Potwarmus flynni, Antoñanzas, 2009

Antoñanzas, Raquel López, 2009, First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea), Zoological Journal of the Linnean Society 156, pp. 664-679 : 672

PROGONOMYS DEBRUIJNI JACOBS, 1978

This species was erected by Jacobs (1978) on the basis of 50 isolated teeth from the Late Miocene YGSP locality 182A, Dhok Pathan Formation, Punjab, Pakistan. The holotype (YGSP 7739, Jacobs, 1978: figs. 15–17) is a left M1. The morphology of the teeth of Progonomys debruijni is very different from that of Potwarmus flynni sp. nov. The M1 of P. debruijni shows the basic murine pattern of three chevrons, lacking in AJ-7, which has a much smaller anterocone and enterostyle, the cusps more alternate, and lacks a well-developed anterostyle. The M2s of P. debruijni are also very distinct, having a prominent anterostyle, which is lacking in AJ11 and AJ12, and a large enterostyle connected to the protocone, which is smaller and isolated in AJ11 and AJ12. The m1 of P. debruijni differs from AJ13, for example, in having a well-developed bilobed anteroconid with a central X-shaped connection pattern, and a variable number of cingular cusps (absent in AJ13).