Glyphidocera fidem , DAVID ADAMSKI, 2005

DAVID ADAMSKI, 2005, Review of Glyphidocera Walsingham of Costa Rica (Lepidoptera: Gelechioidea: Glyphidoceridae), Zootaxa 858, pp. 1-158: 94-95

publication ID 10.5281/zenodo.170793

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Glyphidocera fidem

new species

Glyphidocera fidem  , new species

(Figs. 159–161, 285, 340, Map 28View MAP 28) Diagnosis.— Relationships between Glyphidocera fidem  and its congeners are difficult to determine because the former possesses a combination of features that include male abdomen with sex scales between terga 2–3 and 3–4 [fewer scales between terga 3–4 than between terga 2–3], a widened and densely setose gnathos; dorsal strut of tegumen with arms fused medially from posterolateral margin to base of uncus, digitate process of valva slightly curved laterally, costal furcae of valva equal in size, and apicoventral angle of valval cucullus broadly rounded.

Description.— Head: Vertex and frontoclypeus brown; outer surface of labial palpus brown intermixed with few brownish­yellow scales to near apical margin of segment II; inner surface with segments I –II pale brownish­yellow, segment III brown intermixed with few pale­brownish­yellow scales; scape and flagellum of antenna brown; male with 4 th flagellum dorsally protuberant and curved, forming a deep concavity opposite slightly excavated flagellomeres 5–6; concavity nearly closed by a few setiform scales on apex of 4 th flagellomere; proboscis pale brownish yellow.

Thorax: Tegula and mesonotum brown. Legs brown intermixed with pale brownishyellow scales along apical margins of all segments and tarsomeres. Forewing (Fig. 340): Length 5.2–6.1 mm (n = 18), brown intermixed with few pale­brown scales; cell and surrounding areas paler than marginal areas; cell with 1 or 2 dark­brown spots, 1 spot near middle [ma be present or absent], 1 spot near distal end. Undersurface brown. Hindwing: Brown

Abdomen (Fig. 159): Male with 6 transverse, irregular rows of sex scales on intersegmental membrane between segments 2–3. Intersegmental membrane between terga 3–4 with 2 rows of sex scales along dorsolateral area, and slightly widened medial area between terga 3–4. Sex scales within widened medial area are more sparsely distributed than sex scales on dorsolateral surface.

Male Genitalia (Figs. 160–161): Uncus elongate, slightly constricted above a wide base, slightly rounded distally, apex rounded and setose; gnathos densely setose, protuberant, wide throughout length, slightly upturned apically; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing medially, extending narrowly from posteroventral margin to base of uncus; costa of valva nearly straight from a shallow depression beyond a basal, digitate process to a gradually raised dorsal furca; apex widely bifurcate, inner margin between furcae U­shaped; each furca about equal in size, parallel, and pointed dorsolaterally; sacculus twisted apically slightly beyond midlength, constricting valva, widening distoventrally forming a broad cucullus; cucullus membranous and densely setose; vesica of aedeagus denticulate, bearing a large, conical, apical cornutus; cornutus basally bulbous, with one side slightly shortened, exposing a hollow, basal cavity.

Female Genitalia (Fig. 285): Papillae anales elongate, slightly wider basally than distally; apophysis posterioris and apophysis anterioris about equal in size; eighth sternum slightly narrower than eighth tergum; eighth sternum with posterior margin broadly rounded, anterior margin broadly emarginate; eighth tergum with posterior margin broadly rounded, anterior margin broadly emarginate; antrum enlarged posteriorly from a constricted anterior part, posterior margin with a shallow median notch, posterolateral margin broadly rounded, anterior end with a pair of elongate, spinelike processes within cavity formed from outer wall; ductus bursae short; ductus seminalis large, 4 ­coiled and denticulate basally, ductlike and membranous distally; ductus bursae with a moderately shortened 4 ­coiled, internal support; side opposite ductus bursae with an elongate and widened, internal, platelike support.

Holotype, ɗ, “Quepos, 80 m, P[arque] N[acional] Manuel Antonio, Prov[incia] Punt[arenas], COSTA RICA, R. Zuniga, Abr. 1991, L­S­ 370900, 448800 ”, “ INBio: COSTA RICA: CRI 000, 601719 ” [bar code label], “ INBio, Genitalia Slide by D. Adamski, Sex ɗ, No. 533 ” [yellow label].

Paratypes (7 ɗ, 10 Ψ): 3 ɗ, 4 Ψ, Same label data as above except, “ CRI 000, 601744 ”, “Slide No. 534 ”; “ CRI 000, 593437 ”; “ CRI 000, 601717 ”, “Ψ Slide No. 535 ”; “ CRI 000, 593384 ” (1 Ψ); “ CRI 000, 593325 ” (1 Ψ); “Ene.”, “ CRI 000, 564638 ”, “Slide No. 610 ”; “ CRI 000, 648446 ”, “Ψ Slide No. 536 ”; 3 ɗ, 6 Ψ, “Est. Queb. Bonita, 50 m, Res. Biol. Carara, Prov. Punt., COSTA RICA, R. Zuniga, Ene. 1991, L­N­194500, 469850”, “ CRI 000, 580554 ”, “Slide No. 628 ”; “ CRI 000, 580500 ”, “Slide No. 643 ”; “ CRI 000, 580579 ”, “Ψ Slide No. 640 ”; “Mar.”, “ CRI 000, 648569 ”, “Slide No. 641 ”; “Abr.”, “ CRI 000, 315703 ”, “Ψ Slide No. 630 ”; “Feb.”, “ CRI 000, 680112 ”, “Ψ Slide No. 646 ”; “Jun.”, “ CRI 000, 433567 ”, “Ψ Slide No. 631 ”; “ CRI 000, 433603 ”, “Ψ Slide No. 644 ”; “ CRI 000, 349049 ”, “Ψ Slide No. 645 ”; 1 ɗ, “Est. Los Almendros, Prov. Guana., COSTA RICA, 300 m, 26 –0 Abr. 1995, E. E. Lopez, L­N­ 334850, 369500, # 4790 ”, “ CRI 002, 202091 ”, “Slide No. 505 ”, (10 in INBio, 7 in USNM).

Distribution ( Map 28View MAP 28): Glyphidocera fidem  is known from collecting sites in northwestern Costa Rica along the Cordillera de Guanacaste, east to the south­central part of the country along the Pacific Coast.

Etymology: The species epithet fidem  is derived from the Latin “fides,” meaning truth.


National Biodiversity Institute, Costa Rica


Universidade do Extremo Sul Catarinense, Bairro Universit�rio


Smithsonian Institution, National Museum of Natural History