Stauroderus I. Bolívar, 1897

Sergeev, M. G., Storozhenko, S. Yu. & Benediktov, A. A., 2019, An annotated check-list of Orthoptera of Tuva and adjacent regions. Part 2. Suboder Caelifera. Tridactylidae, Tetrigidae, Acrididae: Melanoplinae, Calliptaminae, and Gomphocerinae (except Gomphocerini), Far Eastern Entomologist 389, pp. 7-44 : 32-33

Genus Stauroderus I. Bolívar, 1897

36(58). Stauroderus scalaris (Fischer de Waldheim, 1846)

Stenobothrus morio: Miram, 1907: 5 .

Stauroderus scalaris: Berezhkov, 1951: 19 ; Ivanova, 1967: 132; Sergeev, 1986: 207; Benediktov,

1997: 117; Vedenina & Bukhvalova, 2001: 103; Tishechkin & Bukhvalova, 2009: 27; Benediktov,

Belyaev 2019: 23.

Stauroderus scalaris scalaris: Berezhkov, 1956: 108 .

MATERIAL. Tuva: W Sayan Mts., Kurtushibinskij Range, 52°16ʹ N, 93°41ʹ E, 1429 m, alpine meadows, 13.VIII 2018, 2 ♂ ( MS); Turan-Uyuk Intermountain Basin, Begreda River , 51°59ʹN, 94°18ʹE ,

829 m, terrace, meadow, 12.VIII 2018, 1 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu (Khapto) River , 51°57ʹN, 95°33ʹE , 1945–2005 m, mountain tundra with Betula and alpine meadow, 10.VIII 2018, 1 ♂, 1 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu

(Khapto) River, 51°56ʹN, 95°30ʹE, 1737–1869 m, southern slope, terraces, flood-plains, meadows, 9–

10.VIII 2018, 7 ♂, 2 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu (Khapto)

River, 51°55ʹ N, 95°2ʹ E, 1432 m, terraces and flood-plain, meadows, 10.VIII 2018, 1 ♂ ( MS); E Tuva,

Academician Obruchev Range, southern slope, Koptu (Khapto) River, 51°52ʹN, 95°26ʹE, 1230 m,

terrace, meadow, 10.VIII 2018, 9 ♂, 1 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope,

Koptu (Khapto) River, 51°50ʹN, 95°25ʹE, 1132–1135 m, terrace and flood-plains, meadows, 11.VIII

2018, 6 ♂, 5 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu (Khapto) River ,

51°44ʹ N, 95°26ʹ E, 958 m, upper terrace, steppe, 11.VIII 2018, 4 ♂, 2 ♀ ( MS); E Tuva, Academician

Obruchev Range, southern slope, Koptu (Khapto) River, 51°44ʹN, 95°26ʹE, 1006 m, southern slope,

stony steppe, 11.VIII 2018, 1 ♂ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu

(Khapto) River, 51°37ʹN, 95°21ʹE, 788–843 m, southern slope and upper terraces, steppes, 11.VIII 2018,

4 ♂, 7 ♀ ( MS); E Tuva, Academician Obruchev Range, southern slope, Koptu (Khapto) River , 51°37ʹN ,

95°21ʹE, 781–784 m, lower terrace and flood-plain, meadow, 11.VIII 2018, 9 ♂, 4 ♀ ( MS); 16 km N

Boyarovka settlement, 15.VII 2013, 4 ♂, 4 ♀ (M. Prostshalykin, V. Loktionov); 14 km SW Shagonar

City, Shagonar River, 51°27ʹN, 92°45ʹE, 583 m, lower terrace, meadow between bushes, 17–18.VI 2017,

34 ♂, 32 ♀, 17 larvae ( MS); 14 km SW Shagonar City, Shagonar River , 51°27ʹN, 92°45ʹE, 580 m, upper flood-plain, pebbles and sand, 20.VI 2017 , 1 ♂ ( MS); 14 km SW Shagonar City, Shagonar River ,

51°27ʹN, 92°45ʹE, 583 m, upper terrace, dry meadow, 17.VI 2017, 6 ♂, 5 ♀, 5 larvae ( MS); 14 km SW

Shagonar City, Shagonar River, 51°28ʹN, 92°44ʹE, 610 m, steppe with Stipa , 23.VI 2017, 5 ♂, 4 ♀, 5

larvae ( MS); 14 km SW Shagonar City, Shagonar River, 51°27ʹN, 92°45ʹE, 580–582 m, dry meadow,

23–26.VI 2017, 2 ♂, 7 ♀, 4 larvae ( MS); 14 km SW Shagonar City, Shagonar River , 51°28ʹN, 92°45ʹE ,

585–600 m, semi-desert and dry steppe, 25–26.VI 2017, 5 ♂ ( MS); 6 km SE Baj-Haak settlement ,

Sosnovka settlement, 51°08ʹN, 94°32ʹE, slope of balka with steppe, 21.VII 2014, 2 ♂, 2 ♀ ( SS); 6 km SE

Baj-Haak settlement, Sosnovka settlement, 51°08ʹN, 94°32ʹE, plakor, mixed steppe, 20.VII 2014, 1 ♂, 3 ♀

( SS); SE Ulug-Khem Intermountain Basin, Uzun-Kharagan River, near Balgazyn settlement, 50°57ʹN,

95°16ʹE, 940 m, piedmont plain, balka, meadow between larches, 8.VII 1978, 1 ♂ ( MS); SE UlugKhem Intermountain Basin, Uzun-Kharagan River , near Balgazyn settlement, 50°57ʹN, 95°16ʹE, 940 m ,

upper terrace, meadow with Caragana bushes, 8.VII 1978, 1 ♂ ( MS); SE Ulug-Khem Intermountain

Basin, Uzun-Kharagan River, near Balgazyn settlement, 50°57ʹN, 95°15ʹE, 886 m, lower terrace,

meadow, 8.VII 1978, 1 ♂ ( MS); SE Ulug-Khem Intermountain Basin, Shuurmak River , left side, 15 km

S Balgazyn settlement, 50°53ʹN, 95°12ʹE, 900–950 m, piedmont plain of E Tannu-Ola Mts., meadow,

9.VII 1978, 1 ♂, 1 ♀ ( MS); E Tannu-Ola Mts. , Shuurmak River, 5 km N Shuurmak settlement, 50°42ʹN ,

95°20ʹE, 1086 m, lower terrace, meadow, 11.VII 1978, 1 ♂ ( MS); Shuurmak River near Shuurmak settlement, 1170 m, 12.VII 2011, 5 ♂ (M. Prostshalykin, V. Loktionov); E Tannu-Ola Mts., Shuurmak

Pass, 50°38ʹN, 95°11ʹE, 1492 m, short meadow, 5.VII 2017, 5 ♂, 9 ♀ ( MS); E Tannu-Ola Mts. , 13 km

NEE Samagaltaj settlement, Kaldak-Khamar (Shuurmak) Pass, 50°38ʹN, 95°11ʹE, 1550 m, open larch forest, 19.VII 2014, 1 ♂, 1 ♀ ( SS); the same locality, 1500 m, rangeland, 19.VII 2014, 2 ♂, 1 ♀ ( SS); the same locality, 1500 m, 11.VII 2013, 3 ♂ (M. Prostshalykin, V. Loktionov); NE Uvs-Nuur Intermountain

Basin, Shivelig River , 50°45ʹN, 94°34ʹE, 1280–1301 m, terraces, dry meadows, 8.VII 2017, some specimens observed ( MS); Dyttyg-Khem River, 12 km SW Samagaltai settlement, 50°38ʹ N, 95°19ʹ E ,

17.VII 2014, 1 ♂ ( SS); Dyttyg-Khem River , 13 km SW Samagaltai settlement , 6–10.VII 2013, 5 ♂, 1 ♀

(M. Prostshalykin, V. Loktionov); Uvs-Nuur Intermountain Basin, S Tore Lake , 50°01′N, 95°04′E ,

1154-1166 m, 07.VII 2017, sand semi-desert and upper terraces with birch forest, 4 ♂, 2 ♀ ( MS); UvsNuur Intermountain Basin, S Tore Lake , 50°02′N, 95°03′E , 1152 m, flood-plain, meadow, 07.VII 2017,

3 ♂ ( MS); 31 km NEE Erzin settlement, Erzin River , 50°21ʹN, 95°34ʹE, 1100 m, flood-plain, meadow near cliff, 18.VII 2014 , 2 ♂, 1 ♀ ( SS); 25 km NEE Erzin settlement, Belyj Medved Mt. , 50°21ʹN ,

95°27ʹE, steppe, 16.VII 2014, 2♂ ( SS). Krasnoyarsk Region: southern part, West Sayan Mts. , Us

Intermountain Basin, near abandoned Idzhim settlement, 52°21′N, 93°13′E, 780–800 m, southern slope,

dry mountain meadow and steppe, 22.VII 1995, 8 ♂, 11 ♀ ( MS); the same locality, 52°20′N, 93°20′E ,

700–750 m, low terrace, meadow, 22.VII 1995, 1 ♀ ( MS); the same locality, 52°16′N, 93°07′E , 680–700

m, terraces and southern slope, steppes, 22–23.VII 1995, 3 ♀ ( MS) .

DISTRIBUTION. Tuva: WSE, ET, KHE, UKH, KKH, ETO, UVS, SAN. – Europe

(except the North), S Siberia (up to Buryatia); Asia Minor, Caucasus, Kazakhstan, Tien

Shan, Pamiro-Alay, NW China, Mongolia [nominotypical subspecies]; Azerbaijan [ssp.

znojkoi (Miram, 1938)], NW Iran [ ssp. demavendi Popov, 1951 ].

ECOLOGY. Common species (especially during the last decade) in the steppes of the central part of Tuva. Often associated with dry meadows with high vegetation.