Coprophanaeus (Metallophanaeus) pertyi ( Olsoufieff, 1924 )

Edmonds, W. D. & Zidek, J., 2010, A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae), Insecta Mundi 2010 (129), pp. 1-111 : 33-35

Coprophanaeus (Metallophanaeus) pertyi ( Olsoufieff, 1924)

Fig. 51, 57 , 62 , 80-84

Phanaeus pertyi Olsoufieff, 1924: 71

Phanaeus thalassinus sensu Felsche, 1901 , nec Perty, 1830 (syn. by Martínez and Pereira 1967: 66) Phanaeus alvarengai Pereira and d’Andretta, 1955: 253 (syn. by Martínez and Pereira 1967: 67) Coprophanaeus pertyi (Olsoufieff) (recomb. by Edmonds 1972: 841)

Type. P. pertyi – lectotype male, Zoologische Staatssammlung, Munich (des. Arnaud 2002a: 6) ; P. alvarengai – holotype male, Museo de Zoologia, Universidade de São Paulo, São Paulo .

Diagnosis. General – Paraocular areas (genae) nearly flat, strongly and densely punctate, with short carina lateral to eyes. Posteromedian portion of pronotum appearing smoother and shinier than remain- der of disk, simply punctate, less strongly and densely so near basal fossae; posterior angle normal, basal fossae small, incorporated into inner ends of distinct sulcus paralleling posterior margin. Salient anteromedian angle of metasternum (seen in profile, Fig. 57 ) rounded. Striae (x5) weakly impressed, simple; bases of striae 1-4 impressed, not distinctly fossate. Elytral interstriae weakly convex. Ventral surface of protibia lateral to longitudinal carina mostly smooth, usually with single or double row of shallow, confluent punctures paralleling longitudinal carina ( Fig. 51 , arrow). Pygidium lacking basal groove. Length 14-20 mm. Elytra dull black; pronotum, head, pygidium and venter with strong bluishgreen metallic reflections.

Male ( Fig. 80-82 ) – Head with evenly tapering long horn, base of horn abruptly swollen. Pronotum of large male with deep anteromedian concavity at summit marked by a lobate tumosity. Tips of parameres only slightly swollen laterally, lacking distinct flanges.

Female ( Fig. 83-84 ) – Cephalic carina simple, scarcely elevated, at most only weakly trituberculate. Pronotum with deep, oval anteromedian concavity, bearing strongly raised, trituberculate, quadrate process adjacent to anterior margin.

Specimens examined – 48.

Distribution. Caatinga province ( Fig. 62 ).

Collection Records. BRAZIL: Bahia – Cruz das Almas (Apr) ; Maracas (Feb); Feira de Santana (Jun). Ceará – Maranguape, Tabatinga, 38 o 43’48"W ; 04 o 00’42"S, 126 ( Mar ) ; Maranguape, Cagado, 38 o 45’05"W 04 o 01’57"S, 114 m (Apr). Maranguape, Sao Benedito, Sanctuario da Nossa Senhora de Penha, 38 o 40’48"W 03 o 52’29"S, 96 m ( Mar ) ; Juazeiro do Norte. Minas Gerais – Aguas Vermelhas (Dec). Paraíba – Mamanguape (Jul); Soledade (Mar); São José dos Cordeiros [Fazenda Almas], 7 o 28’S 36 o 53’W, 650 m (Feb). Pernambuco – Brejo Novo ( Mar ) ; Garanhuns (Mar). Piauí – São Raimundo Nonato [ Parque Nacional Serra da Capivara ] (Jan). Rio Grande do Norte – Canguaretama (Aug) ; Natal (Jun, Nov).

Comments. Phanaeus pertyi was proposed as a new name for the taxon Felsche (1901) erroneously regarded as Phanaeus thalassinus Perty ( Olsoufieff 1924) (see Comments under C. thalassinus below). Subsequently, Martínez and Pereira (1967) synonymized Phanaeus alvarengai Martínez and Pereira (1955) and P. pertyi . In accordance with Article 53.3 of the International Code of Zoological Nomenclature ( ICZN 1999), Phanaeus alvarengai Martínez and Pereira is the senior primary homonym of Phanaeus alvarengai Arnaud (1984) , currently still placed in the genus Phanaeus ( Edmonds 1994; Arnaud 2002c). The fact that Martínez and Pereira placed their taxon in the subgenus Coprophanaeus of Phanaeus does not, in accordance with Article 57.4 of the Code ( ICZN 1999), shield Arnaud’s name from unavailability, and, therefore, the name alvarengai sensu Arnaud must be replaced. We have referred the issue to the species’ author, Patrick Arnaud, for rectification.

Figures

Figure 62. Approximate geographic distribution of Coprophanaeus (Metallophanaeus) species.

Figure 50-61. Characters of the Coprophanaeus subgenus Metallophanaeus. 50) Ventral view of left protibia of C. saphirinus. 51) Same, C. pertyi. 52) Dorsal view of posterior portion of pronotum of C. thalassinus. 53) Same, C. pessoai. 54) Dorsal view of left elytron of C. punctatus. 55) Same, C. pessoai. 56) Posterior view of pygidium of C. horus. 57) Lateral view of metasternal angle of C. pertyi. 58) Same, C. pessoai. 59) Same, C. saphirinus. 60) Dorsal view of right side of head of C. horus. 61) Same, C. pessoai.

Figure 80-84. Coprophanaeus pertyi. 80-81) Male habitus. 82) Aedeagus (dorsal view on left; lateral view on right). 83-84) Female habitus.