Protohermes impunctatus, Liu, Xingyue, Hayashi, Fumio & Yang, Ding, 2008

Liu, Xingyue, Hayashi, Fumio & Yang, Ding, 2008, The Protohermes guangxiensis species-group (Megaloptera: Corydalidae), with descriptions of four new species, Zootaxa 1851, pp. 29-42: 36

publication ID 10.5281/zenodo.183412

persistent identifier

treatment provided by


scientific name

Protohermes impunctatus

sp. nov.

Protohermes impunctatus   sp. nov.

( Figs. 3 View FIGURES 1 – 5 , 20– 24 View FIGURES 20 – 24 )

Diagnosis. This species is distinguished by its immaculate, yellowish head. Additionally, it has two pair of prothoracic marks, the anterior pair being slightly longer than the posterior one. The male 10 th tergite is nearly rounded in caudal view, with a short, blunt ventral process.

Male. Body length 30 mm (n= 1); forewing length 43 mm, hindwing length 39 mm.

Head yellowish-brown, immaculate; postocular spine absent. Compound eyes greyish-brown; ocelli yellow, medially margined brown. Antenna blackish-brown with scape and pedicel yellowish-brown. Mouthparts yellowish-brown; mandibles with distal half blackish-brown; maxillary and labial palpi with distal one or two segments dark.

Thorax yellowish-brown; pronotum with two pairs of black vittae near lateral margins, anterior pair slightly longer than posterior pair; mesonotum laterally with pair of brown marks. Thorax with pilous yellow setae that are much longer on meso – and metathorax than on prothorax. Legs yellow, with yellowish short dense setae; fore and mid legs with tips of tibiae and entire tarsi pale brown; hind legs with tips of 1 st tarsomere and entire 2 nd to 4 th tarsomeres brown; tarsal claws reddish-brown. Wings pale greyish-brown, with several yellowish marks. Forewing with numerous brownish stripes on costal cellules; basally with an irregular mark; medially with many irregular marks on almost all crossveins, transversely arranged into three lines; and with a large round mark at distal 1 / 3. Hindwing much paler than forewing, with an indistinct round mark at distal 1 / 3. Veins pale yellow, slightly darker at wing tips. Rs 9 –branched, last branch bifurcate; 6–9 crossveins between R 1 and Rs; M 1 + 2 4 to 6 -branched, M 3 + 4 2 -branched; 1 A 3 -branched.

Abdomen yellowish-brown, laterally much darker. Ninth tergum ( Fig. 21 View FIGURES 20 – 24 ) much wider than long with oblique lateral margins, arcuately incised anteriorly, slightly produced posteriorly; internal inflection V – shaped; posterolateral portions strongly produced and slenderly clavate. Ninth sternum ( Fig. 22 View FIGURES 20 – 24 ) short; posterior margin with a rather wide trapezoidal incision, forming two broad subtriangular posterior processes produced laterally. Ninth gonostylus ( Fig. 22 View FIGURES 20 – 24 ) elongate unguiform with apex curved medially. Tenth tergite ( Fig. 21 View FIGURES 20 – 24 ) subcylindrical, short, about 0.5 times as long as 9 th tergum, with posterolateral corner strongly produced, and with tip slightly incised; tip ( Fig. 23 View FIGURES 20 – 24 ) in caudal view broad and subtriangular, with a short blunt ventral process directed medially and bearing short dense setae. Tenth sternum ( Fig. 24 View FIGURES 20 – 24 ) arched; dorsomedial process well developed, subtriangular; ventromedial process present; lateral lobe short, proximally rather thick, with tip strongly narrowed and curved medially.

Female. Unknown.

Type material. Holotype ɗ, MYANMAR: Karan Province, Dawna Range, Sud –qst, 16.4667 °N, 98.3333 °E, 25.XI. 1997 ( CAU).

Distribution. Myanmar (Karan Province).

Etymology. The specific epithet ‘ impunctatus   ’ refers to the immaculate head of the new species.

Remarks. The new species appears to be closely related to P. piaoacanus   in having the similar pale legs and the structure of the male 10 th tergite, but it can be easily separated from P. piaoacanus   by the immaculate head and the feebly developed ventral process of the male 10 th tegum. In P. piaoacanus   , the head possesses four pairs of black marks, and the ventral process of the male 10 th tergite is much longer.


China Agricultural University