Ummidia cuicatec, Godwin & Bond, 2021

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia cuicatec sp. nov. Figs 43 , 44 , Map 5

Type material.

HOLOTYPE ♂ (UMM154) from Cuicatlán, Oaxaca, Mexico, 17.7984 -96.9519 6, 752 m a.s.l., 13.vi.1944, coll. H Wagner, AMNH. PARATYPE: 1 ♀ (UMM415, AMNH) from Portillo de Nejapa, Oaxaca, Mexico, 16.061 -95.9797 6, 635 m a.s.l., 20.vii.1966, coll. CM Bogert, AMNH.

Etymology.

The specific epithet is a noun taken in apposition and is in reference to the indigenous people group for whom the locality is named, the Cuicatec.

Diagnosis.

Ummidia cuicatec is a relatively large species in which males can be differentiated from all other species by having a long, sinuous embolus with an extreme distal curve. Males can be differentiated from U. anaya , U. huascazaloya , U. zilchi , U. riverai , and U. hondurena by having fewer spines on prolateral (2 vs 7-28) and retrolateral (10 vs 15-42) faces of tibia I and from U. riverai by lacking a pale dorsal heart patch on the abdomen. Females can be differentiated from all other geographically proximal species except U. pustulosa and U. frankellerae by having spermathecae that curve laterally and end in a distal coil with bulbs facing laterally. Males disperse in June.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Specimen in poor condition, metatarsus I appears to have been damaged. General coloration. Carapace and chelicerae reddish black 10R 2.5/1, legs very dusky red 10R 2.5/2. Abdomen very dark brown 7.5YR 2.5/3. Cephalothorax. Carapace 8.96 long, 8.53 wide. Pars cephalica 5.83 long. Foveal groove procurved, 0.78 long, 1.63 wide. All eyes on low tubercle. AER procurved. PER straight. Eye group 0.97 long, 1.64 wide, AME 0.37, PME 0.26, ALE 0.58, PLE 0.31. Sternum sparsely setose, STRl 5.21, STRw 4.86. Chelicerae with anterior tooth row comprising six teeth, posterior margin with four teeth. Palpal endites with 19 cuspules spread over proximal half of endite face, lacking distal endite cuspules, ENDw 1.83, ENDl 3.25. Labium with two cuspules, LBw 1.72, LBl 1.33. Rastellum with many small teeth on process. Abdomen setose. Legs. F1 7.78; F1w 2.12; P1 3.7; Ti1 5.62; Mt1 4.24; Tr1 2.02; F3 5.8; F3w 2.39; P3 3.01; Ti3 3.52; Sd3 2.06; Mt3 3.22; Tr3 2.55; F4 7.7; F4w 2.41; P4 3.45; Ti4 4.93; Mt4 5.51; Tr4 3.1. Retrolateral face of tarsus IV with distinct comb of spinules, which is surrounded in setae. Leg I spination pattern: TSp 0, TSpv 2, TSrd 0, TSr 0, TSrv 10, MtSp 3, MtSr 7, TrSp 3, TrSr 4. Pedipalps. PTl 4.28, PTw 1.5, Bl 3.63. Embolus relatively long and sinuous with strong bend distally.

Variation, males.

Known only from male type specimen.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. Right palp and leg IV and left leg II Ti-TR removed and in vial. Left palp missing. General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 2.5/4. Abdomen reddish black 2.5YR 2.5/1, spinnerets dark reddish brown 5YR 3/4. Cephalothorax. Carapace 8.98 long, 8.54 wide. Pars cephalica 6.4 long. Foveal groove procurved, 0.84 long, 2.27 wide. All eyes on moderate tubercle. AER procurved. PER procurved. Eye group 1.15 long, 1.93 wide, AME 0.43, PME 0.31, ALE 0.65, PLE 0.43. Sternum very setose around outer third and thicker posteriorly, STRl 5.83, STRw 5.54. Chelicerae with anterior row comprising four teeth, posterior margin with seven teeth. Palpal endites with 24 cuspules spread across proximal half of endite and 78 cuspules distally, ENDw 1.88, ENDl 3.42. Labium with seven cuspules, LBw 1.87, LBl 1.33. Rastellum with many strong spines on process and up cheliceral face for ~ 3 ' length of process. Abdomen. Evenly setose with pale speckles clustered at apodemes. Legs. F1 5.17; F1w 2.09; P1 3.53; Ti1 3.09; Mt1 2.24; Tr1 1.29; F3 4.83; F3w 2.58; P3 3.26; Ti3 2.93, Sd3 2.3; Mt3 2.26; Tr3 2.04; F4 6.36; F4w 2.76; P4 3.45; Ti4 3.67; Mt4 3.85; Tr4 1.91. Retrolateral face tarsus IV with brush of spinules, retrolaterally and ventrally setose. Pedipalps. PF 4.62, PP 2.91, PTi 3.01, PTr 2.56. Spermathecae curved laterally with distal coil, bulbs facing laterally.

Variation, females.

Known only from female type specimen.

Figures

Map 5. Locality records for southern Mexico and Central American species.

Figure 43. Ummidia cuicatec sp. nov. from Oaxaca, Mexico. A male habitus illustration UMM 154 B female habitus illustration UMM 415. Scale bars: 4.0 mm.

Figure 44. Ummidia cuicatec sp. nov. from Oaxaca, Mexico. A-D male holotype (UMM 154) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 415). Scale bars: 1.0 mm.