Amphimenes giganteus Fedorenko

Fedorenko, Dmitry N., 2010, New or little-known species of the genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) from Vietnam, ZooKeys 65, pp. 17-50 : 29-30

Amphimenes giganteus Fedorenko sp. n. Figs 9223140

Description.

Body length 8.5 –10.6/8.3– 10.2 mm, width 3.2-4.1 mm. Dorsum black, mouthparts and antennae red, clypeus and labrum mostly brown; reflexed side margin of pronotum dark brown to reddish-brown, that of elytra translucent reddish at the very base only. Gula brownish-red. Tarsi and tibiae, latter all along or apically, as well as all trochanters and procoxa red or brownish-red. Antennae often infuscated toward apex to brownish-red.

Eyes rather small and slightly flattened, a little longer than tempora; these smoothly extending into neck in dorsal view; posterior supraorbital seta situated about 1/3 distance between eye back margin and pronotal front margin. Frontal foveae fairly deep and reaching level to eye front margin. Antennae very long, surpassing pronotal base by last three or more joints, 3rd antennomere 1.88-2.05 (mean 1.95) times as long as 2nd, 8th 2.73-3.36 (mean 3.11) times as long as wide.

Pronotum subcordate, 1.18-1.26 (mean 1.22) times as wide as long, 1.37-1.49 (mean 1.43) times as wide as head, conspicuously sinuate before hind angles, with front angles pointed and approaching neck; hind angles very obtuse and strongly sloping forward. Base medial part weakly convex backward, often almost straight, with a very shallow border not extended to lateral lobes. Disc rather flat, mid-line superficial throughout its length and weakly separated from transverse basal depression, lateral basal foveae weak, each usually almost extended to anterior margin as a very shallow submarginal depression parallel to side margin. Paramedian foveae lengthwise, very shallow to indistinct.

Elytra elliptic, 1.41-1.48 (mean 1.45) times as long as wide, 1.49-1.69 (mean 1.60) times as wide as pronotum, broadest at about middle, with shoulders strongly rounded from nearly indistinct base; apical truncature strongly sinuate between protruding outer angles and a pointed apex; latter entire or divided at the very tip. D1/EL=0.10-0.14 (mean 0.13), D2/EL=0.61-0.76 (mean 0.70), D3/EL=0.90-0.98 (mean 0.94). Three or four inner striae weaker at base, prescutellary stria very weak to obsolete. Metepisternum very short, 0.96-1.0 times as long as wide.

Proximal pair of ventral setae on last tarsomere rudimentary or absent. Male profemur without ventral tubercle.

Penis (Figs 22, 31, 40) long, narrow, weakly arcuate in lateral view and abruptly curved to the right behind the middle in dorsal view; apical lamella large, long, parallel-sided, widely rounded apically; apical orifice rounded; both ventral striae and microsculpture absent.

Diagnosis.

The present species is easily recognizable among the other congeners by the combination of the large and wingless body, protruding outer angles of the elytral apical truncature, peculiar formula of discal setae, and other characters specified above.

Material.

Holotype ♂ (ZMMU) labelled: "S[outh] Vietnam, Lam Dong Prov. / Bi Doup - Nui Ba [Nature] Reserve / env. Long Lanh / 12°07'N 108°39'44"E / Bi Doup Mt., N. slope / h=1700-1900 m [a s l] / 12.IV.2008 / leg. D Fedorenko" [typewritten] “HOLOTYPE/…” [red typewritten]. Paratypes (ZISP, SIEE), 9 ♂♂, 9 ♀♀, same data but: 10. and 16.IV.2008, 3. and 6.V.2009; 12°11'N 108°42'E, 4 km SSE of Hon Giao Mt., h=1500-1700 m [a s l], 2-3.IV.2008 and 29.IV.2009; 12°10'44"N 108°40'44"E, h=1400-1600 m [a s l], 30. III– 21 (Fedorenko).

Type locality:

Vietnam, Lam Dong Province, Bi Doup Mt, 12°07'N 108°39'44"E.

Geographic distribution.

Known from type locality only.

Life history.

The species is common all over its type locality, where it occurs under fallen deadwood or in its larger open cavities; it has occasionally been found under the exfoliating bark of standing dead trees, sometimes together with Amphimenes bidoupensis .