Thabena retracta ( Walker, 1857 )

Gnezdilov, Vladimir M., 2009, Revisionary notes on some tropical Issidae and Nogodinidae (Hemiptera: Fulgoroidea), Acta Entomologica Musei Nationalis Pragae 49 (1), pp. 75-92 : 78-79

Thabena retracta ( Walker, 1857)

( Figs. 1–4 , 16–24 )

Issus retractus Walker, 1857: 152 .

Thabena retracta: STÅL (1866b) : 393.

Type material examined. HOLOTYPE: J, Borneo ( BMNH).

Additional material examined. MALAYSIA: SARAWAK: Kalulong Mt. , 1800 ft, new forest, 4.xi.1932, 1 ♀, B. M. Hobby & A. W. Moore lgt., Oxford University Exp. ( BMNH) ; Dulit Mt. , 1 J, E. Mjorberg lgt. ( BMNH) ; foot of Dulit Mt. , junction of rivers, primaeval forest, 16.viii.1932, 1 ♀, Tinjar & Lejok lgt., B. M. Hobby & A. W. Moore lgt., Oxford University Exp. ( BMNH). SABAH: Samawang, near Sandakan, jungle, 14.vii.1927, 1 ♀ ( BMNH) ; Bukit Monkobo , 51°48′ N 116°58′ E, base-camp, stunted hill forest, 900 m a.s.l., 7.viii.1987, light trap, 1 J, A. H. Kirk- Spriggs lgt., NMW Sabah Expedition ( NMWC) . SINGAPORE: 1 ♀, H. N. Ridley lgt. ( BMNH) .

Supplementary description ( Figs. 1–4 ). Median keel of metope not reaching its upper margin. Postclypeus with median groove. Coryphe 2.3 times as wide as long, with short median keel from its posterior margin, anterior margin weakly convex. Fore wings with median trifurcate and cubitus anterior simple (vein formula M 3 CuA 1). First metatarsomere with 8–10 intermediate spines arranged in two rows.

Male genitalia ( Figs. 16–24 ). Hind margin of pygofer with basal incision in lateral view ( Fig. 24 ) and, in caudal view, with a pair of large triangular processes with apices directed inwards ( Fig. 23 ). Anal tube in dorsal view narrow basally and very wide medially, weakly narrowing to truncate apex; lateral margins turned down ( Figs. 21–22 ). Anal column short. Phallobase strongly curved in lateral view ( Fig. 16 ), with pair of bulges proximally ( Fig. 17 ). Each dorso-lateral lobe of phallobase with proximal concavity on ventral margin and a pair of distal processes in lateral view; apical process narrow, long, with denticles on lower margin, weakly sclerotised and covering phallotreme, subapical process short, well sclerotized with denticles on upper margin ( Figs. 16, 18 ). Ventral lobe of phallobase long and wide, weakly enlarged towards widely concave apex ( Fig. 17 ). Aedeagus with pair of long ventral hooks (half as long as aedeagus) narrowing apically and bearing denticles on external margins. Apical processes of aedeagus enlarged apically in lateral view. Posterior margin of style straight or convex ( Fig. 19 ). Capitulum of style situated on long neck, hastate in dorsal view ( Fig. 20 ); lateral tooth shaped as wide lobe, apical tooth distinct.

Female genitalia. Posterior margin of sternum VII with large rectangular process medially. Gonoplacs rounded, without keels. Furca weakly sclerotized, with long basal part. Anal tube wide, truncate apically. Anal column short. Each distal part of posterior connective lamina of gonapophyses IX straight, with apical lobe-shaped process. Median field of lamina shaped as single lobe bearing pair of lateral triangular processes. Each lateral field of lamina with long bifurcated process. Gonocoxa VIII with high lobe-shaped posterior margin proximally. Endogonocoxal process bifurcated apically. Anterior connective lamina of gonapophysis VIII with three teeth in apical group and two large teeth in lateral group.

Comments. The male specimen from Sabah differs from the specimen from Sarawak in having a tooth-shaped median process on the posterior margin of pygofer (absent in the specimen from Sarawak) and a convex posterior margin of style (straight in the specimen from Sarawak). I treat these differences as intraspecific variability.

Figures

Figs. 1–6. 1–4 – Thabena retracta (Walker, 1857), specimens from Sarawak: 1 –J, holotype, dorsal view; 2 – same, lateral view; 3 – same, face; 4 –♀, head, dorsal view. 5–6 – Gergithoides carinatifrons Schumacher, 1915, specimens from Taiwan: 5 – J, dorsal view; 6 – ♀, frontal view.

Figs. 16–24. Thabena retracta (Walker, 1857), specimen from Sarawak, male genitalia: 16 – penis, lateral view; 17 – penis, ventral view; 18 – penis, caudo-dorsal view; 19 – style, lateral view; 20 – capitulum of style, dorsal view; 21 – anal tube, dorsal view; 22 – anal tube, lateral view; 23 – pygofer, caudal view; 24 – pygofer, lateral view.