Chileanthicus, Werner, 1966

Kejval, Zbyněk, 2009, Taxonomic revision of the genus Chileanthicus Werner (Coleoptera: Anthicidae) 2180, Zootaxa 2180 (1), pp. 1-82 : 6-9

Key to the species of Chileanthicus (except C. lafertei )

1 Elytra with pale transverse band in posterior half; subapical process of metafemora minute; male sternum VII more or less distinctly emarginate postero-medially, with peg-like setae; species occuring in Chile................................. 2

- Elytra with different colour marking, never with single transverse band in posterior half; subapical process of metafemora usually more prominent; male sternum VII may be slightly modified, but never emarginate posteromedially; species occuring in Madagascar or Australia.............................................................................................. 9

2 Apical portion of tegmen of aedeagus with paired latero-basal incision ( Fig. 52 ); female tergum VII strongly bulging ( Fig. 53 ) ............................................................................................................................................... C. penai Werner

- Apical portion of tegmen lacking latero-basal incisions; female tergum VII at most moderately convex ................ 3

3 Elytra in female tapering apically ( Fig. 240 ); apex of tegmen of aedeagus upturned in lateral view ( Fig. 12 ) ............ .................................................................................................................................................. C. acutipennis sp. nov.

- Elytra in female evenly rounded to truncate apically; apex of tegmen straight to moderately curved ventrad ......... 4

4 Antennae conspicuously slender, slightly enlarged in terminal third; antennomere III more than 3.0 times as long as wide ( Fig. 32 ) .............................................................................................................................................................. 5

- Antennae somewhat stronger, moderately enlarged in distal third, antennomere III less than 2.5 times as long as wide ( Fig. 33 ) .............................................................................................................................................................. 6

5 Paired prongs of male sternite VIII slender, each with more than 10 peg-like setae, rather unevenly scattered ventrally ( Figs 18, 19 ); apical portion of tegmen of aedeagus rather straight in lateral view, and apex wider, rounded in ventral view ( Fig. 20 ) .................................................................................................................. C. elmorado sp. nov.

- Paired prongs of male sternite VIII more robust, each with less than 10 peg-like setae arranged in row ventrally ( Figs 28, 29 ); apical portion of tegmen of aedeagus arcuate in lateral view, and apex narrowed, nearly bluntly pointed in ventral view ( Fig. 31 ) ..................................................................................................................... C. hirsutus sp. nov.

6 Male sternum VII with about 40 peg-like setae ( Fig. 13 ); apical portion of tegmen of aedeagus with conspicuous, long, transverse edge dorsally in basal half, distinctly protruding in lateral view ( Fig. 16 ) ..... C. baculentus sp. nov.

- Male sternum VII with less than 20 peg-like setae; apical portion of tegmen rather smoothly shaped dorsally, dorsal edge indistinct or at least less conspicuous, short, barely protruding from outline of tegmen in lateral view ........... 7

7 Paired prongs of male sternite VIII with peg-like setae arranged somewhat laterally in row ( Figs 46, 47 ); dorsal outline of apical portion of tegmen distinctly sinuous in lateral view, apex of tegmen rather wide, subtriangular in ventral view ( Fig. 48 ) ............................................................................................................................... C. mitis sp. nov.

- Paired prongs of male sternite VIII with peg-like setae rather scattered on ventral side; dorsal outline of apical portion of tegmen at most slightly sinuous/impressed ( Fig. 26 ) or evenly arcuate ( Fig. 41 ) in lateral view, apex of tegmen rather narrowed, bluntly pointed in ventral view (only C. maritimus sp. nov., Fig. 42 ) .................................... 8

8 Male sternum VII moderately and narrowly emarginate posteriorly ( Fig. 22 ); apical portion of tegmen of aedeagus with dorsal transverse edge distant from its base ( Fig. 26 , marked by arrow); female sternum VII modified, with pair of short, bluntly pointed median protrusions ( Fig. 21 ) ..................................................... C. femineus sp. nov. (male)

- Male sternum VIII rather strongly and widely emarginate posteriorly ( Fig. 38 ); apical portion of tegmen of aedeagus with dorsal transverse edge situated near base ( Fig. 41 ); female sternum VII simple, with posterior margin evenly rounded ...................................................................................................................................... C. maritimus sp. nov.

9 Species occuring in Madagascar............................................................................................................................... 10

- Species occuring in Australia................................................................................................................................... 11

10 Body surface matte, very densely and finely punctured ( Fig. 58 ), including lateral sides of pronotum; body setation very short, appressed, erect setae less conspicuous; paired prongs of male sternite VIII bearing peg-like setae ( Fig. 61 ) .................................................................................................................................................. C. dentatus (Dajoz)

- Body surface moderately glossy, its dorsal punctation coarser and sparser ( Fig. 54 ), lateral sides of pronotum partly impunctate and glossy; body setation generally longer and more raised, erect setae more conspicuous; paired prongs of male sternite VIII at most finely setose, lacking peg-like setae ( Fig. 56 ) ................................... C. cursor sp. nov.

11 Metafemora unidentate; with single tooth-like process on inner side subapically ................................................... 12

- Metafemora bidentate or tridentate, with 1–2 processes on inner side and the third one on outer side subapically 41

12 Pronotum with dorso-lateral margins forming sharp edge ( Fig. 241 ; see also C. comes sp. nov. with nearly carinate margins) .................................................................................................................................................................... 13

- Pronotum more or less rounded dorso-laterally (at most with short edge, close behind the widest point of pronotum, see C. comes sp. nov.) ............................................................................................................................................... 16

13 Head with tempora forming blunt edge at level of ventral eye margin (eyes thus seem to be situated rather dorso-laterally; Fig. 241 ); punctation of elytra less conspicuous, simple, punctures distinctly smaller/finer than those on pronotum ........................................................................................................................................................................ 14

- Head with tempora rounded (eyes situated laterally); punctation of elytra conspicuous, its coarse punctures about as large as those on pronotum ....................................................................................................................................... 15

14 Head moderately glossy, distinctly but somewhat sparsely punctured, punctures finer/smaller than those on pronotum ( Fig. 102 ); male characters as in Figs 104–107 ......................................................................... C. discus sp. nov.

- Head matte, distinctly and very densely punctured, similar to pronotum ( Fig. 108 ); male characters as in Figs 110, 111 ........................................................................................................................................................ C. exul sp. nov.

15 Head widely rounded posteriorly in dorsal view ( Fig. 63 ); antennae rather short, antennomeres less elongate ( Fig. 64 ): III 1.8 times, X 1.2 times, XI 1.9 times as long as wide; male characters as in Figs 66, 67 .................................. .................................................................................................................................................. C. angulicollis sp. nov.

- Head angulate posteriorly in dorsal view ( Fig. 129 ); antennae longer, antennomeres more elongate ( Fig. 130 ): III 2.3 times, X 1.7 times, XI 2.1 times as long as wide; male characters as in Figs 132–134 ................. C. inflatus sp. nov.

16 Basal punctation of elytra distinctly double, with coarse and fine punctures ........................................................... 17

- Basal punctation of elytra uniformly fine to somewhat heterogeneous (indistinctly double, its coarse punctures barely discernible) ..................................................................................................................................................... 24

17 Erect setae of elytra conspicuous, much longer than ordinary, subdecumbent setae ............................................... 18

- Erect setae of elytra inconspicuous, about as long as other setae ............................................................................. 20

18 Paler marking and silvery setae of elytra forming narrow band/spot in apical and irregular spot in basal half, extending towards base ( Figs 146 , 235 ), silvery setae less contrasting, rather evenly directed posteriad (not swirled); subapical process of metafemora smaller ( Fig. 147 ); male characters as in Figs 148, 149 .............. C. kinchega sp. nov.

- Paler marking and silvery setae of elytra forming two, narrow transverse bands, silvery setae rather conspicuous, distinctly swirled ( Figs 86 , 141 ); subapical process of metafemora somewhat more conspicuous, with wider base ( Figs 87 , 142 ) ............................................................................................................................................................ 19

19 Male characters as in Figs 143–145 .............................................................................................. C. interruptus (Lea)

- Male characters as in Figs 88, 89 ................................................................................................ C. decipiens sp. nov.

20 Head and pronotum wide, head angulate posteriorly in dorsal view; basal coarse punctures of elytra distinctly larger than fine punctures ( Fig. 155 ); male metatibiae with two longitudinal grooves on inner side, another male characters as in Figs 157–159 ........................................................................................................................... C. latibasis (Lea)

- Head and pronotum narrower, head at most widely rounded posteriorly; basal coarse punctures of elytra at most slightly to moderately larger than fine punctures ( Figs 77 , 123 , 135 , 171 ); male matatibiae lacking grooves on inner side ............................................................................................................................................................................ 21

21 Elytra slightly glossy to matte, very densely punctured ........................................................................................... 22

- Elytra moderately glossy, somewhat less densely punctured ................................................................................... 23

22 Dorsal punctation of head distinctly double, coarse punctures much larger than fine punctures ( Fig. 123 ); the anterior setose band of elytra extending dorso-laterally towards base ( Figs 123 , 234 ); male characters as in Figs 125–128 ................................................................................................................................................... C. improvidus sp. nov.

- Dorsal punctation of head indistinctly double, coarse punctures only moderately larger than fine punctures ( Fig. 77 ); the anterior setose band narrow, at most moderately widened near lateral margin; male characters as in Figs 79, 80 . ........................................................................................................................................................... C. comes sp. nov.

23 Male sternum III with small median protuberance, male metatibiae with small process/denticle on inner side apically ( Fig. 173 ), other male characters as in Figs 174–176 ................................................................... C. melasomus (Lea)

- Male sternum III simple, lacking median protuberance, male metatibiae simple, lacking apical process/denticle, other male characters as in Figs 137–140 ...................................................................................... C. insignis sp. nov.

24 Body largely rufous, elytra with black marking ( Fig. 166 ), evenly and sparsely punctured, rather glossy, uniformly setose; male characters as in Figs 168–170 ............................................................................... C. matthewsi sp. nov.

- Body largely dark coloured, elytra dark, unicoloured or with paler marking, densely punctured, at most moderately glossy, with silvery setose bands/spots ..................................................................................................................... 25

25 Elytra uniformly dark, basal punctation of elytra always uniformly fine ................................................................. 26

- Elytra mostly dark with distinct paler marking (bands/spots, suture, lateral margins), rarely nearly unicoloured (variability, some specimens of C. melasomus and C. quadrimaculatus ); basal punctation of elytra uniformly fine to somewhat heterogeneous (indistinctly double, its coarse punctures barely discernible, see C. melasomus ) ........... 32

26 Silvery/whitish setose bands of elytra with setae at least partly distinctly obliquely to transversely pointing (swirled) ................................................................................................................................................................................... 27

- Silvery/whitish setose bands of elytra with setae rather evenly pointing posteriad as ordinary appressed/subdecumbent setae ( Fig. 238 ) .................................................................................................................................................. 31

27 Head large, distinctly wider (behind eyes) than narrow pronotum ( Fig. 90 ); elytra rather short and ovoid, as long as head and pronotum ( Figs 90 , 232 ); pronotal disc with moderate median longitudinal impression; femora (especially metafemora) long clavate, rather abruptly swollen distally ( Fig. 93 ) ............................................... C. audax sp. nov.

- Head smaller in relation to rather wide pronotum; elytra more elongate, longer than head and pronotum; pronotal disc evenly convex, median longitudinal impression at most indicated anteriorly; femora clavate, but their basal narrow portion rather evenly widening ( Figs 186, 189 ) ................................................................................................ 28

28 Pronotum nearly trapezoidal, widely rounded to angulate anteriorly in dorsal view, strongly narrowing posteriad ( Fig. 183 ); punctation of head and pronotum simple, punctures much larger than those on elytra; basal punctures of elytra arranged in short, obligue rows, separated by fine wrinkles ( Fig. 183 )........................... C. pubifasciatus (Lea)

- Pronotum more or less widely rounded anteriorly and moderately narrowing posteriad in dorsal view ( Figs 72 , 118 , 185 ); punctures of head and pronotum at most moderately larger than those on elytra ( Fig. 185 ), if some of them distinctly larger, than punctation of head and pronotum distinctly double ( Figs 72 , 118 ); basal punctures of elytra irregularly scattered, interspaces rather smooth, unwrinkled ........................................................................................... 29

29 Elytra subtruncate apically; surface of head, pronotum and elytra nearly matte, very densely punctured ( Fig. 185 ); silvery/whitish setae of transverse bands of elytra mostly transversely pointing laterad ( Figs 185 , 237 ) ..................... ........................................................................................................................................................ C. pulcher sp. nov.

- Elytra conjountly rounded apically; surface of head, pronotum and elytra rather moderately glossy, its punctation somewhat less dense ( Figs 72 , 118 ); silvery/whitish setae of transverse bands on elytra differently arranged, swirled medially ( Figs 72 , 118 ) ............................................................................................................................................. 30

30 Male characters as in Figs 120–122 .............................................................................................. C. gammon sp. nov.

- Male characters as in Figs 74–76 ............................................................................................... C. campicola sp. nov.

31 Antennae moderately enlarged in terminal third: antennomere X 1.7 times, XI 2.2 times as long as wide; pronotum trapezoidal, widely rounded to angulate anteriorly in dorsal view, strongly narrowing posteriad ( Fig. 183 ); punctures of head and pronotum much larger than those on elytra, basal punctures of elytra arranged in short, obligue rows, which may appear to be separated by fine wrinkles ( Fig. 183 ) ................................................ C. pubifasciatus (Lea)

- Antennae distinctly enlarged in terminal third ( Fig. 188 ): antennomere X as long as wide, XI 1.8 times as long as wide; pronotum widely rounded anteriorly in dorsal view, moderately narrowing posteriad ( Fig. 187 ); punctures of head and pronotum only moderately larger than those on elytra, basal punctures of elytra irregularly scattered ( Fig. 187 ) ................................................................................................................................................ C. pumilio sp. nov.

32 Silvery setae of elytra both forming setose bands/spots (on paler marking) and scattered on dark coloured surface ( Fig. 233 ) ................................................................................................................................................................... 33

- Silvery setae of elytra restricted to paler coloured marking and forming only setose bands/spots (they may be scattered at most along lateral margins, but never on dorsal side) .................................................................................. 34

33 Antero-lateral outline of pronotum evenly rounded in dorsal view ( Fig. 81 ); male meso- and metatibiae slightly angulately produced on inner side apically, metatibiae uniformly short setose; male sternum III with small median protuberance, another male characters as in Figs 83–85 ........................................................... C. communis sp. nov.

- Antero-lateral outline of pronotum with slight angle at widest place in dorsal view ( Fig. 112 ); male meso- and metatibiae simple apically, metatibiae with a tuft of long setae on inner side apically ( Fig. 115 ); male sternum III simple, male characters as in Figs 116–117 ............................................................................................... C. eyrensis sp. nov.

34 Dorsal surface of head matte, very densely, evenly punctured, punctures nearly contiguous and uniform ( Fig. 204 ); sternum VII with a pair of distinct protuberances ( Fig. 206 ), another male characters as in Figs 207, 208 ................. .................................................................................................................................................. C. tuberculifer sp. nov.

- Dorsal surface of head slightly glossy, rarely nearly matte ( C. comes sp. n.), punctures distinctly separated, often unevenly spaced (sparser in median row) and heterogeneous (more or less distinctly double); male sternum VII simple, lacking paired protuberance ............................................................................................................................... 35

35 Head widely rounded to angulate posteriorly; elytra somewhat widely rounded apically ( Fig. 68 ); elytra nearly matte, very densely punctured, basal punctation simple, punctures somewhat serial (surface appears to be transversely wrinkled); anterior silvery setose band of elytra wide, its posterior margin uneven, and setae along anterior margin pointing mediad ( Fig. 68 ); male characters as in Figs 70, 71 ............................................ C. aprilis sp. nov.

- Head evenly to moderately widely rounded posteriorly; elytra evenly rounded apically; elytra less densely punctured, rarely nearly matte ( C. comes sp. n.), basal punctation simple or somewhat heterogeneous (indistinctly double), punctures rather evenly spaced (interspaces smooth, unwrikled); anterior silvery setose band of elytra rather narrow and even, setae along its anterior margin pointing posteriad ........................................................................ 36

36 Basal punctation of elytra simple, uniformly fine ( Figs 177 , 192 , 212 ); male sternum VII simple, lacking median protuberance .............................................................................................................................................................. 37

- Basal punctation of elytra indistinctly double (its coarse punctures sometimes barely discernible, Figs 135 , 171 ); male sternum VII simple or with median protuberance ............................................................................................ 39

37 Head and pronotum wider, head rather widely rounded posteriorly ( Fig. 177 ); dorsal punctation of head very dense, indistinctly double ( Fig. 177 ); male metatibiae simple and with a tuft of long, stiff setae apically ( Fig. 179 ), another male characters as in Figs 180–182 .............................................................................................. C. paganus sp. nov.

- Head and pronotum narrower, head narrower, at most unevenly rounded posteriorly ( Figs 192 , 212 ); dorsal punctation of head less dense, clearly double, its coarse punctures distinctly larger then fine punctures .......................... 38

38 Head evenly densely punctured, including frons medially; male characters as in Figs 194–196 ................................. ............................................................................................................................................ C. quadrimaculatus (King)

- Head unevenly punctured, punctures distinctly sparser medially, with indication of glossy medial line, especially anteriorly on frons and clypeus; male characters as in Figs 214–216 ......................................... C. volselifer sp. nov.

39 Elytra slightly glossy to matte, punctures densely spaced; pronotum distinctly wider than head ( Fig. 77 ), pronotal disc with nearly carinate lateral margins; male sternum III with median protuberance, male metatibiae uniformly short setose; male characters as in Figs 79–80 ................................................................................. C. comes sp. nov.

- Elytra moderately glossy, punctures less densely spaced; pronotum narrower, pronotal disc more convex, with rounded lateral margins; male sternum VII simple or with median protuberance .................................................... 40

40 Male sternum III with small median protuberance, male metatibiae with small apical process/denticle ( Fig. 173 ), another male characters as in Figs 174–176 ................................................................................. C. melasomus (Lea)

- Male sternum III simple, lacking median protuberance, male metatibiae simple apically, lacking apical process, another male characters as in Figs 137, 140 .................................................................................. C. insignis sp. nov.

41 Metafemora bidentate, with single process on inner side, which is strong, tooth-like, wide basally and strongly narrowing towards pointed apex ( Figs 151 , 201 ) ........................................................................................................... 42

- Metafemora bi- to tridentate, 1–2 processes on inner side narrow, thorn-like shaped ( Figs 95 , 161 ), the smaller one may be reduced ( Fig. 190 ) or resembling angulately produced edge ( Fig. 211 ) ...................................................... 43

42 Body darker coloured, blackish; pronotal disc uneven, with distinct median longitudinal impression; surface of elytra uneven, distinctly impressed dorsally at place of basal setose band; silvery setose bands of elytra very conspicuous, contrasting, setae largely and quite distinctly swirled ( Fig. 239 ); body punctation conspicuously double; male characters as in Figs 198–200 ...................................................................................................... C. speciosus (King)

- Body paler coloured, mostly brownish; pronotal disc and elytra evenly shaped, lacking distinct impressions; silvery setose bands of elytra less conspicuous, narrower, setae moderately swirled medially ( Fig. 150 ); body punctation distinctly double but coarse punctures comparatively less conspicuous ( Fig. 150 ); male characters as in Figs 153, 154 ................................................................................................................................................. C. kingii (MacLeay)

43 Head angulate posteriorly; elytra with single silvery/whitish setose band ( Figs 209 , 225 ), erect setae rather short and inconspicuous; terminal spurs of metatibiae well developed ........................................................ C. uhmanni sp. nov.

- Head at most widely rounded posteriorly; elytra with two silvery/whitish setose bands ( Figs 94 , 160 , 236 ), erect setae rather long and conspicuous; terminal spurs of metatibiae absent or at least strongly reduced and indistinct ..... ................................................................................................................................................................................... 44

44 Setation of metatibiae rather evenly short and fine (except apical fringe of short, stiff setae); male characters as in Figs 162–165 .................................................................................................................................. C. mastersii (King)

- Setation of metatibiae with short, stiff, bluntly pointed setae scattered on outer side (except apical fringe) and numerous conspicuously long, raised setae on inner side distally ( Figs 96, 97 ); male characters as in Figs 98–101 ... ....................................................................................................................................................... C. dentivarius (Lea)

Figures

FIGURES 45–53: Chileanthicus mitis sp. nov., male, holotype: 45, sternum VII; 46, sternite VIII (half), dorsal view; 47, sternite VIII, lateral view; 48, aedeagus, ventral (left) and lateral (right) view; C. penai (Werner), male, paratype (UAIC): 49, sternum VII; 50, sternite VIII (half), dorsal view; 51, abdominal segment IX; 52, aedeagus in ventral (right) and its apical portion in lateral (left) view; 53, apex of abdomen in female, lateral view. Scale (0.5 mm): A, 45, 49, 51; B, 48; C, 53; (0.2 mm): D, 50; E, 46, 47; F, 52.

FIGURES 232–241: Elytra of Chileanthicus species: 232, C. audax sp. nov.; 233, C. communis sp. nov.; 234, C. improvidus sp. nov.; 235, C. kinchega sp. nov.; 236, C. mastersii (King); 237, C. pulcher sp. nov.; 238, C. pumilio sp. nov.; 239, C. speciosus (King); 240, C. acutipennis sp. nov., apices of elytra in female; 241, C. discus sp. nov., head and pronotum in lateral view.

FIGURES 9–16: Chileanthicus acutipennis sp. nov., male, holotype: 9, sternum VII; 10, sternite VIII (half), dorsal view; 11, sternite VIII, lateral view; 12, aedeagus, ventral (left) and lateral (right) view; C. baculentus sp. nov., male, holotype: 13, sternum VII; 14, sternite VIII (half), dorsal view; 15, sternite VIII, lateral view; 16, aedeagus, ventral (right) and lateral (left) view. Scale (0.5 mm): A, 9, 12; B, 10, 11, 13–15; C, 16.

FIGURES 27–33: Chileanthicus hirsutus sp. nov., male, holotype: 27, sternum VII; 28, sternite VIII (half), dorsal view; 29, sternite VIII, lateral view; 30, tergite VIII; 31, aedeagus, ventral (left) and lateral (right) view; antennomeres I–VI: 32, C. hirsutus sp. nov.; 33, C. maritimus sp. nov. Scale (0.5 mm): A, 28, 29; B, 27; C, 31; D, 30, 32; E, 33.

FIGURES 17–26: Chileanthicus elmorado sp. nov., male, holotype: 17, sternum VII; 18, sternite VIII (half), dorsal view; 19, sternite VIII, lateral view; 20, aedeagus in ventral (left) and its apical portion in lateral (right) view; 21, C. femineus sp. nov., female, holotype, sternum VII; C. femineus sp. nov., male, paratype: 22, sternum VII; 23, sternite VIII (half), dorsal view; 24, sternite VIII, lateral view; 25, tergite VIII; 26, aedeagus (lacking apical portion), lateral view. Scale (0.5 mm): A, 20, 26; B, 17, 22; C, 21, 25; D, 18, 19, 23, 24.

FIGURES 34–44: Chilenathicus lafertei (Solier), female, syntype (MNHN): 34, body form; 35, metafemur, lateral view; 36, sternum VII; 37, tergum VII; C. maritimus sp. nov., male, holotype: 38, sternum VII; 39, sternite VIII (half), dorsal view; 40, sternite VIII, lateral view; 41, aedeagus, ventral (right) and lateral (left) view; 42, apex of aedeagus, ventral view; Metafemur, lateral view: 43, C. elmorado sp. nov.; 44, C. femineus sp. nov. Scale (0.5 mm): A, 39, 40; B, 38; C, 41, 42; D, 35, 36, 38, 43, 44; (1 mm): E, 34.

FIGURES 54–62: Chileanthicus cursor sp. nov., male, holotype: 54, punctation of head (top) and elytra (bottom); 55, hind leg; 56, sternite VIII, dorsal view; 57, aedeagus in ventral (left) and its apical portion in lateral (right) view; C. dentatus (Dajoz), male, 48 km ENE Morondava (FMNH): 58, punctation of head (top) and elytra (bottom); 59, apical portion of metafemur with tibia and tarsus; 60, sternite VIII, dorsal view; 61, the same, lateral view; 62– aedeagus in ventral (left) and its apical portion in lateral (right) view. Scale (0.5 mm): A –57, 62; B, 54, 58; C, 55; D, 59; (0.2 mm): E, 56, 60, 61.

FIGURES 102–111: Chileanthicus discus sp. nov., male, holotype: 102, body form and punctation of head (top) and pronotum (bottom); 103, metafemur; 104, apex of metatibia; 105, sternite VIII; 106, segment IX (spiculum); 107, aedeagus in ventral (right) and lateral (left) view; C. exul sp. nov., male, holotype: 108, punctation of head; 109, metafemur; 110, sternite VIII; 111, aedeagus in ventral (left) and lateral (right) view. Scale (1.0 mm): A, 102; B, 103, 109; (0.5 mm): B, 104, 107, 111; C, 106; D, 105, 110.

FIGURES 63–71: Chileanthicus angulicollis sp. nov., male, holotype: 63, body form and punctation of head (top) and elytra (bottom); 64, antenna; 65, metafemur; 66, sternite VIII; 67, apical portion of aedeagus in ventral (right) and lateral (left) view; C. aprilis sp. nov., male, holotype: 68, body form and punctation of head (top) and elytra (bottom); 69, metafemur; 70, sternite VIII; 71, aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 63, 68; B, 65, 69; (0.5 mm): C, 66, 70, 71; D, 67.

FIGURES 123–134: Chileanthicus improvidus sp. nov., male, holotype: 123, body form and punctation of head (top) and pronotum (bottom); 124, metafemur; 125, sternite VIII; 126, segment IX (spiculum); 127, outline of aedeagus in lateral view; 128, apex of aedeagus in ventral (right) and lateral (left) view; C. inflatus sp. nov., male, holotype: 129, body form and punctation of head (top) and elytra (bottom); 130, antenna; 131, metafemur; 132, sternite VIII; 133, aedeagus in ventral view; 134, apex of aedeagus in lateral view. Scale (1.0 mm): A, 123, 129; (0.5 mm): B, 130; C, 124, 126, 127, 131, 133; D, 128, 134; E, 125.

FIGURES 146–154: Chileanthicus kinchega sp. nov., male, holotype: 146, body form and punctation of head (top) and elytra (bottom); 147, metafemur; 148, sternite VIII; 149, apex of aedeagus in ventral (right) and lateral (left) view; C. kingii (MacLeay), male, syntype (SAMA): 150, body form and punctation of head (top) and elytra (bottom); 151, metafemur; 152, apex of metatibia in lateral view; 153, sternite VIII; 154, apex of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 150; B, 146; (0.5 mm): C, 147, 151, 152; (0.2 mm): D, 148, 153; E, 149, 154.

FIGURES 81–89: Chileanthicus communis sp. nov., male, holotype: 81, body form and punctation of head (top) and elytra (bottom); 82, metafemur; 83, sternite VIII; 84, segment IX (spiculum); 85, apical portion of aedeagus in ventral (left) and lateral (right) view; C. decipiens sp. nov., male, holotype: 86, body form and punctation of head (top) and elytra (bottom); 87, metafemur; 88, sternite VIII; 89, aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 81, 86; B, 82, 84, 87; (0.2 mm) C, 83, 89; D, 88; E, 85.

FIGURES 135–145: Chileanthicus insignis sp. nov., male, holotype: 135, body form and punctation of head (top) and elytra (bottom); 136, metafemur; 137, sternite VIII; 138, tergite VIII; 139, segment IX (spiculum); 140, apex of aedeagus in ventral (left) and lateral (right) view; C. interruptus (Lea), male, Townsville (SAMA): 141, body form and punctation of head (top) and elytra (bottom); 142, metafemur; 143, sternite VIII; 144, segment IX (spiculum); 145, apex of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 135, 141; B, 136, 142; (0.5 mm): C, 136; D, 139, 143, 144; (0.2 mm): E, 137; F, 140, 145.

FIGURES 155–159: Chileanthicus latibasis (Lea), male, Brookfield Con. Park (ANIC): 155, body form and punctation of head (top) and elytra (bottom); 156, metafemur; 157, sternite VIII; 158, segment IX (spiculum); 159, apical portion of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 155; B, 156; (0.5 mm): B, 157; C, 158; D, 159.

FIGURES 72–80: Chileanthicus campicola sp. nov., male, holotype: 72, body form and punctation of head (top) and elytra (bottom); 73, metafemur; 74, sternite VIII; 75, apical portion of aedeagus in lateral view; 76, apical portion of tegmen (right) and median lobe (left) of aedeagus in ventral view; C. comes sp. nov., male, holotype: 77, body form and punctation of head (top) and elytra (bottom); 78, metafemur; 79, sternite VIII; 80, apical portion of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 72, 77; B, 72, 78; (0.2 mm): C, 74; D, 79, 80; E, 75, 76.

FIGURES 171–182: Chileanthicus melasomus (Lea), male, South Australia (ANIC): 171, body form and punctation of head (top) and elytra (bottom); 172, metafemur; 173, apex of metatibia; 174, sternite VIII; 175, aedeagus in ventral view; 176, apex of aedeagus in ventral (right) and lateral (left) view; C. paganus sp. nov., male, holotype: 177, body form and punctation of head (top) and elytra (bottom); 178, metafemur; 179, apex of metatibia; 180, sternite VIII; 181, segment IX (spiculum); 182, apex of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 171, 177; B, 172, 175, 178; (0.5 mm): B, 173, 174, 179; C, 181; D, 176, 182.

FIGURES 160–170: Chileanthicus mastersii (King), male, lectotype: 160, body form and punctation of head (top) and elytra (bottom); 161, metafemur; 162, sternite VIII; 163, segment IX (spiculum); 164, aedeagus in ventral (right) and lateral (left) view; 165, apical portion of aedeagus in ventral view; C. matthewsi sp. nov., male, holotype: 166, body form and punctation of head (top) and elytra (bottom); 167, metafemur; 168, apex of metatibia; 169, sternite VIII; 170, apex of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 160; B, 161, 166; C, 164, 167; (0.5 mm): C, 168; D, 163; (0.2 mm): D, 170; E, 162, 165; F, 169.

FIGURES 90–101: Chileanthicus audax sp. nov., female, holotype: 90, body form; 91, punctation of head (top) and elytra (bottom); 92, antenna; 93, metafemur; C. dentivarius (Lea), male, syntype: 94, body form and punctation of head (top) and elytra (bottom); 95, metafemur; 96, metatibia in caudal view; 97, apex of metatibia in lateral view; 98, sternite VIII; 99, segment IX (spiculum); 100, outline of aedeagus and median lobe in ventral view; 101, apical portion of aedeagus in ventral view. Scale (1.0 mm): A, 90; B, 94; (0.5 mm): C, 92, 93, 95, 100; D, 99; E, 96, 97; F, 98, 101.

FIGURES 183–191: Chileanthicus pubifasciatus (Lea), female, holotype: 183, body form and punctation of head (top) and elytra (bottom); 184, distal part of metafemur; C. pulcher sp. nov. (female, holotype): 185, body form and punctation of head (top) and elytra (bottom); 186, metafemur; C. pumilio sp. nov., male, holotype: 187, body form and punctation of head (top) and elytra (bottom); 188, antenna; 189, metafemur; C. mastersii (King), South Australia (SAMA, ANIC): 190, metafemur; 191, apex of tegmen and median lobe of aedeagus in ventral view. Scale (1.0 mm): A, 183, 185; B, 187; (0.5 mm): B, 186, 187; C, 190; D, 184; E, 189.

FIGURES 112–122: Chileanthicus eyrensis sp. nov., male, holotype: 112, body form and punctation of head (top) and pronotum (bottom); 113, antenna; 114, metafemur; 115, apex of metatibia; 116, sternite VIII; 117, apex of aedeagus in ventral (left) and lateral (right) view; C. gammon sp. nov., male, holotype: 118, body form and punctation of head (top) and elytra (bottom); 119, metafemur; 120, sternite VIII; 121, apex of tegmen in ventral (left) and lateral (right) view; 122, apex of median lobe of aedeagus in ventral (left) and lateral (right) view. Scale (1.0 mm): A, 118, 112; B, 113, 114, 119; (0.2 mm): C, 116, 120; D, 115, 121, 122; E, 117.

FIGURES 201–208: Chileanthicus speciosus (King), male, holotype: 201, metafemur and tibia; 202, sternum VII; 203, segment IX (spiculum); C. tuberculifer sp. nov., male, holotype: 204, body form and punctation of head (top) and elytra (bottom); 205, metafemur; 206, sternum VII; 207, sternite VIII; 208, aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 204; B, 205, 206; C, 201; (0.5 mm): B, 208; D, 202, 203; E, 207.

FIGURES 192–200: Chileanthicus quadrimaculatus (King), male, South Australia (SAMA): 192, body form and punctation of head (top) and elytra (bottom); 193, metafemur; 194, sternite VIII; 195, apical portion of aedeagus (left) in lateral and tegmen (right) in ventral view; 196, apex of tegmen in ventral view; C. speciosus (King), male, holotype: 197, body form and punctation of head (top) and elytra (bottom); 198, sternite VIII; 199, tergite VIII; 200, apex of aedeagus in ventral (right) and lateral (left) view. Scale (1.0 mm): A, 192; B, 197; C, 199; D, 193; (0.5 mm): C, 198; D, 194, 195, 196; (0.2 mm): E, 200.

FIGURES 209–219: Chileanthicus uhmanni sp. nov., male, holotype: 209, body form and punctation of head (top) and elytra (bottom); 210, antenna; 211, metafemur; C. volselifer sp. nov., male, holotype: 212, body form and punctation of head (top) and elytra (bottom); 213, metafemur; 214, apex of metatibia; 215, sternite VIII; 216, apex of aedeagus in ventral (right) and lateral (left) view; C. cursor sp. nov. (male, holotype): 217, tergite VIII; 218, segment IX (spiculum); 219, C. quadrimaculatus (King), South Australia (SAMA), male tergite VIII. Scale (1.0 mm): A, 209, 212; (0.5 mm): B, 211, 217, 218, 219; C, 213; D, 210; (0.2 mm): B, 214; E, 215, 216.

FIGURES 220–231: Habitus of Chileanthicus species: 220, C. acutipennis sp. nov. (female); 221, C. maritimus sp. nov.; 222, C. dentatus (Dajoz); 223, C. angulicollis sp. nov.; 224, C. inflatus sp. nov.; 225, C. uhmanni sp. nov.; 226, C. discus sp. nov.; 227, C. mastersii (King); 228, C. pulcher sp. nov.; 229, C. pumilio sp. nov.; 230, C. quadrimaculatus (King); 231, C. speciosus (King).