Stylogaster tanzaniensis, Stuke, Jens-Hermann, 2015

Stuke, Jens-Hermann, 2015, New Conopid records from the Afrotropical Region (Diptera). Part 1: Paramyopa Kröber, Pseudoconops Camras, Stylogaster Macquart, Thecophora Rondani, and Zodion Latreille, Zootaxa 3963 (2), pp. 101-159 : 153-156

Stylogaster tanzaniensis spec. nov.

( Figs 123–132 )

Type material. Holotype ♂: (1) Tanzania: / Mbeya 35kmS / Rt.A345, 2200m / 1.ix.1996 / A. Freidberg; (2) Holotypus / Stylogaster tanzaniensis / spec. nov. / det. Stuke, 2014.

Holotype is deposited in the TAUI. Abdomen dissected, macerated and stored in glycerine in a microvial pinned beneath the specimen. The holotype is otherwise in perfect condition.

Description of holotype (male). Body length about 6.2 mm. Wing-length 5.5 mm. Head-height 1.2 mm. Head. Eye dark brown, with a few small scattered inconspicuous ommatrichia. Facets on inner side of eye slightly enlarged. Ocelli yellow. Ocellar tubercle brown. No ocellar setae on holotype—these may be broken off although no pits are evident. Ocellar triangle occupies almost the entire frons and reaches as far as antennae ( Fig. 124 ). Ocellar triangle brown to yellowish brown. Frons black to light brown lateral to ocellar triangle. No distinct frontoorbital setae. Scape yellowish brown, pedicel and basal flagellomere yellowish brown to brown. Arista threesegmented, dark brown, and situated dorsally on basal flagellomere ( Fig. 125 ). Scape with a few brown setulae dorsally. Pedicel with black setulae dorsally and anteriorly. Basal flagellomere dorsally with a more shining surface than ventrally. Shape of antenna as illustrated in Fig. 125 . One distinct black vertical seta. Face pale yellow, silver pruinose. Postcranium black dorsally and yellow ventrally, distinctly silver pruinose. Postcranium with a line of regularly-arranged small white setulae. Some brown setulae at mouth opening. Proboscis mainly brown to black, except for yellowish brown distal division of labellum and yellowish brown base of proboscis. Labrum ca 2.8 mm, labellum approximately the same length. Thorax. Yellowish brown to light brown, with scutum (with exception of postpronotum), scutellum and mediotergite brown. All setae black. Two notopleural setae, 1 strong supra-alar seta, 2 strong postalar setae, no prescutellar dorsocentral seta, 1 apical scutellar seta, 1 seta on anepimeron and 1 seta above fore coxa on propleuron. Mesoscutum without black semi-adpressed hairs but with very small, scattered brown hairs only. Wing generally clothed in microtrichia, but bc, base of c, base of bm, and base of cup all virtually without microtrichia. Hind margin of wing with brown setulae. Venation as in Fig. 126 . Haltere orange-brown, distal part sharply defined dark brown. Legs whitish yellow, with brown hind femur. Hind tibia with an indistinct broad white ring subapically. Legs with black and brown setulae, lacking any white setulae. Fore coxa with 1–2 setae, mid and hind coxa each with 1 black seta. Hind trochanter lacking any tooth. Mid femur posteriorly in distal half with a row of regularly-arranged black setulae curved at their apices. Ventral side of mid femur without black erect setulae. All tibiae lacking conspicuous setulae. Hind tibia with short black spines on anterior surface. Claws yellowish brown basally, black distally. Pulvilli yellow or yellowish brown. Empodia short, pale brown. Abdomen. Pale brown, with tergite 1 brown medially, tergites 2–4 with brown posterior margin and broad brown median fascia, tergites 5–6 and epandrium largely dark brown. Tergites with semi-adpressed black setulae, and longer setulae laterally on tergite 5. Tergite 1 with several long black setulae laterally, tergite 2 with 2–3 black setae on either side of anterior margin. Terminalia as illustrated in Figs 127–132 . Cercus short, slightly widened laterally, rounded distally ( Fig. 127 ). No teeth or obvious bristles on cerci. Surstylus broad, with a finger-like process anterodorsally. Distal margin of surstylus with two black teeth. Surstylus with a unique anteroventral elongation which has an additional third strong black tooth. Small field of strong erect setulae on inner surface of surstylus. Two short membranous appendages on inner surface of surstylus directed from anterior margin backwards to the posterior margin, and ventrally to the anteroventral elongation. Surstylus with obvious long hairs. Phallus sheath as illustrated in Fig. 130 : Base of phallus sheath hardly protuding and covered with black spines.

Female unknown.

Etymology. The species is named after the state of Tanzania, from which the type material originates.

Discussion. Due to the large basal flagellomere Stylogaster tanzaniensis could key out using Stuke (2012) as Stylogaster hauseri Stuke, 2012 . Stylogaster hauseri is a distinct species with a completely orange-brown scutum and abdomen ( Stuke 2012: 294, Figs 77, 79 ) and a unique field of small black teeth on the surstylus ( Stuke 2012: 295, Figs 82–83 ). Stylogaster tanzaniensis is another species of the Stylogaster nitens group as defined above, and can be distinguished with the key given in the discussion of Stylogaster angolensis .

Distribution. Stylogaster tanzaniensis is only known from the locus typicus in Tanzania.

Figures

FIGURES 123 – 126. Stylogaster tanzaniensis spec. nov. (♂ holotype). 123. Lateral view of habitus. 124. Dorsal view of frons. 125. Lateral view of antenna. 126. Wing.

FIGURES 127 – 132. Stylogaster tanzaniensis spec. nov. (♂ holotype). 127. Lateral view of cercus. 128. Dorsal view of cerci. 129. Lateral view of phallus. 130. Lateral view of phallus sheath. 131. Lateral view of outer surface of surstylus. 132. Lateral view of inner surface of surstylus. mais—membranous appendage on inner surface of surstylus.

FIGURES 76 – 81. Stylogaster fianarantsoensis spec. nov. (♂ holotype). 76. Lateral view of outer surface of surstylus. 77. Lateral view of inner surface of surstylus. 78. Lateral view of phallus sheath. 79. Dorsal view of cercus. 80. Lateral view of cercus. 81. Lateral view of junction of cerci.

FIGURES 82 – 86. Stylogaster ivindoensis spec. nov. (♂ holotype). 82. Lateral view of thorax and head. 83. Dorsal view of frons. 84. Lateral view of antenna. 85. Posterior view of hind tibia. 86. Wing.