Codoserphus calophlebius, Shi, Xiaoqing, Zhao, Yunyun, Shih, Chungkun & Ren, Dong, 2013

Shi, Xiaoqing, Zhao, Yunyun, Shih, Chungkun & Ren, Dong, 2013, New fossil mesoserphid wasps (Insecta, Hymenoptera, Proctotrupoidea) from the Jehol Biota, China, Zootaxa 3710 (6), pp. 591-599 : 593-596

Codoserphus calophlebius sp. nov.

( Fig. 1 A, B and Fig. 3 A, B)

Diagnosis. Fore wing 1-Rs as long as 1-M; 2cu-a anteriorly bent inwards, and adjacent 1m-cu in line.

Description. Holotype male, body length about 7.7 mm, relatively larger than most other genera in the family (see Table 2 in Shih, Feng et Ren, 2011). Head oval and slightly narrower than mesosoma(head width = 1.64 mm, mesosoma width = 1.88 mm). Eyes large and possess two third of the head space from dorsal view. Antenna with at least 20 antennomeres (14 left and 20 right preseved), longer than head and mesosoma combined, reaching middle of the third metasomal segment, and paler than head. Scape swollen and bell-shaped; pedicel small quadrate; first antennomere long, longer than pedicel and scape combined; subsequent flagellomeres gradually decreasing in length from base to apex with the terminal segment shortest.

Fore wing with all veins present ( Fig. 1 A, B) and with seven cells enclosed by tubular veins. 2r-rs arising from about half of pterostigma, slightly oblique apicad, 1.3 times as long as pterostigmal width. C long and extending to apex of fore wing. R robust, cell r closed and triangular; Rs straight and reaching distal C. Cell 1+2r six-sided and surrounded by R, 1-Rs, 1-Rs+M, 2-Rs+M, 2-Rs, and 2r-rs. M+Cu straight and distinct; M and Cu straight in base, slightly curved in apex. 1-Rs as long as 1-M. Cell 1mcu trapezoid and smaller than cua, with 1-M about two times as long as 1m-cu; both 1cu-a and 2cu-a present; 1cu-a slightly distal to 1-M; 2cu-a anteriorly bent inwards and meeting 1m-cu in line.

Hind wing with tubular veins C, R and M preserved, but cell r absent.

Mesosoma slightly broader than head; pronotum short dorsally with length of 0.92 mm and width about 0.2 mm, narrower than head; mesoscutum subquadrate with length of 1.16 mm and width, 1.0 mm; tegula large and triangular, the length of tegula approximately as long as that of fore wing vein 1-M; propodeum areolate.

Metasoma bell-shaped from dorsal view, number of metasomal segments at least six; first to third segments trapezoidal and approximately equal in length, fourth to sixth segments shorter than the former and from trapezoidal to rectangular, the sixth segment trapezoidal and blunt apically.

Fore legs with only part of femur preserved, color paler than pronotum; mid legs with partial femur and tibia, not well preserved; right hind leg with large suboval coxa, part of femur, tibia and tarsus preserved; left hind leg with partial coxa, femur spindle-shaped, narrow basally and gradually widened apically, tibia long and swollen distally, and tarsi partially preserved.

Etymology. The specific name is a Greek word means “good wing venation”.

Measurements (in mm). Head length 1.15, width 1.64; mesosoma length 2.97, width 1.88; Metasoma length 3.55; lengths of metasomal segments are 0.55, 0.66, 0.66, 0.52, 0.42 and 0.52; body length 8.3; length of right antenna 6.06 (length of right flagellomeres are as follows, F1: F2: F3: F4…F19=0.43: 0.53: 0.38: 0.40: 0.37: 0.33: 0.30: 0.30: 0.33: 0.30: 0.30: 0.27: 0.27: 0.23: 0.21: 0.20: 0.16: 0.13: 0.12, with F1 partially covered by head), length of left antenna 5.45; fore wing length 4.85, width 2.12.

Type Materials. Holotype, male, CNU-HYM-LB2012056. A well-preserved almost complete body with antenna, both wings and parts of legs ( Fig. 1 A, B).

Paratypes, CNU-HYM-LB2012058 p/c. Almost complete body with antenna, both wings and parts of legs, but not well preserved ( Fig. 3 A and B).

Figures

FIGURE 1. Holotype of Codoserphus calophlebius Shi, Shih et Ren, gen. et sp. nov. CNU-HYM-LB 2012056. A. Photo. B. Line drawing. Scale bars = 1 mm. (Online figure in color.)

FIGURE 3. Paratypes of Codoserphus calophlebius Shi, Shih et Ren, gen. et sp. nov. A. CNU-HYM-LB 2012058 p.; B. CNU- HYM-LB 2012058 c; C – E. Paratypes of Codoserphus euneurus Shi, Shih et Ren, gen. et sp. nov.; C. CNU-HYM-LB 2012051; D. CNU-HYM-LB 2012052; E. CNU-HYM-LB 2012053. Scale bars = 1 mm. (Online figure in color.)