Thomomys umbrinus (Richardson, 1829)

James L. Patton, 1993, Order Rodentia - Family Geomyidae, Mammal Species of the World (2 nd Edition), Washington and London: Smithsonian Institution Press, pp. 469-476 : 475-476

Thomomys umbrinus (Richardson, 1829) . Quadrupeds, in Fauna Boreali-Americana, 1:202.

TYPE LOCALITY: Originally given as Cadadaguios, southwestern Louisiana, but restricted to Mexico, Veracruz (probably Puebla), vicinity of Boca del Norte, by Bailey (1906) .

DISTRIBUTION: SC Arizona and SW New Mexico ( USA) south to Puebla and Veracruz ( Mexico).

STATUS: IUCN - Rare.

SYNONYMS: albigularis Nelson and Goldman , arriagensis Dalquest, atrodorsalis Nelson and Goldman, atrovarius J. A. Allen , burti Huey, caliginosus Nelson and Goldman, chihuahuae Nelson and Goldman, crassidens Nelson and Goldman, durangi Nelson and Goldman, emotus Goldman, enixus Nelson and Goldman, evexus Nelson and Goldman, eximius Nelson and Goldman, extimus Nelson and Goldman, goldmani Merriam , intermedius Mearns , juntae Anderson, madrensis Nelson and Goldman, martinertsis Nelson and Goldman, musculus Nelson and Goldman , nelsoni Merriam , newmani Dalquest, orizabae Merriam, parviceps Nelson and Goldman, perditus Nelson and Goldman, peregrinus Merriam , potosinus Nelson and Goldman, pullus Hall and Villa , quercinus Burt and Campbell, sheldoni Nelson and Goldman , sonoriensis Nelson and Goldman, supernus Nelson and Goldman, tolucae Nelson and Goldman, vulcanius Nelson and Goldman, zacatecae Nelson and Goldman.

COMMENTS: Hybridization with bottae reported in Patagonia Mtns of S Arizona by Patton and Dingman (1968), Patton (1973), and Hoffmeister (1969). Considered a species separate from bottae by these authors and by Anderson (1966, 1972). Hall (1981:469) included baileyi, bottae , and townsendii in this species based on a his view of the hybridization data in Hoffmeister (1969), Patton (1973), and Thaeler (1968b); he reported intergradation between umbrinus and bottae in Nuevo Leon ( Mexico). Phylogenetic relationships among geographic units of umbrinus were examined by Hafner et al. (1987).