Thomomys bottae (Eydoux and Gervais, 1836)

James L. Patton, 1993, Order Rodentia - Family Geomyidae, Mammal Species of the World (2 nd Edition), Washington and London: Smithsonian Institution Press, pp. 469-476 : 473-474

Thomomys bottae (Eydoux and Gervais, 1836) . Mag. Zool., Paris, 6:23.

TYPE LOCALITY: USA, coast of California (restricted to vicinity of Monterey, Monterey Co., by Baird, 1857) .

DISTRIBUTION: SW and W USA, north to Oregon, east to Colorado, and south to Sinaloa and Nuevo Leon ( Mexico).

SYNONYMS: abbotti Huey , absonus Goldman, abstrusus Hall and Davis, acrirostratus Grinnell, actuosus Kelson, adderrans Huey, affinis Huey , agricolaris Grinnell, 'albatus Grinnell, albicaudatus Hall, 'alexandrae Goldman, alienus Goldman, 'alpinus Merriam, alticolus J. A. Allen, 'altivallis Rhoads, amargosae Grinnell, analogus Goldman, 'angularis Merriam, angustidens Baker, anitae J. A. Allen, 'apache Bailey, 'aphrastus Elliot, ardicola Huey, 'argusensis Huey, aureiventris Hall, 'aureus J. A. Allen, awahnee Merriam, 'baileyi Merriam, basilicae Benson and Tillotson, birdseyei Goldman, bonnevillei Durant, boregoensis Huey, boreorarius Durham, borjasensis Huey, brazierhowelli Huey, brevidens Hall, 'cabezonae Merriam, cactophilus Huey, camargensis Anderson, camoae Burt, caneloensis Lange, 'canus Bailey, carri Lange, catalinae Goldman, catavinensis Huey, cedrinus Huey, centralis Hall, 'cervinus J. A. Allen, chiricahuae Nelson and Goldman, 'chrysonotus Grinnell, cinereus Hall, collinus Goldman, collis Hooper, comobabiensis Huey, concisor Hall and Davis, confinalis Goldman, connectens Hall, contractus Durrant, convergens Nelson and Goldman, convexus Durrant, crassus Chattin, cultellus Kelson, cunicularis Huey , curtatus Hall, depauperatus Grinnell and Hill, depressus Hall, 'desertorum Merriam, desitus Goldman, detumidus Grinnell, 'diaboli Grinnell, dissimilis Goldman, divergens Nelson and Goldman, estanciae Benson and Tillotson, extenuatus Goldman, flavidus Goldman, 'fulvus Woodhouse, fumosus Hall , grahamensis Goldman, growlerensis Huey, guadalupensis Goldman, 'harquahalae Grinnell, homorus Huey, howelli Goldman, hualpaiensis Goldman, hueyi Goldman, humilis Baker, imitabilis Goldman, incomptus Goldman, 'infrapallidus Grinnell, ingens Grinnell, internatus Goldman, 'jacinteus Grinnell and Swarth, jojobae Huey, juarezensis Huey, 'lachuguilla Bailey, lacrymalis Hall, 'laticeps Baird, 'latirostris Merriam, latus Hall and Davis, lenis Goldman, 'leucodon Merriam, levidensis Goldman, limitaris Goldman, limpiae Blair, litoris Burt, lorenzi Huey, lucidus Hall, lucrificus Hall and Durham, ' magdalenae Nelson and Goldman , martirensis J. A. Allen, 'mearnsi Bailey, ' melanotis Grinnell , 'mewa Merriam, minimus Durrant , minor Bailey, modicus Goldman, mohavensis Grinnell, morulus Hooper, ' muralis Goldman , mutabilis Goldman, nnnws Hall, nasutus Hall, navus Merriam, ' neglectus Bailey , nesophilus Durrant, nicholi Goldman, 'nigricans Rhoads, occipitalis Benson and Tillotson, *operarius Merriam, operosus Hatfield, optabilis Goldman, opulentus Goldman, "oreoecus Burt, osgoodi Goldman, paguatae Hooper, pallescens Rhoads , parvulus Goldman , pascalis Merriam, patulus Goldman, *pectoralis Goldman, peramplus Goldman, *perpallidus Merriam, perpes Merriam, pervagus Merriam, pervarius Goldman, phasma Goldman, *phelleoecus Burt, pinalensis Goldman, piutensis Grinnell and Hill, planirostris Burt, pianorum Hooper, pomelli Durrant, *providentialis Grinnell, proximarinus Huey, proximus Burt and Campbell, puertae Grinnell, pusillus Goldman , retractus Baker, rhizophagus Huey, riparius Grinnell and Hill, robustus Durrant, rubidus Youngman, rufidulus Hoffmeister, ruidosae Hall, rupestris Chattin , ruricola Huey, russeolus Nelson and Goldman, sanctidiegi Huey, saxatilis Grinnell, *scapterus Elliot, scotophilus Davis, sevieri Durrant, siccovallis Huey, silvifugus Grinnell, *simulus Nelson and Goldman, *sinaloae Merriam, *solitarius Grinnell, spatiosus Goldman, stansburyi Durrant, *sturgisi Goldman, suboles Goldman, subsimilis Goldman, texensis Bailey , tivius Durrant, *toltecus J.A. Allen, trumbullensis Hall and Davis, tularosae Hall, vanrosseni Huey, varus Hall and Long, vescus Hall and Davis, villai Baker, virgineus Goldman, wahwahensis Durrant, winthropi Nelson and Goldman, xerophilus Huey ('those taxa originally described as distinct species or treated as such in the literature).

COMMENTS: Subgenus Megascapheus . The taxonomic history of this, and related species has been contentious. Hall and Kelson (1959) included bottae within umbrinus but considered baileyi and townsendii as separate species, but Hall (1981) included all four of these taxa within his concept of umbrinus . Anderson (1966, 1972), Hoffmeister (1969, 1986), and Patton and co-workers ( Patton and Dingman, 1968; Patton, 1973; Patton and Smith, 1981) considered bottae (including baileyi) separate from umbrinus . Thaeler (19686), Patton et al. (1984), Patton and Smith (1989), and Rogers (1991«, 6) supported the specific separation of townsendii from Hall's (1981) umbrinus . As envisioned here, bottae includes a number of taxa considered by earlier workers to be full species but which are each now relegated to subspecific status or are recognized as synonyms of other subspecies. Revisions of those subspecies occurring in Arizona and California have been presented by Hoffmeister (1986) and Patton and Smith (1990), respectively. Hoffmeister (1969), Patton and Dingman (1968), and Patton (1973) reported on hybridization between bottae and umbrinus in S Arizona; Thaeler (19686) and Patton et al. (1984) examined hybridization between bottae and townsendii in NE California. For subspecies, see Hall (1981), as modified for Arizona by Hoffmeister (1986), and for California by Patton and Smith (1990).