Amphineurus (Rhamphoneurus) billinghami, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

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Amphineurus (Rhamphoneurus) billinghami

sp. nov.

3.3.6. Amphineurus (Rhamphoneurus) billinghami sp. nov.

Material examined.

Holotype: ♂, Chile, Osorno , Pucotrihue [40°32′S 73°42′W], 10/ 12-IV-1968, Peña (USNM) GoogleMaps *. Paratype: 1 [sex unknown], Chile, Chiloé Is., Chepu [42°2′S 74°1′W], 14-II-1952, Peña (USNM) GoogleMaps .


The species is named after Zacariah D. Billingham, a great expert of Australian craneflies.


This species is characterized by crossvein m-cu uniformly inserted at the fork of bM. Furthermore, this species is characteristized by the male terminalia with a curved sheath of aedeagus, a small right mesal lobe and the most setose ventral branch of gonocoxite. The thorax is also distinguishable by a narrow knob of the halter and katepisternum longer than the meron.


Wing length 6.45 mm, width 1.66 mm. - Coloration: General coloration dark brown. Rostrum and palpus dark brown. First segments of the antenna yellow, remainder dark yellow. Head dark brown. Eyes blackish gray. Thorax dark brown, pleura pale. Scutum without stripes. Halter pale with knob dark yellow. Coxae brownish yellow and legs yellowish-brown. - Head (dorsal view Fig. 13B View Figure 13 , ventral view Fig. 13A View Figure 13 ): Rostrum bifurcated with short appendices; first palpal segment as long as terminal segment; scape thicker than pedicel, terminal flagellomere as long as penultimate. - Thorax (lateral view Fig. 13E View Figure 13 , dorsal view Fig. 13F View Figure 13 ): Thorax with long setae, particularly on scutum. Anatergite as tall as katatergite. Halter with narrow knob. Wing (Fig. 13D View Figure 13 ) without clear markings; veins near fork of bM (Fig. 13C View Figure 13 ) visible; m-cu inserted at fork of bM; M1+2 longer than basal deflection of M1; R2 faded; R2+3+4 shorter than R2+3. - Male terminalia (Fig. 13G View Figure 13 ): Male tergite IX with small V-shaped median notch; posterolateral lobes rounded. Ventral branch of gonocoxite expanded on distal tip. Dorsal branch of gonocoxite nearly half-length of ventral branch; both branches extremely setose. Lobe of gonostylus rounded, longer than wide; longer than half-length of lobe of gonostylus; lobule wider than stem. Clasper of gonostylus with similarly shaped branches; lateral branch rounded; medial branch oblong. Mesal lobes of gonocoxite asymmetrical, both pointed, with several setae on posterior margin; right mesal lobe shorter. Sheath of aedeagus claw-shaped, curved halfway, darkened only on tip.

Remarks. Specimens of this new species were previously identified by C.P. Alexander as A. (R.) nullus Alexander. This species resembles A. (R.) theischingeri sp. nov. but differs mainly in attachment of m-cu, setae concentration in gonocoxite, shape of male tergite IX, and mesal lobe.