Haematotrephus kossacki Witenberg, 1923

Dronen, Norman O. & Blend, Charles K., 2015, Updated keys to the genera in the subfamilies of Cyclocoelidae Stossich, 1902, including a reconsideration of species assignments, species keys and the proposal of a new genus in Szidatitreminae Dronen, 2007, Zootaxa 4053 (1), pp. 1-100 : 55

H. kossacki ( Witenberg, 1923) n. comb.

Type host. Dunlin, Calidris alpina (Linnaeus) (Syn. Tringa alpina Linnaeus ) ( Charadriiformes : Scolopacidae )— Chertkova et al. (1994).

Type locality. Europe; specific type locality unknown; Germany—Chertkova et al. (1994).

Additional hosts. Spotted redshank, Tringa erythropus (Pallas) ; common redshank, Tringa totanus (Linnaeus) ( Charadriiformes : Scolopacidae ) – Murai et al. (1983).

Additional locality. Hungary—Murai et al. (1983).

Previously proposed synonyms. Cyclocoelum nebularium Khan, 1935 (= H. nebularium [ Khan, 1935] n. comb.), Haematotrephus lanceolatum of Stossich, 1902 nec. Wedl, 1858 of Bychowskaja- Pawlowskaja (1953) — Dubois (1959).

Remarks. This species was originally described as Corpopyrum kossacki Witenberg, 1923 (& 1926) and was considered to be the type species for Corpopyrum (= Haematotrephus ) by Witenberg (1923) and Yamaguti (1958, 1971). Macko & Feige (1960) considered this species to be a synonym of Haematotrephus lanceolatum ( Wedl, 1858) and Joyeux & Baer (1927) considered Corpopyrum kossacki to be Cyclocoelum kossacki ( Witenberg, 1923) . It has a pretesticular ovary that forms a triangle with the testes (Haematotrephinae), the genital pore is located near the mid-level of the pharynx (“The sex opening is median, situated high at the mid-level of the pharynx”— Witenberg 1926; likely postpharyngeal) and the vitelline fields are not confluent posteriorly, placing this species in Haematotrephus . Yamaguti (1971, Fig. 1135) illustrated the genital pore as being prepharyngeal, while in Fig. 152 of Bashkirova (1950) it appears to be located more posteriorly(postpharyngeal). Until these inconsistencies are resolved we have retained this species in Haematotrephus . Rudimentary oral sucker appears to be present (“The mouth is subterminal and has the shape of a muscular funnel”— Witenberg 1926).