Themus (Haplothemus) quadratus Wittmer, 1983

Yang, Yuxia, Liu, Haoyu & Yang, Xingke, 2018, A contribution to the knowledge of Themus (Haplothemus) Wittmer from China (Coleoptera: Cantharidae), Zootaxa 4407 (2), pp. 241-253: 242-244

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Themus (Haplothemus) quadratus Wittmer, 1983


Themus (Haplothemus) quadratus Wittmer, 1983 

Figs 1View FIGURES 1–6, 7‒9View FIGURES7–15, 25View FIGURES25–29, 30View FIGURES 30–34

Themus (s.str.) quadratus Wittmer, 1983: 224  , fig. 112. Themus (Haplothemus) quadratus: Švihla, 2008: 185  .

Type material examined. Holotype: one female ( IZAS): [p] “洗象池 \ 1800-2000公尺 \ 采集者: 朱复兴”, [p] “四 川: 峨 眉山 \ 1957. VIII.28 \ 中 国科学院 ”, [p] “ HOLOTYPUS ”, [p] “IOZ (E) 217899”. [one female, China, Sichuan, Emei Shan, Xixiangchi , 1800‒2000 m, 28.VIII.1957, leg. F.X. Zhu]  .

Other material examined. CHINA, Gansu: three males, three females ( MHBU): Wenxian, Huangtuling , 2350m, 8.VII.2003, leg. Y.B. Ba & Y. Yu. 

Distribution. China (Sichuan, Gansu).

Redescription. Male ( Fig. 1View FIGURES 1–6). Head dark brown, black in middle of clypeus and both sides of vertex on dorsum, mouthparts dark brown, darkened at apices of mandibles, antennae black, prothorax dark brown, pronotum with two large irregular black markings on disc, markings more or less conjoint together in middle, sometimes extending to anterior margin but never to posterior or lateral margins, scutellum dark brown, elytra black, legs dark brown, slightly darkened at apical parts of femora and tibiae and tarsi, ventral surface dark brown, black at both sides of each abdominal ventrite except the terminal one, sometimes more or less darkened at meso- and metasterna. Body densely covered with decumbent brown pubescence, mixed with slightly long, semi-erected pubescence on elytra and few long hairs along anterior margin of clypeus.

Head rounded, sparsely and finely punctate, surface lustrous; eyes moderately protruding, head width across eyes slightly wider than anterior margin of pronotum; terminal maxillary palpomeres nearly long-triangular, widest at basal two-fifths length, with outer margins slightly arcuate, inner apical margins slightly sharp and apices acute; antennae slightly thickened, almost extending to basal two-thirds length of elytra, antennomeres II about twice as long as wide, III slightly shorter than II, IV about 1.5 times as long as III, V longest, V ‒X gradually shortened and narrowed one by one, XI slightly shorter than X and pointed at apices, IV ‒VIII each with a small round or oblong impression near middle of outer margin.

Pronotum subquadrate, about 1.2 times as wide as long, anterior margin arcuate, lateral margins nearly parallel, posterior margin nearly straight and narrowly bordered, anterior angles nearly rectangular, posterior angles rounded, disc convex at postero-lateral parts, sparsely and finely punctate like that on head, surface matt.

Elytra about 5.5 times longer than pronotum, 3.0 times as long as humeral width, with lateral margins nearly parallel, disc rugulose-lacunose and coarsely punctate, surface matt.

Aedeagus ( Figs 7‒9View FIGURES7–15): ventral process of each paramere narrow, nearly straight and nearly pointed at apex; conjoint dorsal plate of parameres slightly shorter than ventral processes, with lateral margins nearly parallel, widely and triangularly emarginated in middle and slightly sinuate on both sides of apical margin, latero-apical angles slightly protuberant and sub-rounded; laterophyses depressed and slightly shorter than conjoint dorsal plate, separated on both sides of median lobe and distinctly diverging from each other, acute and slightly hooked at apices, which pointed towards latero-apical angles of conjoint dorsal plate.

Female. Similar to male, but antennae slightly thinner and shorter, nearly extending to elytral mid-length, IV ‒VIII without impressions; pronotum slightly wider, about 1.3 times as wide as long; elytra with lateral margins slightly diverging posteriorly.

Abdominal sternite VIII ( Fig. 30View FIGURES 30–34) triangularly emarginated in middle and bi-sinuately emarginate on both sides of posterior margin, the lateral emarginations both slightly shallower than the middle one, the anterior lateral emargination slightly wider than the posterior one, the portion between the two lateral emarginations nearly rectangular at apex, and that between the middle and posterior lateral emarginations acute at apex.

Internal genitalia ( Fig. 25View FIGURES25–29): vagina stout, with diverticulum and spermatheca situated at apex, median oviduct at basal one-third part of ventral side; diverticulum moderately long and thick sac-shaped, slightly shorter than vagina, distinctly expanded at apical half part, about twice as long as maximal width, rounded at apex; spermatheca short sac-shaped, distinctly shorter than diverticulum, distinctly expanded apically and rounded at apex, and provided with a moderately long and thin accessory gland at base; diverticulum and spermatheca surrounded with a strongly sclerotized ring at base, which is confluent in middle and extending to median oviduct on ventral side, the opening of spermatheca arising from middle of the sclerotized ring on dorsal side.

Body length (both sexes): 17.0‒18.0 mm; width: 3.5‒4.5 mm.

Remarks. The male of T. (H.) quadratus Wittmer, 1983  , which was described on a single female specimen in the original manuscript, is discovered and described here for the first time. At the same time, a related species is recognized and described under the name of T. (H.) quadratiformis  sp. nov.


Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine














Themus (Haplothemus) quadratus Wittmer, 1983

Yang, Yuxia, Liu, Haoyu & Yang, Xingke 2018

Themus (Haplothemus) quadratus: Švihla, 2008: 185

Svihla 2008: 185

Themus (s.str.) quadratus

Wittmer 1983: 224