Dichotomius (Dichotomius) divergens (Luederwaldt, 1923)

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 110

Dichotomius (Dichotomius) divergens (Luederwaldt, 1923) Plate 25A

Pinotus divergens Luederwaldt, 1923: 3 (original description. Type locality: Bucay, west Ecuador, 300 m).

Pinotus divergens : Luederwaldt 1929: 36 (characters in key); Blackwelder 1944: 206 (list of species from Latin America).

Dichotomius divergens : Medina et al. 2001: 138 (cited for Colombia); Carvajal et al. 2011: 320-321 (cited for Ecuador); Krajcik 2012: 91 (complete list of species); Sarmiento-Garcés and Amat-García 2014: 51 (characters in key), 71 (diagnosis).

Dichotomius (Dichotomius) divergens : Chamorro et al. 2018: 94 (cited for Ecuador).

Type specimens.

Pinotus divergens Luederwaldt, 1923. Two syntypes examined deposited at the MZUSP. Lectotype to be designated in a future work on this species group.

Syntype (♂): "W. ECUADOR / Pucay / 1-5.XII.05. F. Ohs. 5. 7. 05 [p]", "COTIPO [p, pink label]", "Pinotus ♂ / divergens L. / Lüd. det. 22 [hw]".

Syntype (♂): "W. ECUADOR / Pucay / 1-5.XII.05. F. Ohs. 5. 7. 05 [p]", "COTIPO [p, pink label]", "Pinotus ♂ / divergens L. / Lüd. det. 22 [hw]".

Distribution.

Ecuador and Colombia.

Records examined.

BOLIVAR: Bosque Protector Filo Palanga, 970 m (1 specimen MUTPL). CARCHI: Tobar Donoso, 300 m (6 specimens MECN). COTOPAXI: Guasaganda km 4, 500 m (7 specimens MQCAZ). EL ORO: Bella María Los Ingleses, 440 m (2 specimens MUTPL); Buenaventura Bajo, 500 m (1 specimen MUTPL); Reserva Jocotoco, 1250 m (1 specimen MQCAZ). ESMERALDAS: Carondelet (8 specimens MQCAZ); Colon del Ónzole (21 specimens MQCAZ; 17 specimens MECN); Chispero (11 specimens MQCAZ; 15 specimens MECN); El Progreso (3 specimens MQCAZ; 8 specimens MECN); Gallinazo (4 specimens MQCAZ); Jeyambi (7 specimens MQCAZ; 11 specimens MECN); Majua (17 specimens MQCAZ; 20 specimens MECN); Los Ajos (2 specimens MQCAZ); Pajonal (9 specimens MQCAZ; 16 specimens MECN); Palma Real (1 specimen CEMT; 25 specimens MQCAZ; 17 specimens MECN); Playa de Oro, La Tabla (26 specimens MQCAZ; 11 specimens MECN); Playa de Oro, Padre Santo (33 specimens MQCAZ; 20 specimens MECN); Playa de Oro, Pote (3 specimens CEMT; 16 specimens MQCAZ; 8 specimens MECN); Playa de Oro, Playa Rica (7 specimens MQCAZ; 3 specimens MECN); Ricauter (1 specimen MQCAZ); Salto del Bravo (9 specimens MQCAZ; 5 specimens MECN); Tjespi (19 specimens MQCAZ; 5 specimens MECN); Zabalito (4 specimens MQCAZ). GUAYAS: Pucay [= Bucay] (1 specimen MZUSP). IMBABURA: Lita, 685 m (2 specimens CEMT; 2 specimens MECN). LOS RÍOS: Río Palenque Science Ctr, 150-220 m (7 specimens MQCAZ). PICHINCHA: Chespi, 1300 m, Bellavista (2 specimens MUTPL); El Encuentro, 620 m, San Miguel de los Bancos (1 specimen MUTPL); El Tigre Río Guayllabamba, Pedro Vicente Maldonado (1 specimen MUTPL); Estación Biológica la Hesperia, 1200 m (4 specimens MUTPL); Río Guayllabamba Guayabilla, 520 m, Manduriacus (3 specimens MUTPL); Río Guayllabamba LLurimaguas, 290 m, Pedro Vicente Maldonado (2 specimens MUTPL); Mangaloma, 720 m, San Miguel de los Bancos (1 specimen MUTPL); Nanegalito, 1000 m (5 specimens MQCAZ); Río Guayllabamba Tortugo, 450 m, Pedro Vicente Maldonado (2 specimens MUTPL); Río Guayllabamba San Roque, 580 m, Pedro Vicente Maldonado (1 specimen MUTPL). SANTO DOMINGO DE LOS TSÁCHILAS: 2 km N de Alluriquin, 1070 m (1 specimen MUTPL); 47 km S de Santo Domingo (3 specimens CEMT); Río Toachi, 900 m (5 specimens MQCAZ); Tinalandia, 650 m (1 specimen CEMT).

Literature records.

GUAYAS: Bucay, 300 m ( Luederwaldt 1923: 4).

Temporal data.

Collected every month of the year.

Remarks.

Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 150-1300 m a.s.l. Collected with pitfall traps baited with carrion, cow dung, and human feces.