Bolbelasmus, Boucomont, 1911
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publication ID |
https://doi.org/ 10.5281/zenodo.5306071 |
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publication LSID |
lsid:zoobank.org:pub:7EFF344A-B2FF-4A14-80AC-3B047298201D |
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DOI |
https://doi.org/10.5281/zenodo.6454369 |
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persistent identifier |
https://treatment.plazi.org/id/2F75BE40-FFA6-BE1C-FEF9-FAB2FBBEF16E |
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treatment provided by |
Marcus |
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scientific name |
Bolbelasmus |
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Identification key for well and medium developed males
1 (10) Scutellar shield impunctate or indistinctly shallowly punctate, frontal horn simple or furcate apically.
2 (5) Frontal horn simple; genae evenly rounded; oblique keel above eyes over whole eyes length; head strongly constricted behind eyes.
3 (4) Frontal horn directed forwards and upwards in lateral view; clypeus with short transversal carina; scutellar shield triangular, approximately as long as wide; body surface brownish, maximal body length 14 mm. Southern parts of Central Europe, Alsace, Italy, Balkan Peninsula, Poland, Russia and Ukraine. ................................... .................................................................... B. unicornis ( Schrank von Paula, 1789)
4 (3) Frontal horn bending not forwards ( Figs 19–20 View Figs 14–20 ); transversal clypeal carina absent; scutellar shield triangular longer than wide; body surface blackish, maximal body length 13 mm ( Figs 6–7 View Figs 5–7 , 38 View Figs 33–44 ); aedeagus as in Fig. 65 View Figs 60–65 ; Algeria, Italy (Sicily) and Tunisia (Fig. 59). ............................................... B. vaulogeri ( Abeille de Perrin, 1898)
5 (2) Frontal horn furcate apically; genae more or less angulate anterolaterally; oblique keel above eyes reaching only to anterior part of eyes; head not strongly constricted behind eyes.
6 (7) Frontal horn short and not longer than broad, horn apex approximately as wide as horn base ( Figs 2 View Figs 1–4 , 15 View Figs 14–20 , 22 View Figs 21–26 ); pronotal punctation denser with exception of basal part, punctures separated approximately by distance equal to their diameters; pronotal base with distinct visible but fine, regularly pinned points; elytral striae well impressed, visible as complete lines; elytral intervals moderately convex; aedeagus as in Fig. 61 View Figs 60–65 ; Spain (Fig. 59). ................................. B. brancoi Hillert & Král sp. nov.
7 (6) Frontal horn long, longer than broad, horn becoming gradually thinner from base to apex ( Figs 1 View Figs 1–4 , 5 View Figs 5–7 , 14, 18 View Figs 14–20 , 21, 25 View Figs 21–26 , 27, 31 View Figs 27–32 , 33, 37 View Figs 33–44 , 57 View Figs 57–58 ); pronotal punctation sparser, punctures separated by approximately three times their diameters; pronotal base with very fine, weakly visible and well separated pinned points; elytral striae not impressed, visible as row of punctures; elytral intervals flat; nothern Africa.
8 (9) Base of frontal horn situated at level of anterior part of eyes; horn sharply keeled from base to half of its length in dorsal aspect ( Figs 1 View Figs 1–4 , 14 View Figs 14–20 , 21 View Figs 21–26 , 27 View Figs 27–32 , 33 View Figs 33–44 ); medial hornlike apophysis of pronotum directed vertically upwards in lateral aspect, subapical protrusion situated basally; lateral hornlike apophyses of pronotum directed forwards and weakly divergent; aedeagus as in Fig. 60 View Figs 60–65 ; Algeria and Morocco (Fig. 59). ............................................................................... B. bocchus ( Erichson, 1841)
9 (8) Base of frontal horn situated at level of posterior part of eyes; horn broadly keeled from base to apex in dorsal view ( Figs 5 View Figs 5–7 , 18 View Figs 14–20 , 25 View Figs 21–26 , 31 View Figs 27–32 , 37 View Figs 33–44 , 57 View Figs 57–58 ); medial hornlike apophysis of pronotum directed obliquely forwards in lateral aspect, subapical protrusion situated near apex; lateral hornlike apophyses of pronotum directed forwards; aedeagus as in Fig 64 View Figs 60–65 ; Egypt, Libya and Tunisia (Fig. 59). ................................................ ............................................... B. nikolajevi Hillert, Arnone, Král & Massa sp. nov.
10 (1) Scutellar shield densely punctate; frontal horn simple apically.
11 (12) Apex of frontal horn approximately as wide as horn base, apex broadly acute-angled, strongly flattened on front part of apex ( Figs 3 View Figs 1–4 , 16 View Figs 14–20 , 23 View Figs 21–26 , 29 View Figs 27–32 , 35 View Figs 33–44 ); aedeagus as in Fig. 62 View Figs 60–65 ; France (including Corsica), Portugal and Spain (Fig. 59). ................................... ....................................................................................... B. gallicus ( Mulsant, 1842)
12 (11) Apex of frontal horn becoming gradually thinner from base to apex, apex acute-angled, only weakly flattened on front part of apex ( Figs 4 View Figs 1–4 , 17 View Figs 14–20 , 24 View Figs 21–26 , 30 View Figs 27–32 , 36 View Figs 33–44 ); aedeagus as in Fig. 63 View Figs 60–65 ; Spain, Gibraltar (Fig. 59). ................ B. howdeni Hillert & Král sp. nov.
Identification key for less developed males
1 (10) Scutellar shield impunctate or weakly shallowly punctate, frontal horn simple or furcate apically.
2 (5) Frontal horn simple; genae regularly rounded; oblique keel above eyes extending over whole eyes length; head strongly constricted behind eyes.
3 (4) Scutellar shield triangular, approximately as long as wide; body surface brownish, maximal body length 12 mm. Southern parts of Central Europe, Alsace, Italy, Balkan Peninsula, Poland, Russia and Ukraine. .... B. unicornis ( Schrank von Paula, 1789)
4 (3) Scutellar shield triangular, longer than wide; body surface blackish, maximal body length 11 mm; aedeagus as in Fig. 65 View Figs 60–65 ; Algeria, Italy (Sicily) and Tunisia (Fig. 59). ....................................................................... B. vaulogeri ( Abeille de Perrin, 1898)
5 (2) Frontal horn furcate apically; genae more or less angulate anterolaterally; oblique keel above eyes reaching only to anterior part of eyes; head not strongly constricted behind eyes.
6 (7) Pronotal punctation denser with exception of basal part, punctures separated approximately by distance equal to their diameters; pronotal base with distinct visible but fine, regularly spaced pinned points, elytral striae well impressed, visible as complete lines; elytral intervals moderately convex; aedeagus as in Fig. 61 View Figs 60–65 ; Spain (Fig. 59). ...................................................................... B. brancoi Hillert & Král sp. nov.
7 (6) Pronotal punctation sparser, punctures separated by approximately three times their diameters pronotal base with very fine, weakly visible and well separated pinned points; elytral striae not impressed, visible as row of punctures; elytral intervals flat; northern Africa.
8 (9) Base of frontal horn situated at level of anterior part of eyes; horn sharply keeled basally only; aedeagus as in Fig. 60 View Figs 60–65 ; Algeria and Morocco (Fig. 59). ....................... ...................................................................................... B. bocchus ( Erichson, 1841)
9 (8) Base of frontal horn situated at level of posterior part of eyes; horn broadly keeled from base to apex in dorsal view; aedeagus as in Fig. 64 View Figs 60–65 ; Egypt, Libya and Tunisia (Fig. 59). ............................... B. nikolajevi Hillert, Arnone, Král & Massa sp. nov.
10 (1) Scutellar shield densely punctate; frontal horn simple apically.
11 (12) Apex of frontal horn flattened on front part of apex; aedeagus as in Fig 62 View Figs 60–65 ; France (including Corsica), Portugal and Spain (Fig. 59). ....... B. gallicus ( Mulsant, 1842)
12 (11) Apex of frontal horn regularly arcuate, not flattened on front part of apex; aedeagus as in Fig. 63 View Figs 60–65 ; Spain, Gibraltar (Fig. 59). ............ B. howdeni Hillert & Král sp. nov.
Identification key for females
1 (10) Scutellar shield impunctate, or weakly shallowly punctate.
2 (5) Frontal tubercle simple ( Figs 41, 42, 44 View Figs 33–44 ); genae regularly rounded; oblique keel above eyes over whole length of eyes; head strongly constricted behind eyes.
3 (4) Scutellar shield triangular, approximately as long as wide; body surface brownish, minimal body length 12 mm. Southern parts of Central Europe, Alsace, Italy, Balkan Peninsula, Poland, Russia and Ukraine. .... B. unicornis ( Schrank von Paula, 1789)
4 (3) Scutellar shield triangular longer than wide; body surface blackish, minimal body length 11 mm ( Fig 13 View Figs 12–13 ); Algeria, Italy (Sicily) and Tunisia (Fig. 59). ........................ ....................................................................... B. vaulogeri ( Abeille de Perrin, 1898)
5 (2) Frontal tubercle furcate apically ( Figs 39–40, 43 View Figs 33–44 ); genae more or less angulate anterolaterally; oblique keel above eyes reaching only to anterior part of eyes; head not strongly constricted behind eyes.
6 (7) Pronotal punctation denser with exception of basal part, punctures separated approximately by distance equal to their diameters; pronotal base with distinct visible but fine, regularly pinned points; elytral striae well impressed, visible as complete lines; elytral intervals moderately convex ( Fig. 9 View Figs 8–11 ); Spain (Fig. 59). ................................... ............................................................................. B. brancoi Hillert & Král sp. nov.
7 (6) Pronotal punctation sparser, punctures separated by approximately three times their diameters pronotal base with very fine weak visible and well separated pinned points; elytral striae not impressed, visible as row of punctures; elytral intervals flat; northern Africa.
8 (9) Base of frontal tubercle situated at level of anterior part of eyes; tubercle sharply keeled basally only in dorsal view ( Fig. 8 View Figs 8–11 ); Algeria and Morocco (Fig. 59). ............ ...................................................................................... B. bocchus ( Erichson, 1841)
9 (8) Base of frontal tubercle situated at level of posterior part of eyes; horn broadly keeled from base to apex in dorsal view ( Fig. 12 View Figs 12–13 ); Egypt, Libya and Tunisia (Fig. 59). ...... ............................................... B. nikolajevi Hillert, Arnone, Král & Massa sp. nov.
10 (1) Scutellar shield densely punctate.
11 (12) France (including Corse), Portugal and Spain (see distribution map on Fig. 59 for details). ......................................................................... B. gallicus ( Mulsant, 1842)
12 (11) Spain, Gibraltar (see distribution map on Fig. 59, for details). .................................. ............................................................................ B. howdeni Hillert & Král sp. nov.
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