Chassalia javanica (Blume) I.M.Turner

Yu, Tian Yi, Turner, Ian M. & Cheek, Martin, 2021, Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species, European Journal of Taxonomy 738, pp. 1-60 : 26-27

Chassalia javanica (Blume) I.M.Turner

Feddes Repertorium 130: 398 ( Turner 2019). – Coffea javanica Blume, Catalogus van eenige der merkwaardigste zoo in- als uit-heemsche gewassen, te vinden in ‘s Lands Plantentuin te Buitenzorg opgemaakt door C. L. Blume, M.D., Directeur van voorz. tuin: 46 ( Blume 1823). – Chiococca javanica (Blume) Blume, Bijdragen tot de flora van Nederlandsch Indië 16: 958 ( Blume 1826). – Proscephaleium javanicum (Blume) Korth., Nederlandsch Kruidkundige Archief 2 (2): 248 ( Korthals 1851). – Type: INDONESIA • Java, Tjapus; C.L. Blume s.n.; hololectotype: L[ L0057991 ], designated by Turner (2019: 398).

Additional material

MALAYSIA • Borneo, Gommisi Kap, Genderen Stort , G. Djempanga ; Aug. 1912; Amdjah 729; K ! • Borneo, Sarawak, Bintulu, Sungai Penyilam , Pengkalan Resam ; 1 Oct. 2006; Julia S., Tan H.S., Malcom D., Rantai J. et al. S 97155 ; K!, SAR .

Description

Epiphytic shrub, glabrous. Stem terete, slightly canaliculate when dry. Internodes broadening slightly at base and apex, (0.5–)2–4(–6) cm × 2–3 mm. Leaf-blades obovate to elliptic, 9–17 × 2–4.5 cm, apex acute to acuminate, acumen 5 mm long if acuminate, base truncate, midrib raised adaxially and abaxially, secondary nerves 8–11 on each side of the midrib, petioles 0.5–3 cm long. Stipule broadly triangular, 4–5 × 5 mm, sometimes bilobed when broken, margin with terminal colleters, inside with standard colleters 0.5–1 mm long. Inflorescence a compound cyme, 4.5–7 cm long, peduncle 0.8–4 cm long, internodes of rachis 0.5–1.5 cm long, each node of inflorescence including capitulum with 4 bracts, triangular, margin with teeth, 2 broader bracts 0.5–1.5 × 0.5–1 mm, 2 narrower bracts 1.5 × 0.5 mm. Open flowers not seen. Fruits narrowly ellipsoid, 5–6 × 4 mm, with pedicels 5–6 mm long. Pyrene dorsal surface rough, with a central longitudinal groove 1.5–2 mm wide, pyrene ventral surface with a central groove 1.5–2 mm wide and a ridge at centre of the groove 0.5 mm wide and as high as the plane of the endocarpal outgrowth, the 2 grooves between endocarpal outgrowth and pyrene margin 0.1 mm wide or absent. Seed not seen.

Distribution, habitat

Malaysia, Borneo; Indonesia, Kalimantan, Java, Sumatra. Epiphytic in lowland to montane forest.

Remarks

Differs from other Asian species of Chassalia by usually being epiphytic, by having narrowly ellipsoid fruits only 5–6 mm long, pyrenes with dorsal surfaces bearing a longitudinal groove but without a crest, and ventral surfaces with only 1 ridge, reaching as high as the plane of the endocarpal outgrowth.