Themus (Haplothemus) mishmiensis Wittmer, 1973

Yang, Yuxia, Liu, Haoyu & Yang, Xingke, 2018, A contribution to the knowledge of Themus (Haplothemus) Wittmer from China (Coleoptera: Cantharidae), Zootaxa 4407 (2), pp. 241-253: 247-250

publication ID

publication LSID

persistent identifier

treatment provided by


scientific name

Themus (Haplothemus) mishmiensis Wittmer, 1973

stat. nov.

Themus (Haplothemus) mishmiensis Wittmer, 1973  stat. nov.

Figs 4View FIGURES 1–6, 16‒18

Themus (Haplothemus) elongatior mishmiensis Wittmer, 1973: 191  , 210, figs 25, 26.

Type material examined. Paratype: 1♂ ( NHMBAbout NHMB): [p] “ Assam: \ Mishmi Hills, \ 10000ft 12-vi-1923, \ Percy Sladen Expn. \ B. M. 1929-531,”, [h] “28°10ʹN 96°37ʹE”, [p] “ PARATYPUS ”, “ Themus  \ elongatior  ssp. \ mishmiensis  \ Wittm. \ det. W. Wittmer”, [p] “Naturhist. \ Museum Basel \ coll. W. Wittmer”, [p] “ CANTHARIDAE  \ CANTH 00001206”.

Other material examined. CHINA, Xizang: one male ( MHBU), Zayü, 17.VII.2005, leg. A.M. Shi  ; one male (MHBU), Mêdog, Lage, 3300m, 8.VIII.2003, leg. G.D. Ren.

Distribution. China (new record: Xizang); India.

Redescription. Male ( Fig. 4View FIGURES 1–6). Body orange brown, head darkened at anterior part of dorsum, mandibles darkened at apices, pronotum slightly darkened at both sides of disc, legs slightly darkened at tarsi, abdominal ventrites except the terminal one each side with a small round black marking. Body densely covered with decumbent orange brown pubescence, except those black on elytra and a few long hairs along anterior margin of clypeus.

Head rounded, sparsely and finely punctate, surface lustrous; eyes moderately protruding, head width across eyes slightly wider than anterior margin of pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal two-fifths length, with outer margins slightly arcuate, inner apical margins slightly sharp and apices rounded; antennae slightly thickened, almost extending to elytral mid-length, antennomeres II about three times as long as wide, III distinctly shorter than II, IV about 1.5 times as long as III, V longest, V‒X gradually shortened and narrowed one by one, XI slightly shorter than X and pointed at apices, IV‒X each with a small round or oblong impression at apical part or near middle of outer margin.

Pronotum subquadrate, about 1.4 times as wide as long, anterior margin arcuate, lateral margins slightly diverging posteriorly, posterior margin nearly straight and narrowly bordered, anterior angles nearly rectangular, posterior angles rounded, disc convex at postero-lateral parts, sparsely and finely punctate like that on head, surface matt.

Elytra about 6.0 times longer than pronotum, 3.3 times as long as humeral width, with lateral margins nearly parallel, disc rugulose-lacunose and coarsely punctate, surface matt.

Aedeagus (Figs 16‒18): ventral process of each paramere moderately bent inwards in ventral view, abruptly narrowed at base, distinctly protuberant at basal part of outer margin, longitudinally depressed along the whole length of ventral side, rounded and slightly turned ventrally at apex; conjoint dorsal plate of parameres slightly shorter than ventral processes, narrowly emarginated in middle of apical margin, lateral angles widely rounded and around with long hairs; laterophyses distinctly shorter than conjoint dorsal plate, situated on dorsal side of median lobe, conjoint and depressed, widely and triangularly emarginated in middle of apical part, with latero-apical part acute at apex and slightly bent dorsally.

FIGURES 16–24. Aedeagus (16, 19, 22. ventral vieW; 17, 20, 23. dorsal vieW; 18, 21, 24. lateral vieW): 16‒18. Themus (Haplothemus) mishmiensis Wittmer, 1973  ; 19‒21. T. (H.). weishanensis Kopetz, 2016  ; 22‒24. T. (H.) longideverticulum  sp. nov. Scale bars: 1.0 mm.

Remarks. Except the appearance, the structure of the adeagus of T. (H.) mishmiensis  is obviously different from that of T. (H.) elongatior  . Thus, we suggest place T. (H.) mishmiensis  in the specific status. It is reported from China for the first time.


Naturhistorisches Museum, Basel














Themus (Haplothemus) mishmiensis Wittmer, 1973

Yang, Yuxia, Liu, Haoyu & Yang, Xingke 2018


Themus (Haplothemus) elongatior mishmiensis

Wittmer 1973: 191