Juraniscus majeri, Kolibáč & Huang, 2019
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publication ID |
https://doi.org/ 10.4202/app.00550.2018 |
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DOI |
https://doi.org/10.5281/zenodo.10987131 |
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persistent identifier |
https://treatment.plazi.org/id/2F2187DD-FB39-FFFF-8926-009CFD42FB05 |
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treatment provided by |
Felipe |
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scientific name |
Juraniscus majeri |
| status |
sp. nov. |
Juraniscus majeri sp. nov.
Figs. 2B View Fig , 4B View Fig .
ZooBank LSID: urn:lsid:zoobank.org:act:6E188F79-0922-46BE-88CB-748A6ECBA438
Etymology: Named in memory of Karel Majer (1950–2000), an eminent researcher of the melyrid lineage of Cleroidea , for his contribution to the systematics and morphology of the group.
Type material: Holotype NIGP168479 View Materials ; male, counterpart absent; perfect dorsal side of specimen preserved, elytra semitransparent (abdomen and metathorax partly visible).
Type locality: Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.
Type horizon: Haifanggou Formation, Callovian, Middle Jurassic.
Diagnosis.—Antennomeres symmetrical including three terminal ones, progressively dilated from the fifth onwards; pronotum approximately 1.4 times as wide as long; anterior corners of pronotum weakly projecting; meso- and metatarsi with tarsomere 1 conspicuous, slightly shorter than tarsomere 2; tarsal lobes present in tarsomeres 1–4; tarsal claws without denticles.
Description.— Body length (from elytral apex to clypeus): 4.25 mm. For other measurements of visible body parts see Table 3 View Table 3 .
Coloration and sculpture: body elongate, weakly convex; body surface unicolorous, without distinct colour stripes or patches, merely remnants of dark pigmentation. Head with very fine sculpture composed of round punctures; interspaces between punctures as large as puncture diameter. Pronotum with similar fine sculpture but interspaces larger than puncture diameter; pronotal punctures without seta at centre. Elytron without carinae, with sparse sculpture composed of small, round punctures; interspaces the same size as, or larger than, puncture diameter; punctation appears irregular. Body completely without pubescence, short, sparse setae observed only on tibiae and labrum.
Head: frons flat; clypeus not extended anteriorly, distinctly separated by arcuate frontoclypeal suture; antennal socket visible from above; eye relatively large (space between eyes approximately three times eye diameter), distinctly elevated, not emarginate, elliptic, finely facetted; head inconspicuously narrowed behind eyes.
Mouthparts: mandibles medium-sized (not unidentate, slender and sharp), bidentate, acuminate; labrum oblong, weakly emarginate along anterior margin; maxillary palps 3-segmented, terminal palpomere slender, conical.
Antenna: 11-segmented; scape longer and wider than pedicel but not larger than two following antennomeres together; antennomeres symmetrical, progressively dilated from the fifth onwards; whole antenna reaches approximately as far as posterior margin of prothorax. Both antennae recognised in the fossil specimen.
Prothorax: pronotum weakly transverse, approximately 1.4 times as wide as long; anterior margin straight, not emarginate, anterior corners weakly projecting; lateral edge present, evenly rounded; carina along lateral margin of pronotum absent; posterior margin of pronotum arcuate, posterior corners conspicuous but not projecting.
Mesothorax: mesonotum transverse; mesocoxal cavities narrowly separated; membranous wing present.
Metathorax: metaventrite flat and wide, transverse; metacoxal cavities narrowly separated. Elytron with rounded humeral corner, without carinae.
Legs: mesocoxae distinctly oval, metacoxae extended to lateral margin of metathorax; femora very weakly clavate. Meso- and metatibia and meso- and metatarsus clearly visible: tibia without row of distinct spines along outer margin but with conspicuous edge along outer side; metatibia with two apical spines (not visible in mesotibia); tarsomere 1 conspicuous, slightly shorter than tarsomere 2; tarsomere 4 approximately as long as 3; tarsomere 5 as long as 1–4 combined; tarsal lobes distinctly present in tarsomeres 1–4; claws without denticles; empodium probably small (not observed in well-preserved tarsi); tarsal formula unknown, meso- and metatarsi 5-segmented.
Abdomen: six ventrites distinctly visible; cucujoid tegmen with separated parameres present.
Remarks. —Habitus similar to that of Mauroniscidae ( Fig. 4C View Fig ), Rhadalidae , Dasytidae , and Malachiidae , with tendency to soft-bodiedness. Sparse sculpture of elytra irregular, composed of fine punctures as in members of above mentioned families. Structure of antenna ( Fig. 2B View Fig 4 View Fig ) closely comparable with non-serrate but progressively dilated antennomeres in Mauroniscidae (e.g., Amecomycter Majer, 1995 , Mauroniscus Bourgeois, 1911 , Scuromanius Majer, 1995 ), Dasytidae : Danaceinae (e.g., Danacaea Laporte de Castelnau 1836 , Hylodanacaea Pic, 1926 ), and Dasytidae : Chaetomalachiinae (e.g., Dasytiscus Kiesenwetter, 1859 , and Eutrix Majer, 1996). Antennomeres from fourth or fifth distinctly asymmetrical in dasytids and weakly asymmetrical in mauroniscids (cf. Majer 1994a, 1995, 1996, 1999). Tarsomeres 1–4 with small, non-membranous lobes and long, distinct setae directed forwards as in Prionoceridae and some Dasytidae and Malachiidae . Claws simple, without denticles as in mauroniscids; membranous appendages of claws absent or not preserved. Abdomen with six ventrites as in all members of the melyrid lineage apart from Phycosecidae (ventrites I and II connate in Rhadalidae , see Peacock [1987], but their suture is clearly visible). Tegmen with separated parameres as in mauroniscids ( Majer 1995) and exceptionally also in dasytids ( Constantin and Menier 1987; Majer 1999).
Stratigraphic and geographic range. —NE China: Inner Mongolia, Daohugou; Middle Jurassic, Callovian.
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