Cidariplura maraho Wu & Owada, 2013

Wu, Shipher, Owada, Mamoru & Wang, Min, 2019, Review of Cidariplura Butler, 1879 (Lepidoptera, Erebidae, Herminiinae). Part 1: the Cidariplura gladiata species complex, Zootaxa 4668 (4), pp. 489-502 : 496

Cidariplura maraho Wu & Owada, 2013

( Figs 15, 16 , 33 , 42 , 52 )

Cidariplura maraho Wu & Owada, in Wu et al., 2013: 151 , figs 14–16, 41, 42, 62, 73, 82.

Type material. Holotype, ♂, Taiwan, Nantou County, Meifeng , 2,100 m, 29. VI. 2012, TFRI147244 S. Wu & W. C. Chang leg. ( TFRI) ( Fig. 15 ) . Paratypes (9♂ 4♀): the same collecting locality as that of holotype, 1♂, 18. VII. 1990, TFRI00010165 , Y. C. Chang leg.; the same locality, 3♂, 20. VII. 2011, TFRI00128724 ♂, S. Wu & W. C. Chang leg. ( TFRI) ; Hualien, Ci’en , 1,950 m, 2♂, 28. VI. 2011, S. Wu & W. C. Chang leg.; the same locality, 1♂, 18. VII. 2011, S. Wu & W. C. Chang leg. ( TFRI) ; Pilu-Shenmu , 2,000 m, 1♀, 16.VII. 2012, M. Owada & L. C. Shih leg. ( ESRI) ; Nantou, Biluxi, 1♂, 12. VII. 2011, C. M. Fu leg.; Turnyuan , 1♀, 23. VI. 2007, 1,950 m, C. M. Fu leg. ( NMNS) ; Taichung, Wuling , 1,850 m, 1♀, 10–12. VIII. 1990, M. Owada leg. ( NSMT) ; Hualien, Tsu’en , 1,990 m, 1♂, 26. VI. 1989, M. Owada leg. ( NSMT) ; Nantou, Hotso [Lushan Spa], 1♀, 26–29. VI. 1973, M. Owada leg. ( NSMT) .

Additional material examined (2♂ 2♀). Hualien, Tsu’en , 2,000 m, 1♀, 13. VII. 2015, NSMT3284 ♀, M. Owada & L. Shih leg. ( NSMT) ; Nantou, Sunlinksea , 1,700 m, 1♂, 25. VI. 2017, NSMT3283 ♂, M. Owada & L. Shih leg. ( NSMT); Kaohsiung, Tianchi-2, 2,280 m, 1♂ 1♀, 6. VII. 2015, M. Owada & C.-M. Fu leg. ( NSMT) .

Diagnosis. The species is easily distinguished from other species in C. gladiata complex due to the paler ground coloration of the wings and the more contrasting forewing medial region, the less curved transversal lines on both wings, and the medial part of the corpus bursae, which is incised with longitudinal wrinkles.

Distribution and phenology. Endemic to Taiwan. The adults occur from June to August.

Figures

FIGURE 1–22. Habitus of Cidariplura gladiata complex. 1. C. gladiata, male, NSMT3285, Honshu, Japan (NSMT); 2. Ditto, female, NSMT3286, Honshu, Japan (NSMT); 3. Ditto, male, NSMT3301, Iriomote Is., Japan (NSMT); 4. Ditto, female, NSMT3299, Ishigaki Is., Japan (NSMT); 5. Ditto, male, NSMT3289, Taiwan (NSMT); 6. Ditto, female, NSMT3290, Taiwan (NSMT); 7. Ditto, male, NSMT3295, Vietnam (NSMT); 8. Ditto, female, NSMT3296, Vietnam (NSMT); 9. Ditto, male, NSMT-MO169, Korea (NSMT); 10. Ditto, male, NSMT3294, Huangshan, Anhui (NSMT); 11. C. nanling sp. nov., male, holotype, NSMT3291, Nanling, Guangdong (SCAU); 12. Ditto, female, paratype, NSMT3292, Nanling, Guangdong (SCAU); 13. C. luding sp. nov., male, holotype, NSMT3279, Luding, Sichuan (SCAU); 14. Ditto, female, paratype, NSMT3280, Luding, Sichuan (SCAU); 15. C. maraho, male, holotype (TFRI); 16. Ditto, female, paratype, Taiwan (TFRI); 17. C. shanmeii, male, holotype, Taiwan (TFRI); 18. Ditto, female, paratype, Taiwan (TFRI); 19. C. hbun sp. nov., male, holotype, Taiwan (TFRI); 20. Ditto, female, paratype, Taiwan (TFRI); 21. Ditto, male, paratype, NSMT3303, Qiyunshan, Hunan (SCAU); 22. Ditto, female, paratype, NSMT3288, Jiulianshan, Guangdong (SCAU). Scale bar= 10 mm. Photos by Mamoru Owada (1–14, 19, 21, 22), Shipher Wu (15–18, 20).

FIGURE 23–33. Male labial palpus of Cidariplura gladiata complex (equal in scale). 23. C. gladiata, NSMT3285, Honshu, Japan (NSMT); 24. Ditto, Taiwan (TFRI); 25. Ditto, NSMT3295, Vietnam (NSMT); 26. Ditto, NSMT-MO169, Korea (NSMT); 27. Ditto, NSMT3301, Iriomote Is., Japan (NSMT); 28. C. shanmeii, paratype, Taiwan (TFRI); 29. C. luding sp. nov., holotype, NSMT3279, Luding, Sichuan (SCAU); 30. C. hbun sp. nov., paratype, Taiwan (TFRI); 31. C. nanling sp. nov., holotype, NSMT3291, Nanling, Guangdong (SCAU); 32. C. hbun, paratype, NSMT3303, Qiyunshan, Hunan (SCAU); 33. C. maraho, paratype, Taiwan (TFRI). Photos by Mamoru Owada (23, 25–27, 29, 31, 32), Shipher Wu (24, 28, 30, 33).

FIGURE 34–45. Male genitalia of Cidariplura gladiata complex (equal in scale). 34. C. gladiata, NSMT3285, Honshu, Japan (NSMT); 35. Ditto, NSMT3301, Iriomote Is., Japan (NSMT); 36. Ditto, NSMT3289, Taiwan (NSMT); 37. Ditto, NSMT3295, Vietnam (TFRI); 38. Ditto, NSMT-MO169, Korea (NSMT); 39. Ditto, NSMT3294, Huangshan, Anhui (NSMT); 40. C. nanling sp. nov., holotype, NSMT3291, Nanling, Guangdong (SCAU); 41. C. luding sp. nov., holotype, NSMT3279, Luding, Sichuan (SCAU); 42. C. maraho, holotype, TFRI147244, Taiwan (TFRI); 43. C. shanmeii, holotype, TFRI128721, Taiwan (TFRI); 44. C. hbun sp. nov., holotype, TFRI195178, Taiwan (TFRI); 45. Ditto, paratype, NSMT3303, Qiyunshan, Hunan (SCAU); Photos by Mamoru Owada (34–41, 44–45), Shipher Wu (42–43).

FIGURE 46–55. Female genitalia of Cidariplura gladiata complex (equal in scale). 46. C. gladiata, NSMT3286, Honshu, Japan (NSMT); 47. Ditto, NSMT3302, Iriomote Is., Japan (NSMT); 48. Ditto, NSMT3290, Taiwan (NSMT); 49. Ditto, NSMT3296, Vietnam (NSMT); 50. C. nanling sp. nov., paratype, NSMT3292, Nanling, Guangdong (SCAU); 51. C. luding sp. nov., paratype, NSMT3280, Luding, Sichuan (SCAU); 52. C. maraho, paratype, TFRI10165, Taiwan (TFRI); 53. C. shanmeii, paratype, TFRI153189, Taiwan (TFRI); 54. C. hbun sp. nov., paratype, TFRI160565, Taiwan (TFRI); 55. Ditto, paratype, NSMT3288, Jiulianshan, Guangdong (SCAU). Photos by Mamoru Owada (46–51, 55), Shipher Wu (52–54).