Digitonthophagus eucatta Génier

Génier, François & Moretto, Philippe, 2017, Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 4248 (1), pp. 1-110: 29-33

publication ID

10.5281/zenodo.439444

publication LSID

lsid:zoobank.org:pub:71F0AC03-C1FB-40AB-8532-99A638FC91E9

persistent identifier

http://treatment.plazi.org/id/3D5987B7-0763-FFB3-FF17-A12C0273FC96

treatment provided by

Plazi

scientific name

Digitonthophagus eucatta Génier
status

new species

Digitonthophagus eucatta Génier  , new species

http://zoobank.org/urn:lsid:zoobank.org:act: D66876 EF-FA95-493B-B87B-1 D88858 C8406 ( Figs. 11View FIGURES 11 – 14, 31–32View FIGURES 31 – 38. 31, 33, 35, 37, 59View FIGURES 55 – 62, 75View FIGURES 71 – 78, 95–96, 123, 147–149, 187; Map 6)

Type locality. Toulfé (13°53’33”N 01°53’39”W), 325 m, Loroum , Burkina Faso.GoogleMaps 

Diagnosis. Species distributed from Oman westward; left lateral apophysis of FLP sclerite large ( Fig. 147View FIGURES 147 – 158).

Description. Holotype ♂ ( Fig. 11View FIGURES 11 – 14). Measurements. Length 11.5 mm, width 6.5 mm. Head ( Figs. 31–32View FIGURES 31 – 38. 31, 33, 35, 37). Anterior clypeal edge straight on median fourth in dorsal view; clypeofrontal carina broadly arcuate and interrupted at gena. Vertex lacking median tubercle; surface with punctures moderate, scabrous anteriorly, separated by less than a diameter. Horns moderately long, parallel on apical half in frontal view, parallel sided on basal four-fifths, slightly tapering externally before apex; posterointernal edge produced into a low, angular projection; apicointernal surface lacking granules, with a few small scabrous punctures; genal edge upturned and distinctly angulate on anterior third, forming a broad angle with clypeal edge. Pronotum ( Fig. 59View FIGURES 55 – 62). Surface with granulate punctures restricted to anterior half medially, with smaller, irregularly-scattered, simple punctures on posterior half of disc; punctures lacking posterolaterally, with more-or-less distinct minute punctures throughout; anteromedian tubercle atrophied, simply round in lateral view, median longitudinal sulcus well defined; surface behind the eyes slightly concave, surface of anterior angles concave, anterior angles unmodified; anterior half of lateral edge arcuate in dorsal and slightly sinuous anteriorly in lateral view; posterior angles simply arcuate in dorsal view. Anterior hypomeral ridge straight, abruptly curved before anterior angle, anterior hypomeral depression surface light in color medially. Elytra ( Fig. 11View FIGURES 11 – 14). Intervals 2 and 4 with few fine granules on apical declivity only. Legs. Protibial apicointernal tooth enlarged, with dorsal ridge extending to apex. Aedeagus ( Fig. 123View FIGURES 119 – 134). Parameres with dorsal and ventral edges slightly diverging toward apex in lateral view. Internal sac sclerites ( Figs. 147–149View FIGURES 147 – 158). Axial sclerite moderately sclerotized, shorter than subaxial sclerite and almost straight. Subaxial sclerite, larger basally and abruptly tapering in line with apex of axial sclerite, extending straight and interrupted approximately at apex of frontolateral peripheral sclerite apical portion, with fine villi on apical sixth. Frontolateral peripheral sclerite basoventral apophysis moderately developed, angular in lateral view; with a single acute medioventral carinae; right lateral fold produced into a uneven bifurcate process, lateral edge straight; left lateral apophysis present, large; left lateral lobe absent; subapicodorsal lobe well sclerotized, entirely covering apical lobe, with villi much shorter than apical lobe villi, right edge obliquely truncated before apex; apical lobe truncated apically, apical villi regular in shape; subapicoventral lobe atrophied, oriented ventrally

Variation. Measurements (107 ♂♂, 118 ♀♀). Length: male 7.0– 12.5 mm (10.4 ± 1.0 mm), female 8.0– 12.5 mm (10.6 ± 1.0 mm). Female allotype. Cephalic outline in dorsal view as in Fig. 75View FIGURES 71 – 78; vertex with a broadly arcuate transverse carina, dorsal edge straight in frontal view, lateral portion subangular and sloping down posteriorly; anterior pronotal tubercles well developed, external lateral edges rather strongly widening in dorsal view (Fig. 95), anterolateral surface deeply, obliquely concave, anterosuperior edge slightly arcuate in dorsal view, lateral portion of anterosuperior edge slightly upturned (Fig. 96). Protibia short, with external teeth more robust.

Primary type data. Holotype ♂ ( CMNC): [ BURKINA FASO: LORUM / Toulfé, 325m / 13°53’33”N 001°53’39”O / 17.VII.2006, zone sahélienne / steppe arborée, piège lumineux / F. & S. Génier, 2006-41]; [COLÉOPTÈRES / DU / BURKINA FASO / BF015417] barcode label; [HOLOTYPE ♂ / Digitonthophagus  / eucatta  n.sp. / des. F. Génier, 2016] red card. 

MAP 6. Distribution of Digitonthophagus eucatta 

Material examined (403 ♂♂, 237 ♀♀), distribution (Map 6): BENIN  : ATAKORA, Batia (10°53'N, 1°29'E), 24.vii.2009, B. Rainon— 1 ♂ (paratype) ( CMD)GoogleMaps  ; BURKINA FASO: CENTRE, Ouagadougou (12°22'N, 1°31'W), viii.1977, P. Ardoin— 2 ♀♀, 7 ♂♂ (9 paratypes) ( FETC)GoogleMaps  ; CENTRE-NORD, 19 km N Kaya , 314 m (13°8'46.2''N, 0°52'42.1''W), 16.viii.2012, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; CENTRE-OUEST, Forêt de Sorobouli [= Forêt Classée de Baporo ], 270 m (11°47'44''N, 2°53'25''W), 10–15.viii.2005, P. Moretto— 31 ♀♀, 41 ♂♂ (72 paratypes) ( FETC, PMOC)GoogleMaps  ; Koudougou (12°15'N, 2°22'W), vii.1998, J.B. Yaméogo— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; CENTRE- SUD, Réserve de Nazinga (11°9'N, 1°36'W), 9–12.vi.2005, S. Rojkoff— 2 ♀♀, 1 ♂ (paratypes) ( PCSR)GoogleMaps  ; EST, 25 km SE Pama (11°7'49''N, 0°52'38''E), 3–24.ix.1988, F. Génier & M. Sanborne— 2 ♀♀, 6 ♂♂ (8 paratypes) ( CMNC)GoogleMaps  ; Kompienga , 20 km S Pama (11°4'37''N, 0°40'50''E), 1–16.vi.1988, Sanborne, Landry & Tou— 2 ♂♂ (2 paratypes) ( CMNC)GoogleMaps  ; Kompienga , 20 km S Pama (11°4'37''N, 0°40'50''E), 3–24.ix.1988, F. Génier & M. Sanborne— 2 ♀♀, 4 ♂♂ (6 paratypes) ( CMNC)GoogleMaps  ; HAUTS-BASSINS, Aéroport, Bobo-Dioulasso (11°9'47''N, 4°19'12''W), 27.ix.2012, P. Moretto— 1 ♀, 2 ♂♂ (3 paratypes) ( PMOC)GoogleMaps  ; Bobo-Dioulasso (11°12'N, 4°18'W), 29.iv.1953, Hamon— 7 ♀♀, 7 ♂♂ (14 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Bobo-Dioulasso (11°12'N, 4°18'W), 7.v.1953, Hamon— 1 ♀, 2 ♂♂ (3 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Matourkou (11°5'N, 4°22'W), 15.vi.2005, S. Rojkoff— 3 ♂♂ (3 paratypes) ( CMD, PCSR)GoogleMaps  ; site de Samandéni, Bama , 330 m (11°22'24.5''N, 4°33'57''W), vii.2012, F. Rousset— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; NORD, 8 km SW Yako, Passoré , 320 m (12°51'44''N, 2°18'35''W), 7.viii.2005, P. Moretto— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; Toulfé, Lorum , 330 m (13°54'13''N, 1°54'19''W), 9.viii.2005, P. Moretto— 5 ♀♀, 6 ♂♂ (11 paratypes) ( PMOC)GoogleMaps  ; PLATEAU-CENTRAL, Piste de Bomboré (12°15'N, 0°53'W), 11.viii.2012, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; Ziniare (12°35'N, 1°18'W), 19.vi.2005, S. Rojkoff— 1 ♀, 4 ♂♂ (5 paratypes) ( PCSR, PMOC)GoogleMaps  ; SAHEL, 3,7 km NW piste vers Essakane, N’djomga, Dori , 287 m (14°5'20.9''N, 0°1'36.3''W), 1– 5.ix.2013, P. Moretto— 2 ♂♂ (2 paratypes) ( PMOC)GoogleMaps  ; après le pont, route Essakane-Dori, Gorum-Gorum , 271 m (14°9'32.8''N, 0°1'20.4''E), 7.viii.2012, P. Moretto— 2 ♂♂ (2 paratypes) ( PMOC)GoogleMaps  ; Conseil régional, Dori , 283 m (14°2'4.5''N, 0°3'10.5''W), 30.viii.2013, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; Dori (14°2'N, 0°1'W), iv.2007, [anonymous]— 1 ♂ (paratype) ( JFJC)GoogleMaps  ; Dori (14°2'N, 0°1'W), 24.viii.2012, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; Essakane, Gorum-Gorum , 264 m (14°22'40.2''N, 0°5'47.7''E), 13–15.viii.2012, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; Essakane, Gorum-Gorum , 264 m (14°22'40.2''N, 0°5'47.7''E), 22–24.viii.2012, P. Moretto— 2 ♀♀, 2 ♂♂ (4 paratypes) ( PMOC)GoogleMaps  ; Essakane, Gorum-Gorum , 264 m (14°22'40.2''N, 0°5'47.7''E), 10–13.ix.2012, P. Moretto— 1 ♀, 2 ♂♂ (3 paratypes) ( PMOC)GoogleMaps  ; Essakane, Gorum-Gorum , 264 m (14°22'40.2''N, 0°5'47.7''E), 2–4.x.2012, P. Moretto— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; N’djomga, Dori (14°3'55.7''N, 0°3'3.4''W), 24–27.viii.2013, P. Moretto— 4 ♂♂ (4 paratypes) ( PMOC)GoogleMaps  ; SUD-OUEST, Bondigui , 324 m (10°53'56.5''N, 3°34'15''W), v.2012, F. Rousset— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; BALÉ, Boromo , 250 m (11°45'6''N, 2°51'58''W), 10.viii.2005, F. Génier (2005-10)— 1 ♀, 4 ♂♂ (5 paratypes) ( FGICAbout FGIC)GoogleMaps  ; GOURMA, Fada N’Gourma , 300 m (12°3'5''N, 0°21'27''E), 14.vii.2006, F. & S. Génier (2006-35)— 1 ♂ (paratype) ( FGICAbout FGIC)GoogleMaps  ; KOMPIENGA, 15 km E Nadiagou , 155 m (11°4'31''N, 0°56'8''E), 22.viii.2005, F. Génier (2005-40)— 1 ♂ (paratype) ( FGICAbout FGIC)GoogleMaps  ; Kompienga , 160 m (11°5'5''N, 0°42'42''E), 21.viii.2005, F. Génier (2005-39)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Pama , 230 m (11°17'N, 0°42'59''E), 24.viii.2005, F. Génier (2005-44)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Pama , 230 m (11°17'N, 0°42'59''E), 25.viii.2005, F. Génier (2005-45)— 1 ♂ (paratype) ( FGICAbout FGIC)GoogleMaps  ; Pama , 230 m (11°17'N, 0°42'59''E), 26–27.viii.2005, F. Génier (2005-49)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; LOROUM, Toulfé , 330 m (13°54'16''N, 1°54'19''W), 9.viii.2005, F. Génier (2005-08)— 1 ♀, 2 ♂♂ (3 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Toulfé , 330 m (13°54'16''N, 1°54'19''W), 16.vii.2006, F. & S. Génier (2006-38)— 1 ♀, 3 ♂♂ (4 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Toulfé , 330 m (13°54'16''N, 1°54'19''W), 16.vii.2006, F. & S. Génier (2006-40)— 1 ♀, 1 ♂ (paratypes) ( FGICAbout FGIC)GoogleMaps  ; Toulfé , 330 m (13°54'16''N, 1°54'19''W), 17.vii.2006, F. & S. Génier (2006-41)— 22 ♀♀, 21 ♂♂ (holotype, allotype, 41 paratypes) ( CMNC, FGICAbout FGIC)GoogleMaps  ; Toulfé , 330 m (13°54'16''N, 1°54'19''W), 18.vii.2006, F. & S. Génier (2006-42)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Toulfé , 330 m (13°54'16''N, 1°54'19''W), 18.vii.2006, F. & S. Génier (2006-43)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; NAHOURI, Forêt de Nazinga , 275 m (11°9'24''N, 1°36'44''W), 20.vii.2006, F. & S. Génier (2006-46)— 1 ♂ (paratype) ( FGICAbout FGIC)GoogleMaps  ; Forêt de Nazinga, Akwazena , 275 m (11°9'24''N, 1°36'44''W), 26.vii.2006, F. & S. Génier (2006-76)— 3 ♂♂ (3 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Forêt de Nazinga, Boulieselo , 310 m (11°11'50''N, 1°35'9''W), 27.vii.2006, F. & S. Génier (2006-82)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Forêt de Nazinga, Kouzougou , 285 m (11°9'17''N, 1°32'10''W), 21.vii.2006, F. & S. Génier (2006-47)— 1 ♀, 4 ♂♂ (5 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Forêt de Nazinga, Naguio , 270 m (11°7'52''N, 1°34'38''W), 24.vii.2006, F. & S. Génier (2006-66)— 1 ♂ (paratype) ( FGICAbout FGIC)GoogleMaps  ; Tiakané , 320 m (11°11'37''N, 1°14'10''W), 20.viii.2005, F. Génier (2005-29)— 2 ♀♀ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; PASSORÉ, 8 km SW Yako , 320 m (12°51'44''N, 2°18'35''W), 7.viii.2005, F. Génier (2005-03)— 4 ♀♀, 2 ♂♂ (6 paratypes) ( FGICAbout FGIC)GoogleMaps  ; 8 km S.O. Yako, 320 m (12°51'44''N, 2°18'35''W), 7.viii.2005, F. Génier (2005-04)—1 ♀ (paratype) (FGIC); 8 km S.O. Yako, 320 m (12°51'44''N, 2°18'35''W), 7.viii.2005, F. Génier (2005-02)—7 ♂♂ (7 paratypes) (FGIC); SANGUIÉ, Forêt de Sorobouli , 270 m (11°47'44''N, 2°53'25''W), 13.viii.2005, F. Génier (2005-14)— 1 ♀, 1 ♂ (paratypes) ( FGICAbout FGIC)GoogleMaps  ; Forêt de Sorobouli , 270 m (11°47'44''N, 2°53'25''W), 14.viii.2005, F. Génier (2005-18)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Forêt de Sorobouli , 270 m (11°47'44''N, 2°53'25''W), 15.viii.2005, F. Génier (2005-19)— 2 ♀♀, 2 ♂♂ (4 paratypes) ( FGICAbout FGIC)GoogleMaps  ; CAMEROON: EXTREME NORTH, Guétalé, Koza (10°53'N, 13°54'E), 13.vi.1967, [anonymous]— 1 ♀ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; Maroua (10°35'N, 14°19'E), vii.1989, D. Bernaud— 3 ♀♀, 5 ♂♂ (8 paratypes) ( CMD)GoogleMaps  ; Maroua (10°35'N, 14°19'E), vii.1979, G. Onore— 2 ♀♀, 2 ♂♂ (4 paratypes) ( PHWC)GoogleMaps  ; Maroua (10°35'N, 14°19'E), 5.vii.1978, [anonymous]— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; Waza (11°24'N, 14°34'E), 18.viii.2006, C. Vanderbergh— 4 ♀♀, 2 ♂♂ (6 paratypes) ( PMOC)GoogleMaps  ; Waza National Park (11°15'N, 14°39'E), 18.viii.2006, C. Vanderbergh— 3 ♀♀, 4 ♂♂ (7 paratypes) ( PMOC)GoogleMaps  ; NORTH, Garoua (9°18'N, 13°24'E), x.1973, B. de Miré— 1 ♀ (paratype) ( MNHNAbout MNHN); CHADGoogleMaps  : HADJER-LAMIS, Sitje , rive sud Lac Tchad (12°53'N, 14°52'E), 11.viii.2006, C. Vanderbergh— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; LOGONE ORIENTAL, Komé, 80 km Moundou (8°30'N, 16°41'E), vii.2002, T. Garnier— 2 ♂♂ (2 paratypes) ( PMOC)GoogleMaps  ; N’DJAMENA, Farcha (12°7'N, 14°59'E), 28.vi.1967, [anonymous]— 4 ♀♀, 8 ♂♂ (12 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Fort Lamy [= N’Djamena _] (12°7'N, 15°3'E), [no date], M. André— 3 ♀♀, 5 ♂♂ (8 paratypes) ( PMOC)GoogleMaps  ; OUADAI, Massif de Kabla (13°25'N, 21°9'W), 5.viii.1949, P. de Miré— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; ERITREA: ANSEBA, Hagaz (15°42'N, 38°16'E), vii.2003, M. Forti— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; GASH-BARKA, 18 km E Akordat (15°33'N, 38°1'E), vii.2003, M. Forti— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; Tesseney , 1200 m (15°6'36''N, 36°39'27''E), 30.viii.2001, L. & M. Stalmans— 2 ♀♀, 4 ♂♂ (6 paratypes) ( CADB, PMOC)GoogleMaps  ; NORTHERN RED SEA, Nefasit (15°20'N, 39°4'E), vii.2002, Czeppel & Forti— 2 ♀♀, 5 ♂♂ (7 paratypes) ( FETC)GoogleMaps  ; ETHIOPIA: AFAR, Yangudi Rassa National Park (10°33'N, 40°59'E), 9/2/2006, S. Rojkoff— 1 ♀, 1 ♂ (paratypes) ( CMD)GoogleMaps  ; GAMBIA: CENTRAL RIVER, Lower Saloum District, Ballangharr (13°39'N, 15°25'W), 23.viii.1997, D.J. Mann— 2 ♀♀, 5 ♂♂ (7 paratypes) ( OUMNH)GoogleMaps  ; MALI: KIDAL  , Adrar des Ifoghas (19°32'N, 1°35'E), viii.1941, Volkonsky— 2 ♂♂ (2 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Etambar, Kidal (18°26'N, 1°25'E), ix.1941, M. Volkoneky— 1 ♀ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; SEGOU, [Station centrale de] Kogoni (14°44'N, 6°2'W), 19.ix.1955, J.-G. Pointel— 2 ♀♀, 1 ♂ (paratypes) ( MNHNAbout MNHN)GoogleMaps  ; [Station centrale de] Kogoni (14°44'N, 6°2'W), 7.x.1955, J.-G. Pointel— 2 ♀♀ (2 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; [Station centrale de] Kogoni (14°44'N, 6°2'W), 11.x.1955, J.-G. Pointel— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; [Station centrale de] Kogoni (14°44'N, 6°2'W), 24.x.1955, J.-G. Pointel— 1 ♀, 1 ♂ (paratypes) ( MNHNAbout MNHN)GoogleMaps  ; [Station centrale de] Kogoni (14°44'N, 6°2'W), 27.x.1955, J.-G. Pointel— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; [Station centrale de] Kogoni (14°44'N, 6°2'W), 2.xi.1955, J.-G. Pointel— 1 ♀, 2 ♂♂ (3 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; [Station centrale de] Kogoni (14°44'N, 6°2'W), 21.ix.1959, J.-G. Pointel— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; MAURITANIA: GUIDIMAKA, [unspecified locality] (15°22'N, 12°6'W), 1908, G. Audan— 4 ♀♀, 7 ♂♂ (11 paratypes) ( FGICAbout FGIC, MNHNAbout MNHN)GoogleMaps  ; HODH EL GHARBI, Tamchekett (17°14'44''N, 10°40'11''W), x.1947, B. de Miré— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; SÉLIBABY, Tachoot [= Tassota Barene ] (15°26'N, 12°16'W), 20.x.1941, [anonymous]— 1 ♀ (paratype) ( OMOC); NIGERGoogleMaps  : AGADEZ  , Agadez (16°58'N, 7°59'E), 20.viii.1989, [anonymous]— 1 ♀, 1 ♂ (paratypes) ( OMOC)GoogleMaps  ; DOSSO, Toudou (14°7'N, 3°54'E), 10.viii.1989, [anonymous]— 1 ♀, 1 ♂ (paratypes) ( OMOC)GoogleMaps  ; MARADI  , Maradi (13°29'N, 7°6'E), [no date], J. Roggeman— 2 ♂♂ (2 paratypes) ( BMNH)GoogleMaps  ; TAHOUA, Entre Madaoua et Bouza (14°15'N, 6°0'E), 3.vii.1980, P. Leblanc— 2 ♂♂ (2 paratypes) ( FETC)GoogleMaps  ; TILLABERI, Koure (13°20'N, 2°35'E), 19–21.ix.2003, Josso, Juhel & Monfort— 1 ♂ (paratype) ( JFJC)GoogleMaps  ; NIGERIA: KANO, [unspecified locality] (11°59'47''N, 8°31'0''E), v.1952, W.E. S. Merrett— 4 ♂♂ (4 paratypes) ( BMNH)GoogleMaps  ; [unspecified locality] (11°59'47''N, 8°31'0''E), vii.1954, W.E.S. Merrett—2 ♂♂ (2 paratypes) (BMNH); [unspecified locality] (11°59'47''N, 8°31'0''E), v.1953, W.E.S. Merrett—4 ♂♂ (4 paratypes) (BMNH); [unspecified locality] (11°59'47''N, 8°31'0''E), vi.1953, W.E.S. Merrett—5 ♂♂ (5 paratypes) (BMNH); [unspecified locality] (11°59'47''N, 8°31'0''E), vi.1954, W.E. S. Merrett— 1 ♂ (paratype) ( BMNH); OMAN: MINTAQAT MASQATGoogleMaps  , Mascat (23°36'48''N, 58°35'36''E), ix–x, Maindron— 2 ♂♂ (2 paratypes) ( BMNH)GoogleMaps  ; SENEGAL: DAKAR, Cité Gendarmerie, Keur Massar (14°45'51''N, 17°16'45''W), 29.viii.2012, I. de Dinechin— 1 ♂ (paratype) ( CMD)GoogleMaps  ; Dakar (14°46'N, 17°22'W), 1914, Chissadon— 3 ♀♀, 4 ♂♂ (7 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Dakar (14°46'N, 17°22'W), 1902, Waterlot— 1 ♂ (paratype) ( MNHNAbout MNHN); Entrée du parc Marine, Keur MassarGoogleMaps  , Dakar (14°47'11''N, 17°18'43''W), viii.2015, I. de Dinechin— 1 ♀, 1 ♂ (paratypes) ( CMD)GoogleMaps  ; Keur Ndiaye Lô (14°45'25''N, 17°14'11''W), ix.2013, Local collectors— 1 ♂ (paratype) ( CMD)GoogleMaps  ; Keur Ndiaye Lô (14°45'25''N, 17°14'11''W), 18.viii.2013, I. de Dinechin— 1 ♂ (paratype) ( CMD)GoogleMaps  ; Sebikotane (14°45'N, 17°8'W), viii.1971, A. Villiers— 1 ♂ (paratype) ( OMOC)GoogleMaps  ; DIOURBEL, Bambey (14°42'N, 16°28'W), 1945, J. Risbec— 4 ♀♀, 5 ♂♂ (9 paratypes) ( BMNH)GoogleMaps  ; Bambey (14°42'N, 16°28'W), viii.1939, J. Risbec— 3 ♂♂ (3 paratypes) ( BMNH)GoogleMaps  ; FATICK, Diouroup (14°21'N, 16°32'W), viii.2008, [anonymous]— 1 ♀, 1 ♂ (paratypes) ( OMOC)GoogleMaps  ; Diouroup (14°21'N, 16°32'W), 14– 18.viii.2007, P. Moretto— 11 ♀♀, 17 ♂♂ (28 paratypes) ( FGICAbout FGIC, PMOC)GoogleMaps  ; Diouroup (14°21'N, 16°32'W), 19– 20.xi.2009, P. Moretto— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; KAFFRINE, Tamba (13°53'49''N, 15°15'33''W), 2.vii.1948, [anonymous]— 1 ♀, 1 ♂ (paratypes) ( CNCAbout CNC)GoogleMaps  ; KAOLAK, Missirah , 45 m (13°59'26''N, 15°7'4''W), 16.vii.2007, F. Génier (2007-01)— 3 ♀♀, 9 ♂♂ (12 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Missirah , 45 m (13°59'26''N, 15°7'4''W), 17.vii.2007, F. Génier (2007-02)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Nioro du Rip (13°44'43''N, 15°46'23''W), 10– 11.viii.2009, P. Moretto— 1 ♀, 1 ♂ (paratypes) ( PMOC)GoogleMaps  ; LOUGA, Linguère (15°24'N, 15°7'W), ix.1967GoogleMaps  , A. Descarpentries, T. Leye & A. Villiers— 14 ♀♀, 9 ♂♂ (23 paratypes) ( MNHNAbout MNHN); Linguère (15°24'N, 15°7'W), 5– 15.ix.1967GoogleMaps  , A. Descarpentries, T. Leye & A. Villiers— 1 ♀, 1 ♂ (paratypes) ( MNHNAbout MNHN)  ; Louga (15°37'N, 16°13'W), 1945, Risbec— 1 ♂ (paratype) ( MNHNAbout MNHN)GoogleMaps  ; Ndam-Dam (15°36'22''N, 16°23'35''W), xi.1967, A. Descarpentries, T. Leye & A. Villiers— 2 ♀♀, 5 ♂♂ (7 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; SAINT-LOUIS, Commune de Saint-Louis, 7 m (16°4'34''N, 16°22'40''W), 27.viii.2009, F. Génier (2009-33)— 1 ♀ (paratype) ( FGICAbout FGIC)GoogleMaps  ; Commune de Saint-Louis, 7 m (16°4'34''N, 16°22'40''W), 27.viii.2009, P. Moretto— 1 ♀ (paratype) ( PMOC)GoogleMaps  ; Parc National des Oiseaux du Djoudj (16°24'N, 16°14'W), x.1996, [anonymous]— 1 ♂ (paratype) ( PSCW)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), xi.1967, A. Descarpentries, T. Leye & A. Villiers— 8 ♀♀, 10 ♂♂ (18 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 28–31.viii.2009, F. Génier (2009-38)— 4 ♀♀, 5 ♂♂ (9 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 28–30.viii.2009, F. Génier (2009-40)— 2 ♀♀, 2 ♂♂ (4 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 28.viii.2009, F. Génier (2009-36)— 1 ♂ (paratype) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 28.viii– 1.ix.2009, P. Moretto— 15 ♀♀, 15 ♂♂ (30 paratypes) ( PMOC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 30.viii.2009, F. Génier (2009-43)— 1 ♀, 2 ♂♂ (3 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 30.viii.2009, F. Génier (2009-44)— 2 ♀♀, 5 ♂♂ (7 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 31.viii.2009, F. Génier (2009- 48)— 2 ♀♀, 4 ♂♂ (6 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 1.ix.2009, F. Génier (2009-49)— 1 ♀, 2 ♂♂ (3 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Richard-Toll (16°28'N, 15°41'W), 1.ix.2009, F. Génier (2009-50)— 2 ♀♀, 1 ♂ (paratypes) ( FGICAbout FGIC, GWPC)GoogleMaps  ; TAMBACOUNDA, Botou , 52 m (13°48'16''N, 13°34'47''W), 11.vii.2008, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; Dienoun Diala (13°12'47''N, 13°6'44''W), 29.vii.1995, P. Moretto— 1 ♂ (paratype) ( PMOC)GoogleMaps  ; THIÈS, 4.5 km E Joal-Fadiouth , piste de Samba Dia, 11 m (14°11'3''N, 16°48'28''W), 9.viii.2007, F. Génier (2007-65)— 2 ♂♂ (2 paratypes) ( FGICAbout FGIC)GoogleMaps  ; 4.5 km E Joal-Fadiouth , piste de Samba Dia, 11 m (14°11'3''N, 16°48'28''W), 10. viii.2 0 0 7, F. Génier & P. Moretto (2007-67)— 6 ♀♀ (6 paratypes) ( FGICAbout FGIC)GoogleMaps  ; 4.5 km E Joal-Fadiouth , piste de Samba Dia, 11 m (14°11'3''N, 16°48'28''W), 10.viii.2007, F. Génier (2007-68)— 1 ♀, 1 ♂ (paratypes) ( FGICAbout FGIC)GoogleMaps  ; Ngazobil (14°12'N, 16°52'W), viii.1971, A. Villiers— 1 ♂ (paratype) ( OMOC)GoogleMaps  ; Nianing , 8 m (14°19'N, 16°55'43''W), 8.viii.2007, F. Génier (2007-63)— 2 ♀♀, 3 ♂♂ (5 paratypes) ( FGICAbout FGIC)GoogleMaps  ; Nianing , 8 m (14°19'N, 16°55'43''W), 8–9.vii.2008, P. Moretto— 2 ♂♂ (2 paratypes) ( PMOC)GoogleMaps  ; SUDAN: GHARB DARFUR, Zalingei , 1021 m (12°54'N, 23°29'E), vii.1984, P. Ruse— 1 ♀, 1 ♂ (paratypes) ( BMNH)GoogleMaps  ; JANUB DARFUR, Buram (10°51'N, 25°9'E), viii.1984, P. Ruse— 2 ♀♀, 1 ♂ (paratypes) ( BMNH)GoogleMaps  ; KHARTOUM, Environs de Khartoum (15°35'N, 32°32'E), [no date], [anonymous]— 1 ♀, 5 ♂♂ (6 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; Khartoum (15°35'N, 32°32'E), [no date], [anonymous]— 1 ♀, 2 ♂♂ (3 paratypes) ( MNHNAbout MNHN)GoogleMaps  ; SENNAR, Sennar (13°34'N, 33°34'E), [no date], Parreys— 1 ♂ (paratype) ( SMFAbout SMF)GoogleMaps  .

Etymology. Eucatta  is a noun indicative of the close relationship of this African species with the Oriental D. catta  .

Natural history. Specimens with data mostly collected in disturbed area and pastures from southern Sahelian and Sudanese zone. This species is common at light traps (mercury vapor and ultraviolet light). Some specimens were collected in cow, donkey, elephant, and zebu ( Bos indices  ) dung and pitfall traps baited with human dung, toad carcasses, and rumen content.

MNHN

Museum National d'Histoire Naturelle

FGIC

Francois Genier

CNC

Canadian National Collection of Insects, Arachnids, and Nematodes

SMF

Forschungsinstitut und Natur-Museum Senckenberg