Beana mutaneni Pellinen, 2021

Pellinen, Markku J. & Wahlberg, Niklas, 2021, Changes in the genus Beana Walker (Beaninae Zahiri & Holloway) with two new species and removing Beana nitida Tams to a new genus Beanoides (gen. nov.), Zootaxa 4941 (3), pp. 415-424 : 418-421

Beana mutaneni Pellinen sp. nov.

( Fig.s 2 a–e and 3 a–d)

Type material. Holotype Thailand, Lampang, Muban Phichai, 16.v.2009 (slide No PS 1817). http://id.luomus. fi/308027. DNA extract LI-009 (stored at Department of Biology , Lund University, Sweden).

Paratypes. Males : Thailand, Lampang, Muban Phichai, 26.vi.2011 , idem 05.viii.2011 (slide No MJP1149) , idem 27.vii.2012 , idem 10.viii.2016 (slide No MJP1148) , idem 16.viii.2016 , idem 13.xi.2017 . Females: Thailand, Lampang, Muban Phichai , 19.vii.2013 , idem 18.vi.2014 (slide No MJP542) , idem 21.viii.2018 .

The holotype and paratypes are in the collection NHM Helsinki.

Diagnosis. Beana mutaneni sp. nov. is a close relative to Beana penniuncus sp. nov. by size (23 mm), smaller than the other known Beana species (30 mm). Externally B. mutaneni is darker with brownish transverse lines, B. penniuncus is more variegated and transverse lines are white. B. mutaneni has same kind strong sexual diorphism as the type species B. terminigera . Male genitalia are clearly different from those of B. penniuncus having uncus two-branched.

Description. Adult. Forewing length 23–25 mm in males and females. Head with frons unicolorous cream-colored scales, labial palpi curved upwards and apical segment narrower, neck with erect cream-colored and occasional blackish scales, male antennae shortly ciliate, females filiform. Male forewings basal, subbasal area blackish brown, apical part lighter grey, orbicular stigma unclear, reniform black, postmedial line double, submarginal line unclear. Hindwings light grey darkening apically, underside grey. Female forewings similar to male, except triangular dark area filling most part of the basal half of the wing. Abdomen 8 th tergite with flap-like apodemes. Male genitalia ( Fig. 1 a–c): Uncus small, slender, curved with fine tip, valva long, broadening apically, with long distal costal process, central process with brushes of setae, distal saccular process apically with brushes of long setae, saccus narrow, rounded, cucullus apically pointed, aedeagus large, narrow, straight. Female genitalia ( Fig. 1 d ): Papillae anales rigth-angled, apophyses posteriores narrow, twice length of apophyses anteriores, ductus bursae long. narrow, finely scobinate, corpus bursae roundish without scobination and signum.

Distribution. The new species is so far known from Northern Thailand, Lampang district.

Etymolgy. The species name is dedicated to Dr. Marko Mutanen (Ecology and Genetics research unit, University of Oulu, FINLAND).

Figures

FIGURE 1. Maximum likelihood topology showing the phylogenetic relationships of Beana species and the position of Beanoides based on molecular data. Clades not relevant to this study are shown as collapsed. Numbers to the left of each node are the SH-like approximate likelihood/Ultrafast bootstrap values.

FIGURE 2. a. Beana mutaneni male Thailand, Lampang, Muban Phichai 05.viii.2011 b. male Thailand, Lampang, Muban Phichai 26.vi.2016 c. male Thailand, Lampang, Muban Phichai 05.viii.2011 d. female Thailand, Lampang, Muban Phichai 21.viii.2018 e. female Thailand, Lampang, Muban Phichai 18.vi.2014 f. Beana penniuncus male Thailand, Lampang, Muban Phichai 31.viii.2010