Horaeomorphus punctatus D.-Y. Zhou & S.-J. Zhang

Zhou, De-Yao, Zhang, Su-Jiong & Li, Li-Zhen, 2016, Contributions to the knowledge of the genus Horaeomorphus Schaufuss (Coleoptera, Staphylinidae, Scydmaeninae) in mainland China, ZooKeys 572, pp. 51-70: 55-58

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Horaeomorphus punctatus D.-Y. Zhou & S.-J. Zhang

sp. n.

Taxon classification Animalia Coleoptera Staphylinidae

Horaeomorphus punctatus D.-Y. Zhou & S.-J. Zhang  sp. n. Figs 3, 5

Type material

(1 ♂, 1 ♀). Holotype: CHINA: ♂, labeled 'China: Yunnan Prov., Yingjiang Hsien [盈江县], Xima [昔马], 1500-1650m, 20-22.v.2013, Wen-Xuan Bi leg.'. Paratype: 1 ♀, same locality as holotype.


Horaeomorphus punctatus  can be readily separated from all other congeners by its moderately large (2.07-2.36 mm) and stout body, broad pronotum with a row of three pits connected by a shallow, barely notable groove; dense and coarse punctation on elytra, unmodified metatrochanters in male, aedeagal parameres lacking setae at apices and shape of spermatheca.


Male. BL 2.36 mm; body (Fig. 3A) moderately large, flattened, dark reddish-brown, legs and palpi slightly lighter. Head broadest at large, finely faceted and moderately convex eyes, HL 0.34 mm, HW 0.46 mm; tempora shorter than eye in dorsal view, in anterior third nearly parallel, then strongly bent and in posterior third nearly transverse to long axis of head; vertex strongly transverse and weakly convex, with pair of small but distinct pits located near posterior margins of supra-antennal tubercles; frons weakly convex; supra-antennal tubercles strongly raised. Punctures on vertex and frons fine and inconspicuous; setation moderately long, sparse. Antennae (Fig. 5A) short, AnL 0.87 mm, relative lengths of antennomeres: 1.0 : 0.9 : 1.0 : 0.9 : 0.9 : 0.8 : 0.8 : 0.8 : 0.9 : 1.0 : 1.5. Pronotum broad, nearly circular, slightly longer than wide, widest at middle, PL 0.76 mm, PWm 0.72 mm, PWb 0.47 mm; posterior collar short, well delimited from disc by constriction and row of three shallow pits connected by shallow groove; punctation dense and coarse; setation moderately long. Elytra broad, flattened and distinctly impressed in middle at about anterior third; EL 1.26 mm, EW 0.84 mm, EI 1.50. Punctures coarse, more distinct than those on pronotum, sharply marked and separated by spaces as wide as diameters of punctures; setation long, moderately dense, erect to suberect. Hindwings fully developed. Metatrochanter (Fig. 5I) unmodified, with distinct ventral edge. Aedeagus (Fig. 5 B–D) relatively slender, with median lobe strongly narrowing toward apex, AeL 0.57 mm; endophallus (Fig. 5 F–G) relatively small and complicated, with sclerotized central portion and two moderately darkly sclerotized and curved structures; parameres (Fig. 5E) slender, minimally longer than median lobe, without apical setae.

Female. Similar to male, but with smaller body and less contractive pronotal base. BL 2.07 mm, HL 0.33 mm, HW 0.44 mm, AnL 0.75 mm, relative lengths of antennomeres: 0.8 : 0.8 : 0.9 : 0.8 : 0.7 : 0.7 : 0.6 : 0.7 : 0.6 : 1.0 : 1.2. PL 0.63 mm, PWm 0.58 mm, PWb 0.44 mm; EL 1.11 mm, EW 0.74 mm, EI 1.50, spermatheca (Fig. 5H) spherical, SpL 0.08 mm, with complicated internal structures.


Horaeomorphus punctatus  is similar to Horaeomorphus mesaios  Jałoszyński & Nomura, 2004 (Vietnam: Ninh Pinh), Horaeomorphus valdepunctatus  Franz, 1984 (Malaysia: Pahang), Horaeomorphus sarawakensis  Franz, 1992 (W Malaysia) and Horaeomorphus samosirensis  Jałoszyński, 2009 (Indonesia: Sumatra) in habitus, and is also very similar to Horaeomorphus mesaios  in the shape of aedeagus and structures of endophallus; but it can be identified by its relatively large body (2.07-2.36 mm), the pronotum with three basal pits; unmodified metafemora, non-protruded metatrochanters in males and achaetous parameral apex.


Southern China: Yunnan.


The specific epithet refers to the coarse punctation of the elytra.