ASPREDINIDAE Adams, 1854

Published, First, 2007, Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types, Zootaxa 1418, pp. 1-628 : 57-62

ASPREDINIDAE Adams, 1854

Aspredinidae Adams , in Adams et al., 1854: 107. Type genus: Aspredo Scopoli, 1777 .

Bunocephalini Bleeker, 1858b : xi, 328. Type genus: Bunocephalus Kner, 1855 .

Platystacinae Eigenmann & Eigenmann, 1890: 9, 13, 20. Type genus: Platystacus Bloch, 1794 .

Hoplomizoninae [sic Hoplomyzontinae ] Fernández-Yépez, 1950b:113. Type genus: Hoplomyzon Myers, 1942 .

Taxonomy summary: Gill (1891a); Friel (2003); Myers (1960); Mees (1987b, Aspredininae ); Mees (1988b Bunocephalinae ).

Keys: Mees (1988b, Aspredininae ).

Revision: Stewart (1985b, Hoplomyzontinae ).

Phylogeny: Friel (1994); Diogo et al. (2001).

12 genera, 36 species; no named fossil taxa.

ACANTHOBUNOCEPHALUS Friel, 1995 Acanthobunocephalus Friel, 1995: 90 . Type species: Acanthobunocephalus nicoi Friel, 1995 . Type by original designation. Gender: Masculine. Acanthobunocephalus nicoi Friel, 1995 Acanthobunocephalus nicoi Friel, 1995: 90 , fig. 1. Type locality: Río Sipapo, 200 m from Salto Remo, 4º34'N, 67º18'W, Venezuela. Holotype: MCNG 29000. Distribution: Upper Orinoco River basin of Venezuela, including Sipapo, Ramoni and Casiquiare rivers (Friel, 2003).

AMARALIA Fowler, 1954 Amaralia Fowler, 1954: 40 . Type species: Bunocephalus hypsiurus Kner, 1855 . Type by original designation. Gen- der: Feminine. Amaralia hypsiura (Kner, 1855) Bunocephalus hypsiurus Kner, 1855a: 98 , pl. 1 (fig. 1). Type locality: Rio Branco [ Brazil]. Syntypes: NMW 47626 (2). Distribution: Throughout the Amazon River basin (Friel, 2003).

ASPREDINICHTHYS Bleeker, 1858

Aspredinichthys Bleeker, 1858b: 328 , 329. Type species: Aspredo tibicen Valenciennes, 1840 . Type by monotypy. Gender: Masculine.

Chamaigenes Eigenmann, 1910: 380 . Type species: Aspredo filamentosus Valenciennes, 1840 . Type by original designation. Gender: Masculine.

Revison: Mees (1987b).

Aspredinichthys filamentosus (Valenciennes, 1840)

Aspredo filamentosus Valenciennes , in Cuvier & Valenciennes, 1840b: 437 (324 of Strasbourg deluxe edition), pl. 450. Type locality: Cayenne. Syntypes: MNHN 0000-4401 (4), MNHN b-0594 (2).

Distribution: Lower portions of rivers and in coastal waters of northern South America from Venezuela to northern Brazil (Friel, 2003).

Aspredinichthys tibicen (Valenciennes, 1840)

Aspredo tibicen Valenciennes , in Cuvier & Valenciennes, 1840b: 438 (325 of Strasbourg deluxe edition). Type locality: Surinam. Holotype: RMNH 3111 .

Distribution: Lower portions of rivers and in coastal waters of northern South America from Venezuela to northern Brazil (Friel, 2003).

ASPREDO Scopoli, 1777

Aspredo Scopoli, 1777: 453 . Type species: Silurus aspredo Linnaeus, 1758 . Type by absolute tautonymy. Gender: Masculine.

Aspredo aspredo (Linnaeus, 1758)

Silurus aspredo Linnaeus, 1758: 304 . Type locality: in Americae fluviis. Syntype: ZMUU Linnaean Coll. 48 (1); also based on descriptions in Gronovius (1754: 8), which apparently is based on the same Linnaean specimen, and on Klein (1775).

Platystacus laevis Bloch, 1794: 58 . Type locality: [No locality stated]. Syntypes: ZMB 3153 (3), and several literature sources, including Silurus aspredo Linnaeus .

Aspredo sicuephorus Valenciennes , in Cuvier & Valenciennes, 1840b: 439 (326 of Strasbourg deluxe edition). Type locality: La Mana [ French Guiana]. Holotype: MNHN 0000-6362.

Aspredo batrachus Gronow , in Gray, 1854: 137. Type locality: fluminibus Americes Meridionalis. Syntype or holotype: ZMUU Linnaean Coll. 48 (1); inasmuch as Gronovius (1754: 8) based the description on an earlier Linnaean description.

Aspredo sicyephorus Günther, 1864: 269 . Type locality: Mana [ French Guiana]. Holotype: MNHN 0000-6362. Unjustified emendation of Aspredo sicuephorus Valenciennes, 1840 .

Distribution: Lower portions of rivers from Venezuela to northern Brazil (Friel, 2003).

BUNOCEPHALUS Kner, 1855

Aspredo Swainson, 1838: 332 . Type species: Aspredo gronovii Swainson, 1838 . Type by monotypy. Gender: Masculine. Preoccupied by Aspredo Scopoli, 1777 ; apparently not replaced.

Bunocephalus Kner, 1855a: 95 . Type species: Platystacus verrucosus Bloch, 1794 . Type by subsequent designation by Bleeker (1862–63: 19). Gender: Masculine.

Platystacus Bleeker, 1858b: 329 . Type species: Bunocephalichthys verrucosus Bloch, 1794 . Type by subsequent designation by Jordan (1919a: 279). Gender : Masculine.

Dysichthys Cope, 1874b: 133 . Type species: Dysichthys coracoideus Cope, 1874 . Type by monotypy. Gender: Masculine.

Agmus Eigenmann, 1910: 379 . Type species: Bunocephalus scabriceps Eigenmann & Eigenmann, 1889 . Type by original designation. Gender: Masculine.

Remarks: Ferraris (1991) clarified type species of Bunocephalus , but see Mees (1997) for alternate opinion. Many of the species listed here were placed in Dysichthys by Mees (1988b, 1989), but Dysichthys was placed in the synonymy of Bunocephalus in Friel (2003).

Review: Mees (1989, as Dysichthys , in part).

Bunocephalus aleuropsis Cope, 1870

Bunocephalus aleuropsis Cope, 1870b: 568 . Type locality: Pebas , Eastern Ecuador [now Peru]. Syntypes: ANSP 8286–8288 (3); one syntype illustrated in Mees (1989: 206, fig. 4).

Bunocephalus melas Cope, 1874b: 132 . Type locality: Nauta. Holotype: ANSP 21235 (in pieces).

Distribution: Amazon and Orinoco River basins (Friel, 2003).

Bunocephalus amaurus Eigenmann, 1912

Bunocephalus amaurus Eigenmann, 1912b: 126 , fig. 24; pl. 2 (fig. 2). Type locality: Konawaruk , British Guiana. Holotype: FMNH 53121 .

Bunocephalus amaurus aloikae Hoedeman, 1961: 130 . Type locality: Litany R., Aloiké village, French Guiana. Holotype: ZMA 102229 .

Bunocephalus amaurus sipaliwini Hoedeman, 1961: 130 , fig. 1. Type locality: Sipaliwini, Suriname, 20 km from frontier with Brazil. Holotype: ZMA 102228 .

Distribution: Coastal rivers of northern South America between Orinoco to Amazon mouths (Friel, 2003).

Remarks: Redescribed in Mees (1989: 216) as a subspecies of Dysichthys coracoideus .

Bunocephalus amazonicus (Mees, 1989)

Dysichthys amazonicus Mees, 1989: 241 , fig. 21. Type locality: Bolivia: creek near Todos Santos, upper course of R. Mamoré , Cochabamba. Holotype: ZMA 109246 (37 mm SL specimen).

Distribution: Upper and middle Amazon River basin (Friel, 2003).

Bunocephalus bifidus Eigenmann, 1942

Bunocephalus bifidus Eigenmann , in Eigenmann & Allen, 1942: 86, pl. 2 (fig. 2). Type locality: creek, Yurimaguas [ Huallaga River , Peru]. Holotype: CAS 35105.

Distribution: Amazon River basin of Bolivia, Brazil, Ecuador, and Peru (Friel, 2003).

Remarks: Name appeared several times (Pearson, 1924: 9; Pearson 1937: 111; Fowler, 1940c: 94) as a nomen nudum prior to being made available in Eigenmann & Allen (1942).

Bunocephalus chamaizelus Eigenmann, 1912

Bunocephalus chamaizelus Eigenmann, 1912b: 127 , fig. 24; pl. 2 (fig. 1). Type locality: Erukin , British Guiana. Holotype: FMNH 53122 .

Distribution: Essequibo River basin (Friel, 2003).

Bunocephalus colombianus Eigenmann, 1912

Bunocephalus colombianus Eigenmann, 1912a: 10 . Type locality: Raspadura [ Colombia]. Holotype: FMNH 56038 .

Distribution: Atrato and Magdalena River basins (Friel, 2003); also Cauca, San Jorge, Sinu and Patía basins, Colombia (Maldonado-Ocampo et al., 2005).

Bunocephalus coracoideus (Cope, 1874)

Dysichthys coracoideus Cope, 1874b: 133 . Type locality: Nauta [ Peru]. Syntypes: ANSP 21212–21215 (4).

Bunocephalus bicolor Steindachner, 1882b: 176 . Type locality: Aus dem Huallaga [ Peru]. Holotype: MTD F345 . Species illustrated and described in more detail in Steindachner (1882c: 8, pl. 2, figs. 1, 1a, 1b).

Bunocephalus haggini Eigenmann & Allen, 1942: 86 , pl. 2 (fig. 1). Type locality: Amazon at Iquitos [ Peru]. Holotype: CAS 35107.

Distribution: Amazon River basin of Bolivia, Brazil, and Peru (Friel, 2003).

Bunocephalus doriae Boulenger, 1902

Bunocephalus doriae Boulenger, 1902b: 286 . Type locality: Villa Rica, Paraguay; and Posadas , Río Parana [ Argentina]. Syntypes: BMNH 1902.2.10.22–23 (2), MSNG 14274 (1) , MSNG 14275 (3).

Bunocephalus retropinnis Eigenmann , in Eigenmann & Allen, 1942: 85, pl. 2 (fig. 3). Type locality: Cacequi, Brazil. Holotype: CAS 35246.

Distribution: Paraguay-Paraná and Uruguay River basins, of Argentina, Brazil, Paraguay and Uruguay (Friel, 2003).

Bunocephalus iheringii Boulenger, 1891

Bunocephalus iheringii Boulenger, 1891: 235 , pl. 26 (fig. 2). Type locality: Not stated [apparently from Rio Grande do Sul, Brazil]. Syntypes: BMNH 1891.3.16.77–78 (2).

Bunocephalus salathei Myers, 1927: 125 . Type locality: Brazil: Morro Ajudo , about 100 km from Rio de Janeiro. Holotype: MCZ 31583.

Bunocephalus minutus Güntert, 1942: 28 , figs. 1–2. Type locality: Paraguay, Dept. Villetta. Syntypes: NMBA 5296– 98 (3), NMBA 5299–5300 (2).

Bunocephalus carvalhoi Miranda Ribeiro , 1944a: 1, figure on p. 2. Type locality: Magé, Estado do Rio de Janeiro [ Brazil]. Holotype: MNRJ 1060 .

Distribution: Atlantic, Paraguay and Uruguay River basins of Brazil and Paraguay (Friel, 2003).

Bunocephalus knerii Steindachner, 1882

Bunocephalus Knerii Steindachner, 1882b: 176 . Type locality: Canelos [ Ecuador]. Syntype: NMW 47628. Species illustrated and described in more detail in Steindachner (1882c: 9, pl. 2, figs. 2, 2a, 2b).

Distribution: Upper Amazon River basin of Colombia, Ecuador, and Peru (Friel, 2003).

Bunocephalus larai Ihering, 1930

Bunocephalus larai Ihering, 1930: 101 , pl. 13 (figs. 3–3a). Type locality: Rio Piracicaba , junto ao Salto; ao 20 klm abaixo (ilha das Flechas) [São Paulo, Brazil]. Syntypes: Whereabouts unknown.

Distribution: Paraná River basin of Brazil (Friel, 2003).

Bunocephalus quadriradiatus (Mees, 1989)

Dysichthys quadriradiatus Mees, 1989: 244 , figs. 23, 24. Type locality: Peru: Samiria , Chinguito [= Cocha Shinguita or Shirguita]. Holotype: MHNG 2157.21 .

Distribution: Upper Amazon River basin of Peru (Friel, 2003).

Bunocephalus rugosus Eigenmann & Kennedy, 1903

Bunocephalus rugosus Eigenmann & Kennedy, 1903: 498 . Type locality: Laguna near Arroyo Chagalalina [ Paraguay]. Holotype: IU 9819 (whereabouts unknown).

Dysichthys australe Eigenmann & Ward , in Eigenmann, McAtee & Ward, 1907: 113, pl. 31. Type locality: Corumba [ Paraguay]. Holotype: CAS 35240.

Distribution: Paraguay-Paraná River basin of Brazil and Paraguay (Friel, 2003).

Bunocephalus verrucosus (Walbaum, 1792)

Silurus verrucosus Walbaum, 1792: 574 . Type locality: [No locality stated]. No types known. Based on Gronovius (1754, pl. 5, fig. 3).

Platystacus verrucosus Bloch, 1794: 63 , pl. 373 (fig. 3). Type locality: Wahrscheinlich gerhört er in Surinam zu Hause [= Suriname]. On Gronovius (1754: 5, pl. 5, fig. 3; and 1763: 102, no. 325).

Aspredo gronovii Swainson, 1838: 332 , fig. 80. Type locality: [No locality stated]. No types known.

Aspredo verrucosa Gronow , in Gray, 1854: 137. Type locality: America Meridionali. No types known; based on Gronovius (1763: no. 325, pl. 5, fig. 3.).

Bunocephalichthys gronovii Bleeker, 1858b: 329 . Type locality: Am. Merid. No types known; based on Gronovius (1754: 153, pl. 5, fig. 3).

Bunocephalus scabriceps Eigenmann & Eigenmann, 1889b: 49 . Type locality: Jutahy [ Brazil]. Lectotype: MCZ 7967 ; lectotype illustrated and designated in figure caption of Eigenmann (1912: pl. 4) as 42 mm specimen.

Agmus lyriformis Eigenmann, 1912b: 128 , fig. 25; pl. 3. Type locality: Gluck Island [Rupununi River], British Guiana. Holotype: FMNH 53120 .

Distribution: Rivers of the Guianas and throughout the Amazon River basin (Friel, 2003).

DUPOUYICHTHYS Schultz, 1944 Dupouyichthys Schultz, 1944c: 244 . Type species: Dupouyichthys sapito Schultz, 1944 . Type by original designation. Gender: Masculine.

Dupouyichthys sapito Schultz, 1944 Dupouyichthys sapito Schultz, 1944c: 245 , pl. 4 (fig. d); fig. 4b. Type locality: Río Motatán , at the bridge 22 km north of Motatán , Venezuela. Holotype: USNM 121072 . Distribution: Magdalena River and Lake Maracaibo basins (Friel, 2003). Remarks: Redescribed in Mojica et al. (2000).

ERNSTICHTHYS Fernández-Yépez, 1953

Ernstichthys Fernández-Yépez, 1953: 3 , 4. Type species: Ernstichthys anduzei Fernández-Yépez, 1953 . Type by original designation. Gender: Masculine.

Ernstichthys anduzei Fernández-Yépez, 1953

Ernstichthys anduzei Fernández-Yépez, 1953: 5 , fig. 1. Type locality: Río Salinas , northeast of El Baúl, Estado Cojedes, Venezuela. Holotype: MHNLS 28 .

Distribution: Orinoco River basin of Venezuela (Friel, 2003).

Ernstichthys intonsus Stewart, 1985

Ernstichthys intonsus Stewart, 1985b: 16 , figs. 1f, 3, 9. Type locality: Río Napo at Añangu, Napo Prov, 0º30.8'S, 76º24.0'W, Ecuador. Holotype: FMNH 94603 .

Distribution: Napo River basin of Ecuador (Friel, 2003).

Ernstichthys megistus (Orcés, 1961)

Hoplomyzon megistus Orcés, 1961: 3 , figs. 1–2. Type locality: Small river off of the Río Chicherota, Río Bobonaza

system, Prov. Pastaza, e. Ecuador. Holotype: Author’s personal coll. 4305 (current whereabouts unknown). Distribution : Bobonaza and Marañon River basins of Ecuador and Peru (Friel, 2003).

HOPLOMYZON Myers, 1942

Hoplomyzon Myers, 1942: 94 . Type species: Hoplomyzon atrizona Myers, 1942 . Type by original designation. Gen- der: Masculine.

Hoplomyzon atrizona Myers, 1942

Hoplomyzon atrizona Myers, 1942: 95 , fig. 3. Type locality: Brook trib. to Río Zulia, at Estacion Tachira, 60 km north of San Cristobal, Venezuela, elev. ca. 150 m. Holotype: SU 36494 .

Hoplomyzon atrizona petroleus Schultz, 1944c: 248 , pl. 4 (fig. C); fig. 4a. Type locality: Río Motatán , 4 km above Motatán, Maracaibo basin, Venezuela. Holotype: USNM 121070 .

Distribution: Lake Maracaibo basin (Friel, 2003).

Hoplomyzon papillatus Stewart, 1985

Hoplomyzon papillatus Stewart, 1985b: 8 , figs. 1b, 2b, 4b, 5. Type locality: Río Aguarico , 1 km upstream from confluence with Río Shushufindi, Napo Prov., 0º17'S, 76º24.5'W, Ecuador. Holotype: FMNH 94908 .

Distribution: Napo and Portuguesa River basins of Ecuador and Venezuela, respectively (Friel, 2003).

Hoplomyzon sexpapilostoma Taphorn & Marrero, 1990

Hoplomyzon sexpapilostoma Taphorn & Marrero, 1990: 4 , figs. 1–4. Type locality: Río Masparro at site of Masparro Dam, Barinas, Venezuela, 8º50'40"N, 70º06'00"W. Holotype: MCNG 18669 .

Distribution: Orinoco River basin of Venezuela (Friel, 2003).

MICROMYZON Friel & Lundberg, 1996 Micromyzon Friel & Lundberg, 1996: 642 . Type species: Micromyzon akamai Friel & Lundberg, 1996 . Type by original designation. Gender: Masculine. Micromyzon akamai Friel & Lundberg, 1996 Micromyzon akamai Friel & Lundberg, 1996: 643 , fig. 1. Type locality: Rio Tocantins, above confluence with Rio Pará, 2º02'S, 49º17'W, Pará, Brazil, 10– 14 m. Holotype: MZUSP 48550. Distribution: Lower Amazon River basin (Friel, 2003).

PLATYSTACUS Bloch, 1794

Platystacus Bloch, 1794: 52 . Type species: Platystacus cotylephorus Bloch, 1794 . Type by subsequent designation by Bleeker (1862–63: 19). Gender: Masculine.

Platysomatos Bloch, 1797: 115 . Type species: Platystacus cotylephorus Bloch, 1794 . Type by being a replacement name. Unneeded new name for Platystacus Bloch, 1794 . Gender: Masculine. See Harry (1953) for comments on this name.

Cotylephorus Swainson, 1838: 354. Type species: Cotylephorus blochii Swainson, 1838. Type by monotypy. Gen- der: Masculine.

Platystacus cotylephorus Bloch, 1794

Platystacus cotylephorus Bloch, 1794: 54 , pl. 372. Type locality: Ostindischen Gewässer [in error; certainly from South America]. Syntypes: ZMB 3157 (2), and several specimens associated with literature sources .

Silurus hexadactylus La Cepède, 1803: 58 , 82. Type locality: [No locality stated]. Holotype: MNHN b-0029.

Cotylephorus Blochii Swainson, 1838: 354, fig. 90. Type locality: Ostindischen Gewässer [in error; certainly from South America]. Syntypes: ZMB 3157. Unneeded new name for Platystacus cotylephorus Bloch, 1794 , apparently to avoid tautonymy.

Aspredo sexcirrhis Valenciennes , in Cuvier & Valenciennes, 1840b: 441 (326 of Strasbourg deluxe edition). Type locality: Surinam. Syntypes: MNHN a-9313 (1), MNHN b-0029 (1).

Aspredo spectrum Gronow , in Gray, 1854: 137. Type locality: in fluminibus Americes Meridionalis. Types: Whereabouts unknown; based on Gronovius (1763: no. 324), and Seba (1734–65: pl. 29, fig. 9).

Platystacus nematophorus Bleeker, 1862a: 371 . Type locality: Surinama. Holotype: RMNH 3105; holotype illustrated in Bleeker (1864: 96, pl. 1, fig. 1).

Distribution: Coastal waters and lower portions of rivers of northern South America, from Venezuela to northern Brazil (Friel, 2003).

PTEROBUNOCEPHALUS Fowler, 1943

Pterobunocephalus Fowler, 1943c: 1 . Type species: Bunocephalus albifasciatus Fowler, 1943 . Type by original designation. Gender: Masculine.

Petacara Böhlke, 1959: 2 . Type species: Bunocephalus dolichurus Delsman, 1941 . Type by original designation. Gender: Neuter.

Pterobunocephalus depressus (Haseman, 1911)

Bunocephalus depressus Haseman, 1911a: 319 , pl. 48 (fig. 1), pl. 49. Type locality: Río Machupo near San Joaquín, Bolivia. Holotype: FMNH 54338 .

Bunocephalus albifasciatus Fowler, 1943c: 2 , fig. 1. Type locality: Todos Santos , Bolivia. Holotype: ANSP 69193 .

Distribution: Amazon, Orinoco and Paraguay River basins (Friel, 2003).

Pterobunocephalus dolichurus (Delsman, 1941)

Bunocephalus dolichurus Delsman, 1941: 77 , fig. 12. Type locality: Trombetas-river , near Obidos [ Brazil]. Holotype: IRSNB 359 .

Distribution: Amazon River basin of Brazil and Peru (Friel, 2003).

XYLIPHIUS Eigenmann, 1912

Xyliphius Eigenmann, 1912a: 10 . Type species: Xyliphius magdalenae Eigenmann, 1912 . Type by monotypy. Gen- der: Masculine. Spelled two ways in original description: Xyliphius and Xiliphius , first reviser not yet identified, but may not have been formalized until noted by Eschmeyer & Bailey, in Eschmeyer (1990).

Xyliphius barbatus Alonso de Arámburu & Arámburu, 1962

Xyliphius barbatus Alonso de Arámburu & Arámburu, 1962: 219 , figs. 1–2. Type locality: río Paraná en Rosario, provincia de Santa Fe, Argentina. Holotype: MLP 6798 .

Xyliphius labrosus Risso & Risso, 1964a: 11 . Apparently an unneeded new name for Xyliphius barbatus Alonso de Arámburu & Arámburu, 1962 .

Distribution: Paraguay-Paraná River basin, Argentina (Friel, 2003).

Xyliphius kryptos Taphorn & Lilystrom, 1983

Xiliphius kryptos Taphorn & Lilystrom, 1983: 43 , fig. 1. Type locality: Cerca del Puente de la carretera 6 sobre el Río Aricuaisá, Estado Zulia, Venezuela. Holotype: MCNG 1224 .

Distribution: Lake Maracaibo basin (Friel, 2003).

Xyliphius lepturus Orcés, 1962

Xyliphius lepturus Orcés, 1962: 50 , figs. 1–2. Type locality: Cerca de Desembocadura del rio Pucayacu en el Bobonaza [ Ecuador]. Holotype: Author’s personal collection 1307 (current whereabouts unknown).

Distribution: Upper Amazon and Orinoco River basins of Colombia, Ecuador and Venezuela (Friel, 2003).

Xyliphius lombarderoi Risso & Risso, 1964

Xyliphius lombarderoi Risso & Risso, 1964a: 12 , pl. 1 (figs. 1–2). Type locality: el Riacho Barranqueras [ Paraná , Argentina]. Holotype: Authors’ personal collection (current whereabouts unknown).

Distribution: Paraná River basin, Argentina (Friel, 2003).

Xyliphius magdalenae Eigenmann, 1912

Xyliphius magdalenae Eigenmann, 1912a: 10 . Type locality: Girardot , Colombia. Holotype: FMNH 56039 ; holotype illustrated in Eigenmann (1922: 27, pl. 1, fig. 1).

Distribution: Magdalena River basin, Colombia (Friel, 2003; Maldonado-Ocampo et al., 2005).

Xyliphius melanopterus Orcés, 1962

Xyliphius melanopterus Orcés, 1962: 52 , figs. 3–4. Type locality: el bajo Pucayacu no lejos de su desembocadura en el Bobonaza [ Ecuador]. Holotype: Author’s personal coll. 2021 (current whereabouts unknown).

Distribution: Upper Amazon and Orinoco River basins (Friel, 2003).