Aulacus insularis, Jennings & Parslow & Austin, 2018
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publication ID |
https://doi.org/ 10.11646/zootaxa.4538.1.1 |
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publication LSID |
lsid:zoobank.org:pub:E4D8D897-3CBE-4FA6-95C3-143A945229A4 |
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DOI |
https://doi.org/10.5281/zenodo.3798390 |
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persistent identifier |
https://treatment.plazi.org/id/3E4187A0-1755-4E00-FF54-F963FA0CFA39 |
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treatment provided by |
Plazi |
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scientific name |
Aulacus insularis |
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Key to Australian Aulacus Jurine View in CoL View at ENA (females only)
We have not included in the key four species known only from males, viz., A. flindersbaudini Jennings, Austin & Stevens , a largely reddish brown species which is endemic to Kangaroo Island, South Australia; A. insularis sp. nov., which is readily separated from all other Australian species by the two distinctive cream bands on metasomal segments 2 and 3 ( Fig. 26A View FIGURE 26 ); A. naumanni sp. nov., which has a colour combination different from all other Australian species (see comments below under this species), and A. vespiformis Kieffer , which has distinctive banding on the metathorax ( Figs 50A, B View FIGURE 50 ) and anterior half of fore wing distinctively fuscous ( Fig. 50A View FIGURE 50 ).
1 Mesoscutum in lateral view angular antero-dorsally (e.g., Fig. 18E View FIGURE 18 ).......................................... 2
- Mesoscutum in lateral view rounded antero-dorsally (e.g., Fig. 2E View FIGURE 2 ).......................................... 20
2(1) Metasoma clavate ( Figs 9A, B View FIGURE 9 ); face punctate-reticulate; fore wing vein 2-Rs+M very long; mesothorax pale orange in the anterior half and largely dark brown in the posterior half; brown band either side of fore wing veins 1-Rs and 1-M and apically in costal cell [NSW and Queensland]................................................ A. bicolor sp. nov.
- Metasoma ovate (e.g., Figs 29A, B View FIGURE 29 ); other features not in combination........................................ 3
3(2) Fore wing with brown infuscation (spot) at apex (e.g., Fig. 1 View FIGURE 1 )............................................... 4
- Fore wing lacking brown infuscation (spot) at apex...................................................... 15
4(3) Hind wing venation incomplete, variable loss of one or more veins M+Cu, Cu, r-m and 2-M....................... 5
- Hind wing venation complete (e.g., Fig. 1 View FIGURE 1 ).............................................................. 7
5(4) Hind tarsal claw long, 3.0× length tarsomere 5; face rugose with a few punctures medially; head and mesosoma black, metasoma orange; ovipositor guide about 2/3 rd distance along coxa, oblique ( Fig. 51F View FIGURE 51 ) [Victoria]..... A. walkeri sp. nov.
- Hind tarsal claw short, about half length of tarsomere 5; other features not in combination......................... 6
6(5) Face rugose; head and mesosoma blackish brown, metasomal T1 and T2 orange, remainder of metasoma blackish brown ( Figs 29A, B View FIGURE 29 ); ovipositor guide about ¾ way along coxae, oblique, distal fringe of long setae, long setae in groove [NSW]............................................................................ A. kiwarrakensis sp. nov.
- Face punctate-rugose; head and mesosoma black, metasoma orange ( Figs 30A, B View FIGURE 30 ); ovipositor guide medial, oblique, setae on lower distal margin, lacking setae in groove [Tasmania]...................................... A. leai sp. nov.
7(4) Ovipositor guide on hind coxa transverse (e.g., Fig. 5D View FIGURE 5 ).................................................... 8
- Ovipositor guide on hind coxa oblique (e.g., Fig. 51F View FIGURE 51 )..................................................... 9
8(7) Ovipositor guide on hind coxa distal; face rugulose-reticulate; frons without lateral medial carina above toruli; head black, meso- and metasoma orange ( Figs 36A, B View FIGURE 36 ) [Victoria]....................................... A. neboissi sp. nov.
- Ovipositor guide on hind coxa medial; face punctate-rugose; frons with lateral medial carina above toruli; largely dark black-brown species ( Figs 25A, B View FIGURE 25 ) [Queensland]........................................... A. hackeri sp. nov.
9(7) Ovipositor guide on hind coxa distal (e.g., Fig. 12F View FIGURE 12 )...................................................... 10
- Ovipositor guide on hind coxa medial (e.g., Fig. 7F View FIGURE 7 )...................................................... 14
10(9) Frons with lateral medial carina above toruli; face imbricate; head, scape, pedicel, propleuron, legs and metasoma orange, except for variable amounts of black on metasomal T1, mesosoma black ( Figs 18A, B View FIGURE 18 ) [NSW and Victoria]............................................................................................. A. elegans sp. nov.
- Frons without lateral medial carina above toruli; other features not in combination.............................. 11
11(10) Face imbricate; vertex punctate ( Fig. 33C View FIGURE 33 ); posterior margin of head distinctly concave in dorsal view ( Fig. 33C View FIGURE 33 ); metasoma dark brown, head, scape, pedicel, flagellomere 1 and propodeum reddish brown, legs and metasoma brown ( Figs 33A, B View FIGURE 33 ) [NSW and Queensland]....................................................... A. minutus sp. nov.
- Face other than imbricate; other features not in combination............................................... 12
12(11) Face rugulose; body length 8.1 mm, excluding ovipositor; ovipositor 5.0 mm; head and mesosoma black, metasoma largely orange, ovipositor sheaths black, ovipositor brown ( Figs 45A, B View FIGURE 45 ) [Tasmania]................ A. smithi sp. nov.
- Face rugulose-reticulate or rugose-reticulate; other features not in combination................................ 13
13(11) Body length excluding ovipositor, 8.0 mm; ovipositor 5.3 (5.0–5.7) mm; vertex rugose-reticulate, almost strigate-reticulate ( Fig. 53C View FIGURE 53 ); head and mesosoma black, metasomal T1 and T2 largely orange with remainin g segments blackish brown ( Figs 53A, B View FIGURE 53 ) [NSW]............................................................... A. williamsi sp. nov.
- Body length excluding ovipositor, 6.5 mm; ovipositor 3.7 mm; vertex rugose-reticulate, punctate-reticulate near ocelli ( Fig. 35C View FIGURE 35 ); body orange-brown [Queensland]................................................ A. nebo sp. nov.
14(9) Body length, excluding ovipositor, 9.45 (9.4–9.5) mm; ovipositor longer than body, 12.5 (12.0–13.0) mm [Victoria]...................................................................................... A. confusus sp. nov.
- Body length, excluding ovipositor, 7.5 mm; ovipositor shorter than body, 5.7 mm [Queensland].................................................................................................. A. mareebaensis sp. nov.
15(3) Face with punctures, sculpturing either punctate-rugose or punctate-reticulate................................. 16
- Face lacking punctures, sculpturing either rugose or rugulose.............................................. 17
16(15) Face punctate-rugose; antenna black except flagellomeres 4-7 cream ( Figs. 43A, B View FIGURE 43 ), ovipositor sheath brown; metasoma 1.3× length of mesosoma, and T1 and T2 narrow, glabrous, smooth tending to imbricate with scattered, small punctures posteriorly [Queensland].............................................................. A. scitulus sp. nov.
- Face punctate-reticulate; antenna black except flagellomeres 5–7 white ( Figs 37A, B View FIGURE 37 ); ovipositor sheath black except tip pale orange; metasoma 1.19 (1.12–1.21)× length of mesosoma, and T1 and T2 broad, several shallow and indistinct punctures antero-dorsally Tasmania, Victoria, Queensland]................................... A. obcordellus sp. nov.
17(15) Face rugulose (wrinkles fine); head, meso- and metasoma black, tibae and tarsi largely light brown; body length, excluding ovipositor, 10.5 mm [South Australia]................................................. A. moerens Westwood
- Face rugose (wrinkles coarse, e.g., Fig. 27C View FIGURE 27 ); body length excluding ovipositor, ≤ 9.0 mm....................... 18
18(17) Metasomal T1 and T2 rugulose dorsally, smooth laterally, lacking punctures; body length 7.6 mm, excluding ovipositor [ovipositor and sheath largely missing on holotype]; black except antenna orange-yellow, mandibles, fore and mid trochanters and femora brown, fore and mid tibiae and tarsi pale brown, hind legs dark brown, metasoma dark brown ( Fig. 21A View FIGURE 21 ) [Victoria]................................................................. A. flavimanus (Kieffer)
- Metasomal T1 and T2 with at least a few punctures; other features not in combination........................... 19
19(18) Metasomal T1 and T2 glabrous, except for a few indistinct, shallow punctures; body length 6.8 (5.8–7.3) mm, excluding ovipositor; ovipositor 2.8 (2.30–3.35) mm; ovipositor guide absent [Victoria]..................... A. brabyi sp. nov. - Metasomal T1 and T2 weakly rugulose-punctate dorsally; body length 9.0 mm, excluding ovipositor; ovipositor short, 3.5 mm; ovipositor guide medial, transverse [Victoria]........................................ A. flavicornis Kieffer
20(1) Metasoma clavate (e.g., Figs 9A, B View FIGURE 9 ).................................................................. 21
- Metasoma ovate (e.g., Figs 29A, B View FIGURE 29 )................................................................... 38
21(20) Fore wing with brown spot at apex (see Fig. 1 View FIGURE 1 )........................................................ 23
- Fore wing without brown spot at apex................................................................. 33
22(21) Hind wing venation incomplete, veins M+Cu, Cu, r-m and 2-M absent....................................... 23
- Hind wing venation complete, veins M+Cu, Cu, r-m and 2-M present (e.g., Fig. 1 View FIGURE 1 ).............................. 24
23(22) Ovipositor guide absent ( Fig. 15F View FIGURE 15 ); ovipositor 6.3 (6.0–6.5) mm; metasoma 1.80 (1.60–1.92)× length of mesosoma; metasoma dark brown [Victoria]..................................................... A. dandenongensis sp. nov.
- Ovipositor guide medial, oblique; ovipositor very short, 3.95 mm; metasoma 1.5× length of mesosoma; metasoma dark blackish brown with pale brown band dorsally before posterior margin of T2 ( Fig. 28B View FIGURE 28 ) [Queensland].................................................................................................. A. kittelae sp. nov.
24(22) Ovipositor guide absent; posterior margin of head not concave in dorsal view ( Fig. 54C View FIGURE 54 ) [Queensland]................................................................................................ A. wrightae sp. nov.
- Ovipositor guide present, either transverse or oblique; posterior margin of head concave in dorsal view............. 25
25(24) Ovipositor guide transverse......................................................................... 26
- Ovipositor guide oblique........................................................................... 28
26(24) Head with lateral medial carina present above toruli (e.g., Figs 11C View FIGURE 11 , 43F View FIGURE 43 )..................................... 27
- Head without lateral medial carina above toruli [South Australia]........... A. belairensis Jennings, Austin & Stevens
27(25) Ovipositor guide somewhat caudad, narrow ( Fig. 31G View FIGURE 31 ); head and mesosoma black, metasoma orange ( Figs 31A, B View FIGURE 31 ) [Queensland, NSW, ACT, Victoria]................................................. A. longiventris (Kieffer)
- Ovipositor guide medial, broad; body orange-brown [WA]................... A. houstoni Jennings, Austin & Stevens
28(25) Occipital carina present laterally, weak (narrow) ( Fig. 7D View FIGURE 7 ); body length excluding ovipositor 7.5 mm; ovipositor short, 3.0 mm [Queensland]................................................................ A. bamagensis sp. nov.
- Occipital carina absent; other features not in combination................................................. 29
29(28) Face with punctures............................................................................... 30
- Face without punctures............................................................................. 31
30(28) Face punctate-reticulate; frons without lateral medial carina above toruli; metasomal T2 with distinctive cream band dorso-laterally ( Figs 17A, B View FIGURE 17 ) [Queensland]................................................. A. doddi sp. nov.
- Face punctate-rugose; frons with lateral medial carina above toruli; body yellowish-brown, including metasomal T2 ( Figs 41A, B View FIGURE 41 ) [Queensland]................................................................. A. quickei sp. nov.
31(29) Head and mesosoma red-brown ( Figs 27A, B View FIGURE 27 ); face rugose ( Fig. 27C View FIGURE 27 ); ovipositor guide somewhat medial [NSW]...................................................................................... A. jamberoo sp. nov.
- Head and mesosoma largely black; other features not in combination........................................ 32
32(31) Body black except femora, tibiae and posterior metasomal T1 orange, tarsomeres predominantly white ( Figs 2A, B View FIGURE 2 ); face rugose; ovipositor guide somewhat caudal [Queensland and NSW].......................... A. albimanus (Kieffer)
- Body black except propleuron and ventral half of pronotum orange, metasomal T1 and T2 orange, and remainder dark brown ( Figs 13A, B View FIGURE 13 ); face rugulose; ovipositor guide medial [Queensland]........................ A. burnsi sp. nov.
33(21) Hind wing venation incomplete...................................................................... 34
- Hind wing venation complete (see Fig. 1 View FIGURE 1 ).............................................................. 35
34(33) Ovipositor guide transverse; head and mesosoma dark brown, metasoma yellow brown [Western Australia]............................................................................ A. mcmillani Jennings, Austin & Stevens
- Ovipositor guide somewhat oblique; head and mesosoma black, metasoma orange ( Figs 24A, B View FIGURE 24 ) [Queensland]........................................................................................... A glorious sp. nov.
35(33) Occipital carina present laterally, weak (narrow); face punctate-rugulose; mesosoma and metasoma about equal in length; ovipositor guide more or less oblique [South Australia]....................... A. grossi Jennings, Austin & Stevens
- Occipital carina absent; other features not in combination................................................. 36
36(35) Frons with lateral medial carina above toruli [Tasmania]..................................... A. simsoni sp. nov.
- Frons without lateral medial carina above toruli......................................................... 37
37(36) Face punctate-rugose ( Fig. 42C View FIGURE 42 ); body dark brown-black [ACT, NSW, Victoria].................... A. rieki sp. nov.
- Face without punctures, rugose-reticulate; head and mesosoma black, metasoma orange except for variable amounts of black on T1 ( Figs 23A, B View FIGURE 23 ) [NSW, Queensland]........................................ A. fuscicornis (Cameron)
38(21) Ovipositor guide absent............................................................................ 39
- Ovipositor guide present............................................................................ 42
39(38) Frons with lateral carina above toruli (e.g., Figs 11C View FIGURE 11 , 43F View FIGURE 43 )................................................. 40
- Frons without lateral carina above toruli............................................................... 41
40(39) Face rugulose-reticulate; head and mesothorax black, metasoma dark brown ( Figs 38A, B View FIGURE 38 ) [NSW].............................................................................................. A. pallidicaudis (Cameron)
- Face rugose; body black except legs, and metasomal T1 and T2 with variable amounts of light brown ( Figs 4A, B View FIGURE 4 ) [Queensland, South Australia]................................................................ A. aquilus sp. nov.
41(39) Face punctate-reticulate; head and mesosoma brown except some orange on gena, propleuron and lateral pronotum, scape, pedicel, legs and metasoma orange ( Figs 22A, B View FIGURE 22 ) [NSW].................................... A. froggatti sp. nov.
- Face rugose; body black except scape orange, fore and mid trochanters dark orange, all femora orange, fore and mid tar- someres 1–4 cream, all tarsomeres 5 brown, hind tibiae dark brown and first two-thirds basitarsus dark brown, apical onethird basitarsus and tarsomeres 2–4 cream ( Figs 8A, B View FIGURE 8 ) [Tasmania]........................... A. bashfordi sp. nov.
42(38) Ovipositor guide transverse......................................................................... 43
- Ovipositor guide oblique........................................................................... 45
43(42) Body entirely black ( Fig. 42A View FIGURE 42 ); length 11.4 mm, excluding ovipositor; ovipositor 17.0 mm [NSW, Queensland]..................................................................................... A. planiceps (Szépligeti)
- Body generally dark brown or head largely orange and mesosoma black...................................... 44
44(43) Length 6.2 (5.6–6.6) mm, excluding ovipositor; ovipositor 7.6 (7.5–7.7) mm; head orange except for variable amounts of dark brown medially on frons and vertex, mesosoma black ( Figs 16A, B View FIGURE 16 ) [Victoria]................ A. deansi sp. nov.
- Length 12.5 mm, excluding ovipositor; ovipositor long, 14.3 mm; generally dark brown, with variable amounts of black on frons and face, pronotum, mesoscutum, metanotum, mesopleuron, mesepimeron, basally on mid and hind coxae, hind trochanter and trochantellus ( Figs 19A, B View FIGURE 19 ) [Queensland]...................................... A. festivus (Kieffer)
45(42) Ovipositor guide somewhat distal.................................................................... 46
- Ovipositor guide medial............................................................................ 48
46(45) Face rugose, punctate-rugose laterally; ovipositor guide distal ( Fig. 12F View FIGURE 12 ); length 7.3 mm, excluding ovipositor; ovipositor 9.7 mm; head and metasoma dark brown, mesosoma black [ACT].............................. A. broadi sp. nov.
- Face lacking punctures; other features not in combination................................................. 47
47(46) Face rugulose-reticulate; head and mesosoma black, metasoma black except somites 1–3 orange, with variable amounts of black anteriorly on T1 ( Figs 10A, B View FIGURE 10 ); metasoma 1.22 (1.15–1.26)× length of mesosoma [NSW]................................................................................................ A. boonanghiensis sp. nov.
- Face rugulose; head black, meso- and metasoma orange ( Figs 6A, B View FIGURE 6 ); metasoma 1.3 (1.2–1.4)× length of mesosoma [Tasmania]............................................................................. A. atriceps Kieffer
48(45) Head black...................................................................................... 49
- Head orange to dark brown......................................................................... 54
49(48) Mesosoma and metasoma orange..................................................................... 50
- Mesosoma black, metasoma variable in colour, but not entirely orange....................................... 51
50(49) Face rugulose; length 9.6 mm, excluding ovipositor; ovipositor long, 10.6 mm [Queensland]...... A. umbackae sp. nov.
- Face punctate-rugose; length 8.0 mm, excluding ovipositor; ovipositor short, 6.5 mm [Tasmania]................................................................................................... A. warraensis sp. nov.
51(49) Ovipositor short, <6.0 mm.......................................................................... 52
- Ovipositor long,> 9.0 mm.......................................................................... 53
52(51) Length 7.0 mm, excluding ovipositor; ovipositor short, 5.5 mm; head 1.35× wider than long when viewed dorsally; face rugulose; metasoma predominantly dark brown, T1 and T2 paler ( Fig. 48A View FIGURE 48 ) [Victoria].......... A. truncatus (Kieffer)
- Length 7.1 mm, excluding ovipositor; ovipositor 5.5 mm; head 1.10× wider than long when viewed dorsally; face rugose; metasoma dark brown ( Fig. 46B View FIGURE 46 ) [Tasmania]........................................... A. tasmanicus sp. nov.
53(51) Metasoma 1.6 (1.5–1.7)× length of mesosoma; ovipositor 11.0 (10.5–11.5) mm; metasoma with T1 black, T2 orange, T3– T8 dark brown ( Fig. 43B View FIGURE 43 ) [ACT and NSW]................................................. A. anici sp. nov.
- Metasoma 1.3× length of mesosoma; ovipositor 9.4 mm; metasoma dark brown except T2 generally paler ( Fig. 5B View FIGURE 5 ) [Queensland]....................................................................... A. aroueti (Girault)
54(48) Large specimens, length 16.5 mm, excluding ovipositor; ovipositor 18.8 mm; head and body largely orange-red; ovipositor guide medial and oblique [Western Australia]............................. A. douglasi Jennings, Austin & Stevens
- Small specimens. length, excluding ovipositor, less than 10 mm; other features not in combination................. 55
55(54) Head dark brown, meso- and metasoma pale orange ( Figs 32A, B View FIGURE 32 ); length 6.8 mm, excluding ovipositor; ovipositor 6.1 mm; metasoma 1.4× length of mesosoma [Victoria]........................................ A. pallidus sp. nov.
- Head and metasoma orange, mesosoma brown ( Figs 47A, B View FIGURE 47 ); length 7.6 (6.5–8.25) mm, excluding ovipositor; ovipositor 13.6 (13–14.2) mm; metasoma 1.5× length of mesosoma [Victoria]............................. A. tiernyi sp. nov.
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