Lysiteles

Tang, Guo, Yin, Chang-Ming, Peng, Xian-Jin, Ubick, Darrell & Griswold, Charles, 2008, The crab spiders of the genus Lysiteles from Yunnan Province, China (Araneae: Thomisidae), Zootaxa 1742, pp. 1-41 : 3-4

Key to species of the genus Lysiteles from Yunnan Province

1 Male ............................................................................................................................................................................. 2

- Female ........................................................................................................................................................................ 18

2 Palp with 2 tibial apophyses (VTA and RTA)............................................................................................................... 6

- Palp not as above mentioned......................................................................................................................................... 3

3 Palp with 3 tibial apophyses (VTA, RTA and ITA) ...................................................................................................... 4

- Palp with 1 tibial apophysis (RTA) ( Fig. 20 h ) ........................................................................ L. uniprocessus sp. nov.

4 Embolus slender, filiform .............................................................................................................................. L. dentatus

- Embolus not slender …................................................................................................................................................. 5

5 ITA transversal digitform, embolus tube-shaped ( Fig. 19 j ) ....................................................... L. transversus sp. nov.

- ITA blunt, embolus short screw-shaped ......................................................................................................... L. conicus

6 Ratio of width of RTA / tibia 0.40–0.50, RTA with some modifications...................................................................... 7

- Ratio of width of RTA / tibia 0.10–0.30, RTA without modifications ........................................................................ 10

7 RTA with some small denticles and some black hairs on the surface........................................................................... 8

- RTA with a basal process .............................................................................................................................................. 9

8 Ratio of width of embolus/bulb = 0.33, RTA with some black hairs ( Fig. 18 c ) ........................................... L. torsivus

- Ratio of width of embolus/bulb = 0.08, RTA with some denticles ( Fig. 8 e ) ................................ L. distortus sp. nov.

9 Ratio of width of embolus/bulb = 0.20, RTA spatulate, cymbium without process ( Fig. 17 j ). L. subdianicus sp. nov.

- Ratio of width of embolus/bulb = 0.08, RTA digitiform, cymbium with an anterior process ( Fig. 7 i ) ....... L. dianicus

10 Ratio of length of embolus/bulb = 0.90–1.20 ............................................................................................................ 11

- Ratio of length of embolus/bulb = 0.25–0.50 ............................................................................................................. 12

11 Shield of prosoma without markings ( Figs 15 a–d ), ratio of width of embolus / bulb = 0.12 ( Fig. 15 j ) ....... L. salticus

- Shield of prosoma with 2 pairs of black markings ( Figs 4 a–c ), ratio of width of embolus / bulb = 0.20 ( Fig. 4 j ) ...... ........................................................................................................................................................ L. conflatus sp. nov.

12 Bulb with a small ATA ( Fig. 3 j ) .................................................................................................. L. clavellatus sp. nov.

- Bulb without ATA ...................................................................................................................................................... 13

13 Embolus small nail-shaped ........................................................................................................................................ 14

- Embolus cone-shaped ................................................................................................................................................ 15

14 Opisthosoma dorsum yellow with 10 pairs of small black dots ( Figs 14 a–d ) ........................................... L. punctiger

- Opisthosoma dorsum light brown with 2 pairs of joint black stripes posteriorly ( Figs 11 a–b ) ........... L. kunmingensis

15 Body color light with typical markings, embolus twist basally ................................................................................. 16

- Body color dark, dorsum markings various greatly; embolus not as the aboved ..................................................... 17

16 VTA with 5 bristles in a row, RTA bifid apically ......................................................................................... L. ambrosii

- VTA without bristles, RTA curved apically, beak-shaped .......................................................................... L. bhutanus

17 Dorsal shield of prosoma black ( Figs 13 a–d ) .................................................................................................. L. niger

- Dorsal shield of prosoma gray, usually with stripes laterally ( Figs 12 a–d ) ..................................................... L. maius

18 Epigyne with atrium ................................................................................................................................................... 19

- Epigyne without artium .............................................................................................................................................. 26

19 Epigyne with median septum ..................................................................................................................................... 20

- Epigyne without median septum................................................................................................................................. 22

20 Ratio of width of copulatory openings / atrium = 0.2 ............................................................................................... 21

- Ratio of width of copulatory openings / atrium = 0.33, copulatory ducts arcuate curved ( Figs 1 d, e ) ......................... ......................................................................................................................................................... L. arcuatus sp. nov.

21 Prosoma yellow with 1 pair of brown stripes, dorsum with some small markings ( Figs 5 a–c ); copulatory ducts rugate ( Figs 5 e, g ) .................................................................................................................................... L. corrugus sp. nov.

- Prosoma blackish brown, dorsum with large markings ( Fig. 9 a ), copulatory ducts sinuous ( Figs 9 b–e ) .................. ................................................................................................................................................................ L. guoi sp. nov.

22 Ratio of width of atrium / spermatheca = 0.5–1 ......................................................................................................... 23

- Ratio of width of atrium / spermatheca = 2–3 ........................................................................................................... 24

23 Ratio of length of copulatory ducts / spermatheca = 1 ( Figs 6 f, g ) ............................................... L. curvatus sp. nov.

- Ratio of length of copulatory ducts / spermatheca = 0.5 ( Figs 12 k, l ) ............................................................. L. maius

24 Atrium with anterior rim ( Figs 19 h , k ) ..................................................................................... L. transversus sp. nov.

- Atrium without anterior rim........................................................................................................................................ 25

25 Copalatory ducts twist helix-shaped ( Figs 7 j, k ) ......................................................................................... L. dianicus

- Copalatory ducts curved simply ( Figs 17 l ) ............................................................................... L. subdianicus sp. nov.

26 Epigyne with transversal sclerotized plate ................................................................................................................. 32

- Epigyne without transversal sclerotized plate............................................................................................................. 27

27 Epigyne with hood or median septum ........................................................................................................................ 28

- Epigyne without hood and median septum ................................................................................................................ 29

28 Epigyne with a pairs of hoods ( Fig 2 d) ...................................................................................... L. auriculatus sp. nov.

- Epigyne with a median septum ( Fig 16 e ) ...................................................................................... L. spirellus sp. nov.

29 Copulatory openings visible .......................................................................................................................... L. ambrosi

- Copulatory openings invisible .................................................................................................................................... 30

30 Copulatory ducts visible ............................................................................................................................................. 31

- Copulatory ducts invisible ( Figs 14 i, j ) ........................................................................................ L. punctiger sp. nov.

31 Prosoma with a pair of paramedian stripes, spermathecae pyriform ( Figs. 3 c, d, l ) .................. L. clavellatus sp. nov.

- Prosome without markings, spermathecae global ( Figs 20 i, j ) ............................................... L. uniprocessus sp. nov.

32 Ratio of length of copulatory ducts / spermatheca = 0.5–1.0 ..................................................................................... 33

- Ratio of length of copulatory ducts / spermatheca = 1.2–3.0 .................................................................................... 36

33 Ratio of length of transversal slerotized plate / spermatheca = 0.8, the shield of prosoma brown with black paramed- ian stripes or black ..................................................................................................................................................... 34

- Ratio of length of transversal slerotized plate / spermatheca = 1.5, the shield of prosoma yellow without markings .. ..................................................................................................................................................................................... 35

34 Dorsum of opisthosoma with 2 pairs of joint black stripes posteriorly, spermatheca pyriform (Figs a, b, i) ................ ............................................................................................................................................................... L. kunmingensis

- Dorsum of opisthosoma with various black markings, spermatheca global ( Figs 13 a–d , l) ........................... L. niger

35 Transversal slerotized plate anteriorly, much developed ............................................................................ L. bhutanus

- Transversal slerotized plate medially, very thin ............................................................................................. L. conicus

36 Copulatory ducts slender, Ratio of width of copulatory ducts / spermatheca = 0.2 ................................................... 37

- Copulatory ducts thick, Ratio of width of copulatory ducts / spermatheca = 0.5–1.0 ................................................ 38

37 Ratio of length of copulatory ducts / spermatheca = 3.0 ............................................................................. L. dentatus

- Ratio of length of copulatory ducts / spermatheca = 1.2 ( Fig 10 e ) ...................................................... L. himalayensis

38 Ratio of width of copulatory ducts / spermatheca = 0.8–1.0 ...................................................................................... 39

- Ratio of width of copulatory ducts / spermatheca = 0.5 ( Fig 15 l ) .................................................................. L. saltus

39 The shield of prosoma yellow, copulatory ducts straight ( Figs 4 a–c ) ........................................... L. conflatus sp. nov.

- The shield of prosoma black, copulatory ducts twist ....................................................................................... L. davidi

Figures

FIGURES 2 a- – e. Lysiteles auriculatus sp. nov. a Female, habitus; b, d Epigyne; c, e Vulva; H hood. Scale bars: a = 1; b – e = 0.1.

FIGURES 20 a – j. Lysiteles uniprocessus sp. nov. a – c Habitus (a male; b, c female); d – f, h Left palp (d prolateral, e, h ventral, f retrolateral view); g, i Epigyne. j Vulva. Scale bars: a – c = 1; d – j = 0.1.

FIGURES 19 a – i. Lysiteles transversus sp. nov. a – c Habitus (a male; b – c female); d – g Left palp (d prolateral, e ventral, f retrolateral, g dorsal view); h Epigyne; Scale bars: a – c = 1; d – i = 0.1.

FIGURES 19 j – l. Lysiteles transversus sp. nov. j Left palp, k Epigyne; l Vulva. Scale bars: k – l = 0.1.

FIGURES 18 a – e. Lysiteles torsivus Zhang, Zhu & Tso, 2006. a Male, habitus; b – e Left palp (b prolateral, c – d ventral, e retrolateral view). Scale bars: a = 1; b – e = 0.1.

FIGURES 8 a – e. Lysiteles distortus sp. nov. a Male, habitus; b – e Left palp (b prolateral, c – d ventral, d retrolateral view). Scale bars: a = 1; b – e = 0.1.

FIGURES 17 a – l. Lysiteles subdianicus sp. nov. a – d Habitus (a, b male; c, d female); e – g, j Left palp (e prolateral, f, j ventral, g retrolateral view); h, k Epigyne; i, l Vulva. Scale bars: a – d = 1; e – l = 0.1.

FIGURES 7 a – f. Lysiteles dianicus Song & Zhao, 1994. a – b Habitus (a male, b female); c – f Left palp (c prolateral, d, i ventral, e retrolateral, f dorsal view). Scale bars: a – b = 1; c – f = 0.1.

FIGURES 7 i – k. Lysiteles dianicus Song & Zhao, 1994. i Left palp, j Epigyne; k Vulva; A atrium. Scale bars: i – k = 0.1.

FIGURES 15 a – l. Lysiteles saltus Ono, 1979. a – d Habitus (a, b male; c, d female); e – g, j Left palp (e prolateral, f, j ventral, g retrolateral view); h, k Epigyne; i, l Vulva. Scale bars: a – d = 1; e – l = 0.1.

FIGURES 4 a – g. Lysiteles conflatus sp. nov. a – c Habitus (a male, b – c female); d – g, j Left palp (d prolateral, e, j ventral, f retrolateral, g dorsal view). Scale bars: a – c = 1; d – g = 0.1.

FIGURES 3 a – l. Lysiteles clavellatus sp. nov. a – d Habitus (a male, b – d female); e – g, j Left palp (e prolateral, f, j ventral, g retrolateral view); h – i, k Epigyne. l Vulva; ATA apical tegular apophysis. Scale bars: a – d = 1; e – l = 0.1.

FIGURES 14 a – j. Lysiteles punctiger Ono, 2001. a – d Habitus (a, b male; c, d female); e – f, h Left palp (e, h ventral, f retrolateral view); g, i Epigyne, j Vulva. Scale bars: a – d = 1; e – j = 0.1.

FIGURES 11 a – i. Lysiteles kunmingensis Song & Zhao, 1994. a – b Habitus (a male, b female); c – e, g Left palp (c prolateral, c, g ventral, e retrolateral view); f, h Epigyne; i Vulva. Scale bars: a – b = 1; c – i = 0.1.

FIGURES 13 a – k. Lysiteles niger Ono, 1979. a – d Habitus (a male, b – d female); e – g, j Left palp (e prolateral, f, j ventral, g retrolateral view); h, k Epigyne; i, l Vulva. Scale bars: a – d = 1; e – l = 0.1.

FIGURES 12 a – l. Lysiteles maius Ono, 1979. a – d Habitus (a, b male; c, d female); e – g, j Left palp (e prolateral, f, j ventral, g retrolateral view); h, k Epigyne; i, l Vulva. Scale bars: a – d = 1; e – l = 0.1.

FIGURES 1 a – e. Lysiteles arcuatus sp. nov. a Female, habitus; b, d Epigyne; c, e Vulva; A atrium; MS median septum; WOA width of atrium. Scale bars: a = 1; b – e = 0.1.

FIGURES 5 a – g. Lysiteles corrugus sp. nov. a – c Female, habitus; d, f Epigyne; e, g Vulva; MS median septum. Scale bars: a – c = 1; d – g = 0.1.

FIGURES 9 a – e. Lysiteles guoi sp. nov. a Female, habitus; b – c Epigyne; d – e Vulva. Scale bars: a = 1; b – e = 0.1.

FIGURES 6 a – g. Lysiteles curvatus sp. nov. a – c Female, habitus; d, f Epigyne; e, g Vulva. Scale bars: a – c = 1; d – g = 0.1.

FIGURES 16 a – f. Lysiteles spirellus sp. nov. a, b Female, habitus; c, e Epigyne; d, f Vulva; MS median septum. Scale bars: a – b = 1; e – f = 0.1.

FIGURES 10 a – e. Lysiteles himalayensis Ono, 1979. a Female, habitus; b, d Epigyne; c, e Vulva, TSP transversal sclerotized plate. Scale bars: a = 1; b – e = 0.1.