Coprophanaeus, Olsoufieff, 1924

Edmonds, W. D. & Zidek, J., 2010, A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae), Insecta Mundi 2010 (129), pp. 1-111 : 89

Coprophanaeus (C.) parvulus ( Olsoufieff, 1924)

Fig. 229 , 238 , 264-266

Phanaeus parvulus Olsoufieff, 1924: 67

Coprophanaeus parvulus (Olsoufieff) (recomb. by Blackwelder 1944: 209)

Type. Lectotype male (des. by Arnaud, 1982a: 116), Muséum National d’Histoire Naturelle, Paris (examined by photo).

Diagnosis. General – Dorsum black with weak green to greenish red reflections along margin of pronotum. Posterior (ventral) surface of protibia smooth except for row of coarse punctures paralleling longitudinal carina (as in Fig. 230 , arrow). Cap of anterior angle of metasternum deeply bifurcated, arms of cap embracing small, round fossa. Anterior surface of metasternum glabrous above anterior margin of metasternal disk, otherwise sparsely, weakly punctured. Basal pronotal fossae effaced ( Fig. 229 ). Length 13-17 mm.

Male – Head horn small, sides of base convergent, central process straight, length equal to that of base. Pronotal prominence ( Fig. 264 ) a pair of rounded, blister-like tumosities separated by longitudinal depression. Apical processes of parameres tooth-like, closely appressed dorsally and clearly visible laterally ( Fig. 266 ).

Female – Cephalic carina trituberculate, weakly bowed anteriorly; length of clypeus greater than that of frons. Pronotal carina only weakly raised, feebly bidentate medially.

Specimen examined – 104.

Distribution. Northern Amazonian subregion ( Guyana, Humid Guyana, Amapá and Roraima provinces) ( Fig. 238 ).

Collection Records. BRAZIL: Amazonas – Manaus (Oct) ; 26 km NE Manaus [Reserva Forestal Ducke] (Feb, Apr, Jul, Sep-Oct, Dec) ; 60 km N Manaus [Fazenda Esteio], 2 o 25’S 59 o 50’W (Jan, Jul) . Pará – Tucurí (Dec); Monte Dourado , 1 o 01’S 52 o 44’W, 100 m. COLOMBIA: Amazonas – Leticia [Tarapacá] (Jul-Aug). FRENCH GUIANA: Crique Plomb (Sep); Belizon (May); Saül (Jul) ; 7 km NW Saül, Mt. La Fumée , 3 o 39’49”N 53 o 13’19”W, 490 m (Jun) ; 18.4 km SSE Roura , 4 o 36’38”N 53 o 13’25”W, 240 m ; Nouveau Chantier; St. Jean du Maroni ; Nouragues Field Station , 4 o 05’N 52 o 40’W (Mar-Apr, Oct-Nov) . SURINAME: Paramaribo – Pallisadenweg (Jul). Sipaliwini – Central Suriname Nature Reserve, environs of Lolopasi field station, 4 o 42.91’N 56 o 12.83'’W, 80 m (Mar) ; Nassau Mt. , 4 o 48’36”N 54 o 31’16”W (Jun) ; Lely Plateau , 4 o 16’N 54 o 44’W, 640 m (Oct) . VENEZUELA: Amazonas – El Infierno [Distr. Atures] (May, Jul). Bolívar – 20 km NW Guri (Jul-Aug) ; 8 km N Guri , 200 m (Jul-Aug) ; Lago Guri Lslands, 7 o 21’N 62 o 52’W, 270 m (Jun-Jul) .

Comments. This is a widely distributed Amazonian species that often occurs in the same communities as C. jasius and C. lancifer . It is similar to C. degallieri (q.v.), but the distributions of the two species as now known are largely separate latitudinally similar to the case of Oxysternon festivum and O. macleayi ( Edmonds and Zidek 2004) . The form of the male head horn and pronotum of this species, as well as C. degallieri , is like that of the smallest males of C. telamon and C. corythus . After the distribution map ( Fig. 238 ) was prepared, we had the opportunity to examine a single specimen of C. parvulus from northern Loreto, Peru (2 o 55’26"S 74 o 49’08"W; ~ 190 km WNW Iquitos), a considerable distance (roughly 850 km) west of the range indicated in Fig. 238 . Its presence there is unsurprising, and we suspect that intervening collection sites will be identified in the future.

Figures

Figure 238. Approximate geographic distribution of Coprophanaeus (s. str.) species in the dardanus species group B.

Figure 228-236. Characters of the Coprophanaeus (s. str.) dardanus species group. 228) Oblique lateral view of metasternum (asterisk marks capped anterior angle of shield), C. degallieri. 229) Same, C. parvulus. 230) Ventral view of left protibia of C. degallieri (arrow marks row of coarse punctures). 231) Same, C. telamon. 232) Dorsal view surface of left elytron of C. ignecinctus. 233) Same, C. milon. 234) Same, C. magnoi. 235) Dorsal view of head of female C. ignecinctus. 236) Same, C. magnoi.

Figure 259-266. Coprophanaeus spp. 259-263) C. degallieri. 259-260) Female habitus. 261-262) Male habitus. 263) Aedeagus (dorsal view below; lateral view above). 264-266) C. parvulus. 264-265) Male habitus. 266) Aedeagus (dorsal view above; lateral view below).