Onthophagus (Onthophagus) cyanellus Bates, 1887

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 182

Onthophagus (Onthophagus) cyanellus Bates, 1887 Plate 40C

Onthophagus cyanellus Bates, 1887: 81 (original description. Type locality: MEXICO: Parada, Jalapa; GUATEMALA: San Jerónimo, Cerro Zunil; COSTA RICA: Río Sucio, Volcán de Irazu; PANAMA, Bugabá, Volcán de Chiriqui).

Onthophagus cyanellus : Gillet 1911a: 205 (complete list of species); Campos 1921: 57 (cited for Ecuador); Boucomont and Gillet 1927: 205 (catalog of species); Boucomont 1932: 323 (distribution); Blackwelder 1944: 211 (list of species for Latin America); Howden and Young 1981: 95 (characters in key), 114 (redescription); Zunino and Halffter 1988: 131 (redescription); 134 (distribution and biology); Kohlmann and Solís 2001: 164 (characters in key), 196 (redescription); Ratcliffe 2002: 17 (cited for Panama); Morón 2003: 69 (cited for Mexico); Pulido-Herrera and Zunino 2007: 101 (cited for Ecuador); Carvajal et al. 2011: 322-323 (cited for Ecuador); Krajcik 2012: 177 (complete list of species); Solís and Kohlmann 2012: 8 (list of species for Costa Rica); Delgado and Curoe 2014: 66 (characters in key, cited for Panama).

Onthophagus (Onthophagus) cyanellus : Chamorro et al. 2018: 97 (cited for Ecuador).

Type specimens.

Onthophagus cyanellus Bates, 1887. The lectotype is deposited at the NHML (see Zunino and Halffter 1988: 132). Locality: V de Chirirquí, 1300-2000 m, Panamá, not examined.

Distribution.

Colombia, Costa Rica, Ecuador, Guatemala, Mexico, Nicaragua, and Panama.

Literature records.

BOLIVAR: Chimbo ( Campos 1921: 57). GUAYAS: Bucay ( Campos 1921: 57).

Temporal data.

It is not known when this species was collected.

Remarks.

It is possible that this species may be found in coastal evergreen foothill forests. Campos (1921) reported this species from Chimbo and Bucay. However, we did not find any other record of this species in the collections examined. The collection method is unknown.