Enicospilus pseudopuncticulatus Shimizu, 2020
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publication ID |
https://dx.doi.org/10.3897/zookeys.990.55542 |
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publication LSID |
lsid:zoobank.org:pub:7B73642C-278D-40F8-9091-B26213C9A704 |
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persistent identifier |
https://treatment.plazi.org/id/24488D4C-E242-4199-B87E-98B67E27BC4A |
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taxon LSID |
lsid:zoobank.org:act:24488D4C-E242-4199-B87E-98B67E27BC4A |
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treatment provided by |
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scientific name |
Enicospilus pseudopuncticulatus Shimizu |
| status |
sp. nov. |
Enicospilus pseudopuncticulatus Shimizu sp. nov. Figure 35 View Figure 35
Etymology.
This species is very close to E. puncticulatus , hence the specific name based on their similarity.
Type series.
Total of 8 specimens (5♀♀3♂♂): Japan (5♀♀3♂♂).
HT: ♀, NIAES, Tsukuba City, Ibaraki Pref., Kantô-Kôshin, JAPAN, 17-26.VII.1989, M. Sharkey leg. (NIAES).
PT: 1♂, Awashima, Niigata Pref., Hokuriku, JAPAN, 7.VII.1937, K. Baba & J. Sawano leg. (MNHA); 1♂, same data as HT (NIAES); 1♀, NIAES, Tsukuba City, Ibaraki Pref., Kantô-Kôshin, JAPAN, 14-19.IV.1989, M. Sharkey leg. (EMUS); 2♀♀, Nokata, Nisshin, Aichi Pref., Tôkai, JAPAN, 10-16.VII.2011 (1♀), 23-30.VII.2011 (1♀), H. Seo & R. Mizutani leg. (MsT) (ELMU); 1♀1♂, Niryou, Takatsuki City, Ōsaka Pref., Kinki, JAPAN, 30.VI-4.VIII.2013, S. Fujie leg. (MsT) (♀, MNHA, SEN176-DDBJ-LC492933; ♂, OMNH, SEN147-DDBJ-LC492932).
Distribution.
Eastern Palaearctic region.
JAPAN: [Hokuriku] Niigata; [ Kantô-Kôshin] Ibaraki; [ Tôkai] Aichi; [Kinki] Ōsaka.
Bionomics.
Unknown.
Differential diagnosis.
Although this species is very close to E. puncticulatus based on both morphology and DNA barcoding (Fig. 6 View Figure 6 ), it is rather readily distinguishable by the linear central sclerite and straight fore wing vein 1cu-a, as in Fig. 35F View Figure 35 .
Description.
Female (n = 5). Body length 20.0-22.0 (HT: ca. 20.5) mm.
Head with GOI = 3.0-3.2 (HT: 3.1) (Fig. 35D View Figure 35 ). Lower face 0.8 × as wide as high, strongly shiny, finely punctate with setae (Fig. 35B View Figure 35 ). Clypeus 1.7-2.0 × (HT: 1.9) as wide as high, moderately punctate with setae as in upper face, moderately convex in profile, ventral margin impressed (Fig. 35B, D View Figure 35 ). Malar space 0.3 × as long as basal mandibular width (Fig. 35B, D View Figure 35 ). Mandible moderately twisted by 25-40° (HT: ca. 30°), rather long, proximally narrowed, distally parallel-sided, outer surface with diagonal setose groove (Fig. 35B, D View Figure 35 ). Upper tooth of mandible 1.8-2.1 × (HT: 2.1) as long as lower (Fig. 35B View Figure 35 ). Frons, vertex and gena strongly shiny with fine setae (Fig. 35B-D View Figure 35 ). Posterior ocellus very close to eye (Fig. 35B-D View Figure 35 ). Ventral end of occipital carina joining oral carina. Antennae with 66-69 (HT: 67) flagellomeres; first flagellomere 1.6-1.7 × (HT: 1.6) as long as second; 20th flagellomere 2.0-2.1 × (HT: 2.0) as long as wide.
Mesosoma entirely very strongly shiny with setae (Fig. 35E View Figure 35 ). Pronotum diagonally striate to moderately punctate (Fig. 35E View Figure 35 ). Mesoscutum 1.5 × as long as maximum width, finely punctate with setae, and evenly rounded in profile (Fig. 35E View Figure 35 ). Notauli absent (Fig. 35E View Figure 35 ). Scutellum moderately convex, finely punctate, with lateral longitudinal carinae along entire length of scutellum. Epicnemium densely punctate with setae. Epicnemial carina strong, evenly curved to anterior, dorsal end not reaching anterior margin of mesopleuron (Fig. 35E View Figure 35 ). Mesopleuron entirely longitudinally striate to punctostriate (Fig. 35E View Figure 35 ). Submetapleural carina broadened anteriorly (Fig. 35E View Figure 35 ). Metapleuron entirely moderately punctate with diagonal striae posteriorly (Fig. 35E View Figure 35 ). Propodeum declivous in profile; anterior transverse carina complete; anterior area longitudinally striate; spiracular area finely punctate with setae; posterior area finely to moderately rugose; propodeal spiracle elliptical and not joining pleural carina by ridge (Fig. 35E View Figure 35 ).
Wings (Fig. 35F View Figure 35 ). Fore wing length 13.0-14.0 (HT: ca. 13.0) mm with AI = 0.3-0.6 (HT: 0.5), CI = 0.2-0.3 (HT: 0.3), DI = 0.4, ICI = 0.5, SDI = 1.1-1.2 (HT: 1.1), SI = 0.2, SRI = 0.3; vein 1m-cu&M moderately sinuous; vein 2r&RS slightly sinuous; vein RS evenly curved; fenestra and sclerites of discosubmarginal cell as in Fig. 35F View Figure 35 ; proximal sclerite rounded triangular, not confluent with distal sclerite, strongly pigmented; central sclerite linear and pigmented, subparallel to 2r&RS, positioned in medio-distal part of fenestra; distal sclerite moderately pigmented; proximal corner of marginal cell evenly setose; posterodistal corner of second discal cell 95-100° (HT: ca. 95°) and of subbasal cell 75-85° (HT: ca. 75°); vein 1cu-a antefurcal to M&RS by 0.1 × length of 1cu-a. Hind wing with NI = 1.3-1.4 (HT: 1.4); vein RS slightly evenly curved; vein RA with 7-8 (HT: 7) uniform hamuli.
Legs. Hind leg with coxa in profile 1.9-2.0 × (HT: 1.9) as long as deep; basitarsus 2.0-2.1 × (HT: 2.0) as long as second tarsomere; fourth tarsomere 4.5-4.6 × (HT: 4.6) as long as wide; tarsal claw simply pectinate.
Metasoma with DMI = 1.3-1.4 (HT: 1.3), PI = 2.9-3.3 (HT: 3.2), THI = 2.2-2.6 (HT: 2.3); thyridium suboval, moderately sized; ovipositor sheath not longer than posterior depth of metasoma (Fig. 35A View Figure 35 ).
Colour (Fig. 35 View Figure 35 ). Entirely red-brown except for apex of mandible black. Wings hyaline; sclerites pigmented and amber; veins red-brown.
Male (n = 3). Very similar to female.
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