemala: Cerro Zunil 4000–5000 ft. Type depository: BMNH 4 ST. Distribution: Mexico, Guatemala. Notes: comb. nov. ex Philonthus  , distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution ( Asiain et al. 2011).

MCP

Pontificia Universidade Catolica do Rio Grande do Sul

IRSNB

Institut Royal des Sciences Naturelles de Belgique

FMNH

Field Museum of Natural History

USNM

Smithsonian Institution, National Museum of Natural History

NHME

Natuurhistorisch Museum

NMW

Naturhistorisches Museum, Wien

CZUG

Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia

CNIN

Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico

MCZ

Museum of Comparative Zoology

CUIC

Cornell University Insect Collection

ZIN

Russian Academy of Sciences, Zoological Institute, Zoological Museum

ZMUC

Zoological Museum, University of Copenhagen

ZMUH

Zoologisches Institut und Zoologisches Museum, Universitat Hamburg

MRSN

Museo Regionale di Scienze Naturali

MCSN

Museo Civico di Storia Naturale "Giacomo Doria"

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Loc

Belonuchus Nordmann 1837

Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey 2018

2018
Loc

Belonuchus

Nordmann 1837 : 129

Loc

Musicoderus

Sharp 1885 : 455 Lucas 1920 : 425 Bernhauer 1916c: 266 Blackwelder 1943: 421

Loc

Philonthus abnormalis

Sharp 1885 : 397

Loc

Belonuchus aequalis

Sharp 1876a : 161

Loc

Belonuchus agilis

Erichson 1840 : 423

Loc

Belonuchus chevrolati

Fauvel 1863 : 432 Fauvel 1869: 489

Loc

Belonuchus albovariegatus

Bernhauer 1916c : 276

Loc

Linoderus navarretei López-García & Méndez-Rojas 2014 : 86

López-García & Méndez-Rojas 2014 : 86 Chani-Posse & Solodovnikov 2017: 90

Loc

Philonthus alternans

Sharp 1885 : 418 Navarrete-Heredia et al. 2002 : 334

Loc

Belonuchus alternatus

Sharp 1885 : 430 Bernhauer & Schubert (1914: 369) Sharp 1885 : 418

Loc

Belonuchus altissimus

Bernhauer 1916c : 275

Loc

Belonuchus altivagans

Bernhauer 1916c : 272

Loc

Philonthus aluticollis

Bernhauer 1921c : 107 Chani-Posse 2010: 8 Chani-Posse 2011b: 215

Loc

Belonuchus amplipennis

Fauvel 1891 : 117

Loc

Belonuchus amplus

Blackwelder 1943 : 425

Loc

Belonuchus andinus

Bernhauer 1916c : 278

Loc

Belonuchus (Musicoderus) angusticeps

Scheerpeltz 1960 : 117

Loc

Belonuchus angusticollis

Bernhauer 1906 : 330

Loc

Belonuchus antiguae

Blackwelder 1943 : 434

Loc

Belonuchus (Musicoderus) apicicornis

Bernhauer 1916c : 270

Loc

Philonthus apiciventris

Sharp 1885 : 419

Loc

Belonuchus argentinus

Bernhauer 1916c : 278 Bruch (1915: 503)

Loc

Belonuchus armatus

Sharp 1876a : 163

Loc

Belonuchus arrogans

Bernhauer 1916c : 274

Loc

Belonuchus asperatus

Bernhauer 1906 : 336 Scheerpeltz 1960 : 120

Loc

Plociopterus atticida

Wasmann 1925 : 71 Chatzimanolis & Chani-Posse 2016: 214

Loc

Hesperus eidmanni

Scheerpeltz 1936 : 484 Posse 2016 : 214 Chatzimanolis & Chani-Posse 2016: 214

Loc

Belonuchus avidus

Erichson 1840 : 423 Herman 2001d : 1594

Loc

Belonuchus (Musicoderus) barbicornis

Bernhauer 1916c : 271

Loc

Philonthus apiciventris

Sharp 1885 : 418

Loc

Belonuchus batesi

Sharp 1876a : 157

Loc

Belonuchus bidens

Sharp 1885 : 430

Loc

Belonuchus bipunctatus

Bernhauer 1935 : 90

Loc

Belonuchus boops

Sharp 1885 : 435

Loc

Belonuchus boxi

Bernhauer 1934b : 212

Loc

Belonuchus brasilianus

Bernhauer 1917b : 19

Loc

Belonuchus breviceps

Fauvel 1891 : 117

Loc

Belonuchus bruchi

Bernhauer 1912b : 175

Loc

Belonuchus bruchianus

Bernhauer 1911 : 413

Loc

Belonuchus bugnioni

Fauvel 1901 : 86

Loc

Belonuchus eximius

Bernhauer 1917c : 105 Blackwelder 1943: 428

Loc

Belonuchus cacao

Blackwelder 1943 : 434

Loc

Philonthus candens

Erichson 1840 : 460 Bernhauer & Schubert 1914: 369

Loc

Belonuchus cariniventris

Bernhauer 1918 : 225

Loc

Belonuchus castaneus

Chani-Posse 2006 : 366 Bernhauer 1922 : 15

Loc

Belonuchus richteri

Bernhauer 1922 : 15 Bernhauer 1912a : 44 Chani-Posse 2011b : 219

Loc

Belonuchus (Musicoderus) cavipennis

Bernhauer 1916c : 267

Loc

Philonthus celatus

Sharp 1885 : 420

Loc

Musicoderus cephalotes

Sharp 1885 : 455 Scheerpeltz 1933 : 1377

Loc

Belonuchus cifuentesi Rodríguez & Navarrete-Heredia 2016 : 445

Rodríguez & Navarrete-Heredia 2016 : 445

Loc

Belonuchus clypeatus

Sharp 1876a : 160

Loc

Belonuchus coelestinus

Bernhauer 1908 : 330

Loc

Belonuchus coeruleus

Cameron 1922 : 122 Blackwelder 1943: 426

Loc

Belonuchus smithi

Cameron 1922 : 122 Blackwelder, 1943: 426

Loc

Belonuchus (Musicoderus) coelestis

Bernhauer 1916c : 266

Loc

Belonuchus coeruleispennis

Motschulsky 1858 : 668 Motschulsky 1870 : 49 Bernhauer & Schubert 1914 : 369 Herman (2001d : 2522

Loc

Belonuchus cognatus

Sharp 1885 : 429

Loc

Philonthus colon

Sharp 1885 : 419 Navarrete-Heredia et al. 2002 : 334

Loc

Belonuchus (Musicoderus) columbicus

Bernhauer 1916c : 271

Loc

Staphylinus connexus

Say 1830 : 35 Erichson 1840: 927

Loc

Musicoderus convexus

Bernhauer 1908 : 331 Scheerpeltz 1933 : 1377

Loc

Belonuchus (Musicoderus) cordilleranus

Bernhauer 1916c : 270

Loc

Philonthus corticalis

Sharp 1885 : 413

Loc

Belonuchus cribriceps

Bernhauer 1921c : 172

Loc

Belonuchus danforthi

Blackwelder 1943 : 429

Loc

Belonuchus decipiens

Sharp 1876a : 159

Loc

Belonuchus dichrous

Erichson 1840 : 420

Loc

Belonuchus diversicornis

Bernhauer 1917c : 104

Loc

Belonuchus docilis

Sharp 1885 : 435

Loc

Belonuchus dominicus

Blackwelder 1943 : 431

Loc

Belonuchus duplicatus

Blackwelder 1944 : 137

Loc

Belonuchus bruchi

Bernhauer 1934b : 213

Loc

Belonuchus (Musicoderus) ecuadorensis

Bernhauer 1935 : 91

Loc

Staphylinus ephippiatus

Say 1830 : 35 Erichson 1840: 927 Smetana 1995: 746

Loc

Belonuchus simplex

Sharp 1885 : 432 Fauvel 1901: 86 Smetana 1995: 748

Loc

Belonuchus punctiventris

Casey 1915 : 446 Smetana 1995: 746 Smetana 1995: 748

Loc

Belonuchus laticeps

Casey 1915 : 446 Smetana 1995: 746 Smetana 1995: 749

Loc

Belonuchus jacobianus

Casey 1915 : 447 Smetana 1995: 746 Smetana 1995: 749

Loc

Belonuchus schaefferi

Cooper 1933 : 545 Smetana 1995: 746

Loc

Belonuchus erichsoni

Bernhauer 1917d : 223

Loc

Belonuchus erythropterus

Solsky 1868 : 137 Smetana 1995: 752

Loc

Belonuchus excelsus

Bernhauer 1917b : 19

Loc

Philonthus extremus

Sharp 1885 : 415

Loc

Belonuchus falsus

Bernhauer 1917c : 105

Loc

Belonuchus fassli

Bernhauer 1917a : 37

Loc

Philonthus flavicoxis

Bernhauer 1912a : 43 Chani-Posse 2010: 8 Chani-Posse 2011b: 221

Loc

Belonuchus flavipennis

Solsky 1870 : 263

Loc

Belonuchus flavocaudatus

Bernhauer 1921c : 170

Loc

Belonuchus (Musicoderus) forsteri

Scheerpeltz 1960 : 115

Loc

Belonuchus fossulatus

Wasmann 1887 : 414

Loc

Belonuchus frater

Bernhauer 1906 : 333

Loc

Belonuchus gagates

Erichson 1840 : 424

Loc

Belonuchus gowdeyi

Bernhauer 1935 : 91 Smetana 1995: 739 Blackwelder 1943: 424

Loc

Belonuchus minax

Erichson 1840 : 424 Blackwelder 1943: 424 Smetana (1995: 737)

Loc

Musicoderus nigrocaeruleus

Cameron 1913 : 175 Blackwelder 1943: 424 Smetana (1995: 738)

Loc

Belonuchus ganglbaueri

Bernhauer 1906 : 331

Loc

Philonthus godmani

Sharp 1885 : 413

Loc

Plociopterus goeldii

Wasmann 1925 : 72 Chatzimanolis & Chani-Posse 2016: 215

Loc

Musicoderus gracilis

Sharp 1885 : 456

Loc

Staphylinus haemorrhoidalis

Fabricius 1801 : 596 Nordmann 1837: 130 Gmelin 1790 : 2036 Herman 2001a: 42

Loc

Belonuchus hispaniolus

Blackwelder 1943 : 422

Loc

Belonuchus holisoides

Sharp1876a : 161

Loc

Belonuchus iconnicoffi

Plavilstschikov 1929 : 30

Loc

Philonthus ielskii

Solsky 1872 : 296 Bernhauer & Schubert 1914: 370 Herman 2001d : 2527 Asenjo 2004 : 61

Loc

Belonuchus impressifrons

Sharp 1876a : 162

Loc

Philonthus infimus

Sharp 1885 : 414

Loc

Belonuchus iridescens

Solsky 1868 : 140

Loc

Belonuchus irregularis

Bernhauer 1916c : 275

Loc

Philonthus iteratus

Sharp 1887 : 788

Loc

Belonuchus jalappensis

Bernhauer 1917d : 224

Loc

Belonuchus laevipennis

Bernhauer 1942 : 25

Loc

Philonthus latecinctus

Sharp 1885 : 415

Loc

Belonuchus latro

Erichson 1840 : 422

Loc

Belonuchus longiceps

Bernhauer 1906 : 332

Loc

Philonthus lucilius

Sharp 1885 : 415

Loc

Belonuchus magnificus

Bernhauer 1916c : 277

Loc

Belonuchus meridionalis

Bernhauer 1916c : 269

Loc

Belonuchus mexicanus

Solsky 1864 : 440

Loc

Belonuchus mexicanus var. femoralis

Solsky 1864 : 440 Navarrete-Heredia et al. (2002 : 335

Loc

Belonuchus militaris

Bernhauer 1921c : 170

Loc

Belonuchus minor

Bruch 1915 : 504 Fernández et al. (2007 : 38

Loc

Belonuchus (Musicoderus) miripennis

Bernhauer 1916c : 267

Loc

Belonuchus modestus

Fauvel 1891 : 118

Loc

Belonuchus montanellus

Bernhauer 1916c : 272

Loc

Belonuchus montanicus

Blackwelder 1944 : 137

Loc

Belonuchus montanus

Bernhauer 1916c : 273 Fauvel 1895 : 267

Loc

Belonuchus monticola

Bernhauer 1917b : 18

Loc

Belonuchus mordens

Erichson 1840 : 422 Wasmann 1904 : 482

Loc

Belonuchus moritzi

Bernhauer 1906 : 331

Loc

Endeius multipunctatus

Coiffait 1981 : 297 Maus 2008 : 386 Chani-Posse 2011b: 222

Loc

Belonuchus mundus

Erichson 1840 : 425

Loc

Philonthus muticus

Sharp 1876a : 167

Loc

Belonuchus nigerrimus

Bernhauer 1918 : 225

Loc

Belonuchus oakleyi

Blackwelder 1943 : 422

Loc

Philonthus optatus

Sharp 1885 : 427

Loc

Philonthus oxyporinus

Sharp 1885 : 419

Loc

Belonuchus (Musicoderus) palustris

Bernhauer 1916c : 268

Loc

Belonuchus panamensis

Sharp 1885 : 434

Loc

Belonuchus paradoxipunctatus

Bernhauer 1921c : 171

Loc

Belonuchus paradoxus

Bernhauer 1906 : 333

Loc

Belonuchus pectinipes

Bernhauer 1910 : 379

Loc

Belonuchus peruvianus

Plavilstschikov 1929 : 30

Loc

Belonuchus piceolus

Bernhauer 1935 : 90

Loc

Belonuchus pictipennis

Sharp 1885 : 436

Loc

Belonuchus planiceps

Erichson 1840 : 425

Loc

Philonthus platypterus

Sharp 1885 : 412

Loc

Belonuchus pollens

Sharp 1885 : 431 Smetana 1995: 733

Loc

Staphylinus posticalis Laporte de Castelnau 1840 : 176

Castelnau 1840 : 176 Bernhauer & Schubert 1914: 371

Loc

Belonuchus prasinipennis

Bernhauer 1917b : 18

Loc

Belonuchus pulchripennis

Bernhauer 1908 : 330

Loc

Belonuchus purpuripennis

Bernhauer 1916c : 278

Loc

Philonthus (Chroaptomus) richteri

Bernhauer 1912a : 44 Chani-Posse 2006: 365 Chani-Posse 2011b: 223

Loc

Staphylinus rufipennis

Fabricius 1801 : 596 Erichson 1840: 426 Smetana 1995: 726

Loc

Staphylinus formosus

Gravenhorst 1806 : 72 Smetana 1995: 728

Loc

Belonuchus pallipes

Melsheimer 1844 : 35 Smetana 1995: 728

Loc

Cafius apicalis

Dejean 1833 : 63 Erichson 1840: 426

Loc

Philonthus rufiventris

Sharp 1887 : 787

Loc

Philonthus rufocaudus

Sharp 1885 : 412

Loc

Philonthus rufopygus

Sharp 1885 : 411

Loc

Belonuchus rugicollis

Bernhauer 1916c : 276

Loc

Belonuchus satyrus

Erichson 1840 : 421

Loc

Philonthus serraticornis

Sharp 1876a : 179

Loc

Belonuchus setiger

Sharp 1876a : 164

Loc

Belonuchus sharpi

Bernhauer 1907 : 285 Sharp 1876a : 158

Loc

Belonuchus grandiceps

Sharp 1876a : 158 Kraatz 1859a : 95

Loc

Belonuchus silvestris

Bernhauer 1917b : 17

Loc

Belonuchus similis

Sharp 1885 : 436

Loc

Belonuchus simillimus

Bernhauer 1917c : 106

Loc

Belonuchus skalitskyi

Bernhauer 1906 : 334

Loc

Musicoderus spinicornis

Champion 1913 : 4

Loc

Belonuchus stenoderus

Sharp 1885 : 434

Loc

Belonuchus (Musicoderus) subaeneipennis

Bernhauer 1916c : 269

Loc

Belonuchus subaeneus

Bernhauer 1916c : 268

Loc

Belonuchus subsellatus

Bernhauer 1916c : 276

Loc

Belonuchus superbus

Bernhauer 1916c : 273

Loc

Philonthus supernus

Herman 2001a : 48 Bernhauer 1941 : 287

Loc

Philonthus excelsus

Bernhauer 1941 : 287 Cameron 1931 : 361 Cameron 1932 : 147

Loc

Philonthus taczanovskii

Solsky 1872 : 303

Loc

Staphylinus terminalis Laporte de Castelnau 1840 : 176

Castelnau 1840 : 176 Bernhauer & Schubert 1914: 371 Erichson 1839 : 396

Loc

Belonuchus tremolerasi

Bernhauer 1912a : 40

Loc

Belonuchus trinitatis

Blackwelder 1943 : 423

Loc

Philonthus trochanterinus

Sharp 1885 : 417

Loc

Belonuchus tucumanensis

Bernhauer 1934b : 213

Loc

Belonuchus variolosus

Solsky 1870 : 263

Loc

Belonuchus variolaris

Solsky 1872 : 305

Loc

Belonuchus venezolanus

Bernhauer 1917c : 104

Loc

Belonuchus viridipennis Baudi di Selve 1848 : 125

Selve 1848 : 125 Herman 2001c : 1594

Loc

Belonuchus dejectus

Sharp 1885 : 433 Sharp 1887: 789

Loc

Belonuchus vulneratus

Bernhauer 1906 : 335

Loc

Philonthus weiserianus

Bernhauer 1921c : 106 Chani-Posse 2010: 8 Chani-Posse 2011b: 224

Loc

Belonuchus xantholinoides

Bernhauer 1917c : 106

Loc

Belonuchus xanthomelas

Solsky 1868 : 139

Loc

Philonthus xanthopterus

Nordmann 1837 : 99 Erichson 1840: 421 Soukup 1942: 177

Loc

Belonuchus xanthopus

Solsky 1868 : 139

Loc

Philonthus zunilensis

Sharp 1885 : 418