Mitrella petersi ( Hilber, 1879 )

Harzhauser, Mathias & Landau, Bernard M., 2021, The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group, Zootaxa 5025 (1), pp. 1-75 : 29-30

Mitrella petersi ( Hilber, 1879)

Figs 4E, 8G 1 –G 2 , H 1 –H 2

* Columbella Petersi R. Hoern. u. Auing. in coll.— Hilber 1879: 420, pl. 1, fig. 2.

[ Mitrella ] Petersi nov. form.—Hoernes 1880: 126 (pars) [nomen nudum].

Columbella (Mitrella) Petersi nov. form.— Hoernes & Auinger 1880: 97 (pars) [non figs].

Columbella (Mitrella) kostejana n. sp. — Boettger 1906: 16.

Mitrella petersi Hörn u. Auing.— Meznerics 1933: 340.

Pyrene (Mitrella) kostejana (Boettger) — Zilch 1934: 253, pl. 16, figs 87a–b.

M.[itrella]. (A [tilia].) petersi (R. Hörn. et Au.) — Sieber 1958a: 147.

M.[itrella]. (A [tilia].) kostejana Bttg. — Sieber 1958a: 147.

Mitrella (Atilia) petersi (R. Hörn. Au.) — Sieber 1958b: 148.

Pyrene (Mitrella) kostejana Boettger — Toth 1948: 403.

Pyrene (Mitrella) kostejana ( Boettger, 1906) — Bałuk 1995: 235, pl. 32, figs 3–4.

Mitrella kostejana ( Boettger, 1906) — Landau et al. 2013: 190, pl. 29, figs 4–5, pl. 67, fig. 5.

non Atilia (Macrurella) Petersi Hoernes et Auinger — Cossmann 1901: 245 [= Mitrella elongatissima nov. sp.] non Pyrene (Mitrella) petersi Hoernes et Auinger , sp. 1880— Glibert 1952: 311, pl. 7, fig. 10 [= Mitrella gracilis Ivolas &

Peyrot, 1900]. non Pyrene (Mitrella) petersi Hoernes et Auinger — Csepreghy-Meznerics 1956: 403, pl. 7, figs 12–13 [?: Mitrella hilberi

Cossmann, 1901]. non Mitrella (Atilia) petersi Hörnes & Auinger 1880 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 184, pl. 45 [?:

Mitrella fallax ( Hoernes & Auinger, 1880) ] non Columbella (Atilia) petersi Hoernes & Auinger (in Hilber), 1879— Strausz 1966: 292, pl. 42, figs 8–9 [?: Mitrella hilberi

Cossmann, 1901].

Type material. Lectotype (designated herein): NHMW1858 /0020/0002, SL: 6.9 mm, MD: 2.3 mm, Guglitz at St. Florian ( Austria), illustrated in Hilber (1879, pl. 1, fig.1), Figs 4E, 8G 1 –G 2 .

Additional material. SMF 360680 (= XII.2211 a), Senckenberg Museum , Frankfurt / Main, Germany , SL: 5.8 mm, MD: 1.9 mm, Coşteiu de Sus ( Romania), lectotype of Columbella (Mitrella) kostejana illustrated in Zilch (1934, pl. 16, figs 87a–b), Figs 8H 1 –H 2 ; NHMW2020 /0174/0002 (ex B. Landau coll.), 2 spec., Korytnica ( Poland) .

Revised description. Shell moderately small, slender fusiform, with high, weakly cyrtoconoid spire and shallow suture. Low dome-shaped protoconch of about three weakly convex whorls with weak mid-whorl angulation. Teleoconch of seven whorls. Early spire whorls comparatively low, weakly convex. Later whorls increasing in height, flat-sided or faintly convex. Last whorl moderately high, weakly convex, slowly contracting into short base. Shell surface smooth except for few prominent spiral cords over siphonal fasciole. Aperture narrow. Outer lip with internal labial callus pad developed, straight sided with about six denticles. D1 strongest, weakening adapically. Anal canal indistinct, shallow, broad U-shaped. Siphonal canal moderately short, moderately wide with shallow siphonal notch. Columellar callus forming sharply delimited callus rim, detached from base, without columellar denticles; no parietal callus.

Shell measurements and ratios. SL: 11.4–12.1 mm, MD: 3.1–3.6 mm, AA: 30–35°, SL/MD: 3.0–3.2, AL/AW: 3.5–4.3, LWH/AH: 1.3–1.5.

Discussion. The lectotype of this species was described and illustrated by Hilber (1879) from the Styrian Basin. Hilber (1879) already referred to additional specimens from the Vienna Basin stored in the collections of the NHMW, which laid the foundation for future confusion. A year later, these ‘collection specimens’ were included by Hoernes & Auinger (1880) in their descriptions, which strongly distorted the understanding of Mitrella petersi . These specimens differ from Mitrella petersi in their larger size, higher spire and distinctly more slender outline, and are described herein as Mitrella elongatissima nov. sp. Probably due to these misidentifications by Hoernes & Auinger (1880), Boettger (1906) described this species again as Columbella (Mitrella) kostejana , which is a subjective junior synonym.

Palaeoenvironment. Inner neritic lagoonal environments (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Carpathian Foredeep: Korytnica ( Poland) ( Bałuk 1995); Vienna Basin: Perchtoldsdorf ( Austria) ( Toth 1948); Styrian Basin: Guglitz at St. Florian ( Austria) ( Hilber 1879), Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906).

Proto-Mediterranean Sea. Serravallian (middle Miocene): Karaman Basin: Seyithasan ( Turkey) ( Landau et al. 2013).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width.